A quadrat is a frame, traditionally square, used in

ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...

geography Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth description") is a field of science devoted to the study of the lands, features, inhabitants, and ...

and

biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...

to isolate a standard unit of area for study of the distribution of an item over a large area. Modern quadrats can for example be rectangular, circular, or irregular. The quadrat is suitable for sampling

plants Plants are predominantly Photosynthesis, photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all curr ...

, slow-moving animals, and some aquatic organisms. A photo-quadrat is a photographic record of the area framed by a quadrat. It may use a physical frame to indicate the area, or may rely on fixed camera distance and lens

field of view The field of view (FoV) is the extent of the observable world that is seen at any given moment. In the case of optical instruments or sensors it is a solid angle through which a detector is sensitive to electromagnetic radiation. Human ...

to automatically cover the specified area of substrate. Parallel laser pointers mounted on the camera can also be used as scale indicators. The photo is taken perpendicular to the surface, or as close as possible to perpendicular for uneven surfaces.

History

The systematic use of quadrats was developed by the pioneering plant ecologists R. Pound and F. E. Clements between 1898 and 1900. The method was then swiftly applied for many purposes in

, such as the study of

plant succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as ca ...

. Botanists and ecologists such as

Arthur Tansley Sir Arthur George Tansley FLS, FRS (15 August 1871 – 25 November 1955) was an English botanist and a pioneer in the science of ecology. Educated at Highgate School, University College London and Trinity College, Cambridge, Tansley taught a ...

soon took up and modified the method. The ecologist J. E. Weaver applied the use of quadrats to the teaching of ecology in 1918.

Method

When an

ecologist Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...

wants to know how many organisms there are in a particular

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

, it would not be feasible to count them all. Instead, they would be forced to count a smaller representative part of the population, called a sample. Sampling of plants or slowly moving animals (such as snails) can be done using a sampling square called a quadrat. A suitable size of a quadrat depends on the size of the organisms being sampled. For example, to count plants growing on a school field, one could use a quadrat with sides 0.5 or 1 metre in length. Choice of quadrat size depends to a large extent on the type of survey being conducted. For instance, it would be difficult to gain any meaningful results using a 0.5m2 quadrat in a study of a woodland canopy. Botanist and students examining the species distribution using quadrat

Botanist and students examining the species distribution using quadrat

It is important that sampling in an area is carried out at random, to avoid bias. For example, if you were sampling from a school field, but for convenience only placed quadrats next to a path, this might not give a sample that was representative of the whole field. It would be an unrepresentative, or

bias Bias is a disproportionate weight ''in favor of'' or ''against'' an idea or thing, usually in a way that is closed-minded, prejudicial, or unfair. Biases can be innate or learned. People may develop biases for or against an individual, a group ...

ed, sample. One way one can sample randomly is to place the quadrats at coordinates on a numbered grid. Long-term studies may require that the same quadrats be revisited months or even years after initial sampling. Methods of relocating the precise area of study vary widely in accuracy, and include measurement from nearby permanent markers, use of

total station A total station (TS) or total station theodolite (TST) is an electronic/optical instrument used for surveying and building construction. It is an electronic transit theodolite integrated with electronic distance measurement (EDM) to measure ...

theodolites, consumer-grade

GPS The Global Positioning System (GPS), originally Navstar GPS, is a Radionavigation-satellite service, satellite-based radionavigation system owned by the United States government and operated by the United States Space Force. It is one of t ...

, and

differential GPS Differential Global Positioning Systems (DGPSs) supplement and enhance the positional data available from global navigation satellite systems (GNSSs). A DGPS for GPS can increase accuracy by about a thousandfold, from approximately to . DGPSs c ...

References

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External links

Ecological Sampling Methods
Ecology terminology Environmental statistics Environmental Sampling Equipment

History

Method

See also

References

External links