Pseudalopex culpaeus
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The culpeo (''Lycalopex culpaeus''), also known as culpeo zorro, Andean zorro, Andean fox, Paramo wolf, Andean wolf,Comparative ecology of two South American foxes, 'Dusicvon ariseus' and 'culpaeus' by Warren E. Johnson. Doctoral dissertation. Iowa State University; 1992. p2. Accessed July 10, 2021 at https://lib.dr.iastate.edu/cgi/viewcontent.cgi?article=11006&context=rtd and colpeo fox, is a species of South American fox. Regardless of the name, it is not a true fox, but more closely related to wolves and jackals. Its appearance resembles that of foxes due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. It is the second-largest native
canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
on the
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions In geography, regions, otherwise referred to as zones, lands or territories, are areas t ...
after the
maned wolf The maned wolf (''Chrysocyon brachyurus'') is a large canine of South America. It is found in Argentina, Brazil, Bolivia, Peru, and Paraguay, and is almost extinct in Uruguay. Its markings resemble those of foxes, but it is neither a fox nor a w ...
. In appearance, it bears many similarities to the widely recognized red fox. It has grey and reddish fur, a white chin, reddish legs and a stripe on its back that may be barely visible. The culpeo's
diet Diet may refer to: Food * Diet (nutrition), the sum of the food consumed by an organism or group * Dieting, the deliberate selection of food to control body weight or nutrient intake ** Diet food, foods that aid in creating a diet for weight loss ...
consists largely of
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s, rabbits,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s and lizards, and to a lesser extent,
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
material and carrion. The culpeo does attack
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticated ...
on occasion and is, therefore, often hunted or poisoned. In some regions, it has become rare, but overall the species is not threatened with extinction. The culpeo was
domesticated Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
by the
Selk'nam people The Selk'nam, also known as the Onawo or Ona people, are an indigenous people in the Patagonian region of southern Argentina and Chile, including the Tierra del Fuego islands. They were one of the last native groups in South America to be enco ...
of
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
producing the
Fuegian dog The Fuegian dog ( es, perro yagán, perro fueguino), also known as the Yahgan dog, is an extinct domesticated canid. It was a domesticated form of the culpeo (''Lycalopex culpaeus''). The culpeo is similar in build to true foxes (tribe Vulpini) b ...
, which is now extinct.


Description

The culpeo is a canid intermediate in size between a red fox and a coyote. The average weight of the male is , while the typically smaller females average . Overall, a weight range of has been reported. Total length can range from , including a tail of in length.Burnie D and Wilson DE (Eds.) (2005), ''Animal: The Definitive Visual Guide to the World's Wildlife''. DK Adult, The pelt has a grizzled appearance. The neck and shoulders are often tawny to rufous in color, while the upper back is dark. The bushy tail has a black tip.


Range

Its distribution extends from
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
to the southern regions of
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
and
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
. Some populations live in the southern regions of Colombia. It is most common on the western slopes of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
, where it inhabits open country and
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s.


Habitat

The culpeo lives in a wide variety of habitats of western South America. They are found in broadleaf ''
Nothofagus ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gui ...
''
temperate rainforest Temperate rainforests are coniferous or broadleaf forests that occur in the temperate zone and receive heavy rain. Temperate rain forests occur in oceanic moist regions around the world: the Pacific temperate rain forests of North American ...
,
sclerophyllous Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
matorral,
deserts A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
, chaparrals, and
plateaus In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ha ...
, like the
Altiplano The Altiplano (Spanish for "high plain"), Collao (Quechua and Aymara: Qullaw, meaning "place of the Qulla") or Andean Plateau, in west-central South America, is the most extensive high plateau on Earth outside Tibet. The plateau is located at the ...
, up to the tree line ().


Diet

The culpeo is an opportunistic predator that will take any variety of prey. It mainly feeds on rodents and
lagomorph The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (Î»Î±Î³Ï ...
s (especially the introduced
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has b ...
and
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
); however, it occasionally feeds on domestic livestock and young
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco ...
s. Culpeos are considered beneficial because they are significant predators of the rabbits introduced in 1915; such introduced rabbit populations are believed to have allowed culpeos to spread from the Andean foothills across the Patagonian plain. They sometimes take young lambs up to 1 week old. In limited studies, the larger culpeo appears to dominate potential competitors, including
South American gray fox The South American gray fox (''Lycalopex griseus''), also known as the Patagonian fox, the chilla or the gray zorro, is a species of ''Lycalopex'', the "false" foxes. It is endemic to the southern part of South America. Description The South A ...
es,
Geoffroy's cat Geoffroy's cat (''Leopardus geoffroyi'') is a small wild cat native to the southern and central regions of South America. It is about the size of a domestic cat. It is listed as Least Concern on the IUCN Red List because it is widespread and abun ...
s,
pampas cat The Pampas cat (''Leopardus colocola'') is a small wild cat native to South America. It is listed as Near Threatened on the IUCN Red List as habitat conversion and destruction may cause the population to decline in the future. It is named after ...
s,
grison A grison , , also known as a South American wolverine, is any mustelid in the genus ''Galictis''. Native to Central and South America, the genus contains two extant species: the greater grison (''Galictis vittata''), which is found widely in Sou ...
s and various raptorial birds. Its range also overlaps that of the much larger puma, but the size difference ensures that the two species have limited competition. They are known to eat the carcasses of
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', its former genus name) is one of the two wild South American camelids, which live in the high alpine areas of the Andes, the other being the guanaco, which live ...
s. Culpeos have also been observed preying upon introduced beavers. During a period of drought in central Chile's scrublands lagomorphs,
coati Coatis, also known as coatimundis (), are members of the family Procyonidae in the genera ''Nasua'' and ''Nasuella''. They are diurnal mammals native to South America, Central America, Mexico, and the southwestern United States. The name ...
,
goats The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of th ...
, and
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
make up a large amount of their diet, but it is unknown if cattle is hunted or scavenged.


Reproduction

The typical mating period is between August and October. After a gestation period of 55–60 days, the female gives birth usually to between two and five pups.


Classification


Subspecies

File:Lycalopex culpaeus Bolivia.jpg, ''Lycalopex culpaeus andinus'' (Thomas, 1914) File:Lycalopex culpaeus culpaeus in the Buin Zoo, Chile..jpg, ''Lycalopex culpaeus culpaeus'' (Molina, 1782) File:Lycalopex culpaeus lycoides in Ushuaia, Argentina..jpg, ''Lycalopex culpaeus lycoides'' (Philippi, 1896) File:Fox -Puerto Natales, Patagonia, Chile-8.jpg, ''Lycalopex culpaeus magellanicus'' (Gray, 1837) File:Pes horský.jpg, ''Lycalopex culpaeus reissii'' (Hilzheimer, 1906)


Taxonomy

The taxonomy of the culpeo has been the topic of debate due to their high phenetic variability and the scarcity of research, among other things. Over the past three decades, they have been placed variably in the genera '' Dusicyon'' (Clutton-Brock, ''et al.'', 1976; Wozencraft, 1989), ''
Canis ''Canis'' is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and de ...
'' (Langguth, 1975; Van Gelder, 1978), ''
Pseudalopex The South American foxes (''Lycalopex''), commonly called ''raposa'' in Portuguese, or ''zorro'' in Spanish, are a genus from South America of the subfamily Caninae. Despite their name, they are not true foxes, but are a unique canid genus m ...
'' (Berta, 1987; Wozencraft, 1993; Tedford ''et al.'', 1995) and '' Lycalopex'' (Zunino, 1995; Wozencraft, 2005). This canid, like other South American foxes, is still sometimes classified as a member of the genus ''Pseudalopex''. As ''Pseudalopex'' and ''Lycalopex'' have largely come to describe the same genus, either classification is acceptable, although the modern practice is to give ''Lycalopex'' prominence. ::


References

{{Taxonbar, from=Q373558 South American foxes Mammals of the Andes Mammals of Patagonia Mammals of Argentina Mammals of Bolivia Mammals of Chile Mammals of Colombia Mammals of Ecuador Mammals of Peru Fauna of Tierra del Fuego Fauna of the Falkland Islands Carnivorans of South America Mammals described in 1782 Least concern biota of South America