Population model
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A population model is a type of mathematical model that is applied to the study of population dynamics.


Rationale

Model A model is an informative representation of an object, person or system. The term originally denoted the plans of a building in late 16th-century English, and derived via French and Italian ultimately from Latin ''modulus'', a measure. Models c ...
s allow a better understanding of how complex interactions and processes work. Modeling of dynamic interactions in nature can provide a manageable way of understanding how numbers change over time or in relation to each other. Many patterns can be noticed by using population modeling as a tool.
Ecological Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
population modeling is concerned with the changes in parameters such as
population size In population genetics and population ecology, population size (usually denoted ''N'') is the number of individual organisms in a population. Population size is directly associated with amount of genetic drift, and is the underlying cause of effect ...
and age distribution within a population. This might be due to interactions with the environment, individuals of their own species, or other species. Population models are used to determine maximum harvest for agriculturists, to understand the dynamics of biological invasions, and for
environmental conservation *Environmental protection *Nature conservation Nature conservation is the moral philosophy and conservation movement focused on protecting species from extinction, maintaining and restoring habitats, enhancing ecosystem services, and protec ...
. Population models are also used to understand the spread of parasites, viruses, and
disease A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
. Another way populations models are useful are when species become endangered. Population models can track the fragile species and work and curb the decline


History

Late 18th-century biologists began to develop techniques in population modeling in order to understand the dynamics of growing and shrinking of all populations of living organisms.
Thomas Malthus Thomas Robert Malthus (; 13/14 February 1766 – 29 December 1834) was an English cleric, scholar and influential economist in the fields of political economy and demography. In his 1798 book ''An Essay on the Principle of Population'', Mal ...
was one of the first to note that populations grew with a geometric pattern while contemplating the fate of humankind. One of the most basic and milestone models of population growth was the
logistic model of population growth Logistic may refer to: Mathematics * Logistic function, a sigmoid function used in many fields ** Logistic map, a recurrence relation that sometimes exhibits chaos ** Logistic regression, a statistical model using the logistic function ** Logit, ...
formulated by
Pierre François Verhulst Pierre François Verhulst (28 October 1804, Brussels – 15 February 1849, Brussels) was a Belgian mathematician and a doctor in number theory from the University of Ghent in 1825. He is best known for the logistic growth model. Logistic e ...
in 1838. The logistic model takes the shape of a
sigmoid curve A sigmoid function is a mathematical function having a characteristic "S"-shaped curve or sigmoid curve. A common example of a sigmoid function is the logistic function shown in the first figure and defined by the formula: :S(x) = \frac = \ ...
and describes the growth of a population as exponential, followed by a decrease in growth, and bound by a carrying capacity due to environmental pressures. Population modeling became of particular interest to biologists in the 20th century as pressure on limited means of sustenance due to increasing human populations in parts of Europe were noticed by biologist like
Raymond Pearl Raymond Pearl (June 3, 1879 – November 17, 1940) was an American biologist, regarded as one of the founders of biogerontology. He spent most of his career at Johns Hopkins University in Baltimore. Pearl was a prolific writer of academic books, ...
. In 1921 Pearl invited physicist Alfred J. Lotka to assist him in his lab. Lotka developed paired
differential equation In mathematics, a differential equation is an equation that relates one or more unknown functions and their derivatives. In applications, the functions generally represent physical quantities, the derivatives represent their rates of change, an ...
s that showed the effect of a parasite on its prey. Mathematician
Vito Volterra Vito Volterra (, ; 3 May 1860 – 11 October 1940) was an Italian mathematician and physicist, known for his contributions to mathematical biology and integral equations, being one of the founders of functional analysis. Biography Born in An ...
equated the relationship between two species independent from Lotka. Together, Lotka and Volterra formed the Lotka–Volterra model for competition that applies the logistic equation to two species illustrating competition, predation, and parasitism interactions between species. In 1939 contributions to population modeling were given by Patrick Leslie as he began work in biomathematics. Leslie emphasized the importance of constructing a life table in order to understand the effect that key life history strategies played in the dynamics of whole populations. Matrix algebra was used by Leslie in conjunction with life tables to extend the work of Lotka. Matrix models of populations calculate the growth of a population with life history variables. Later,
Robert MacArthur Robert Helmer MacArthur (April 7, 1930 – November 1, 1972) was a Canadian-born American ecologist who made a major impact on many areas of community and population ecology. Early life and education MacArthur was born in Toronto, Ontario, ...
and
E. O. Wilson Edward Osborne Wilson (June 10, 1929 – December 26, 2021) was an American biologist, naturalist, entomologist and writer. According to David Attenborough, Wilson was the world's leading expert in his specialty of myrmecology, the study of an ...
characterized island biogeography. The equilibrium model of island biogeography describes the number of species on an island as an equilibrium of immigration and extinction. The logistic population model, the Lotka–Volterra model of community ecology, life table matrix modeling, the equilibrium model of island biogeography and variations thereof are the basis for ecological population modeling today.


Equations

Logistic growth A logistic function or logistic curve is a common S-shaped curve (sigmoid curve) with equation f(x) = \frac, where For values of x in the domain of real numbers from -\infty to +\infty, the S-curve shown on the right is obtained, with the ...
equation: : \frac = rN\left(1-\frac\right)\,
Competitive Lotka–Volterra equations The competitive Lotka–Volterra equations are a simple model of the population dynamics of species competing for some common resource. They can be further generalised to the Generalized Lotka–Volterra equation to include trophic interactions ...
: : \frac = r_1 N_1\frac\,
Island biogeography Insular biogeography or island biogeography is a field within biogeography that examines the factors that affect the species richness and diversification of isolated natural communities. The theory was originally developed to explain the pattern of ...
: : S = \frac Species–area relationship: : \log(S) = \log(c)+z \log(A)\,


Examples of individual-based models


See also

* Population dynamics *
Population dynamics of fisheries A fishery is an area with an associated fish or aquatic population which is harvested for its commercial or recreational value. Fisheries can be wild or farmed. Population dynamics describes the ways in which a given population grows and shrinks ...
*
Population ecology Population ecology is a sub-field of ecology that deals with the dynamics of species populations and how these populations interact with the environment, such as birth and death rates, and by immigration and emigration. The discipline is import ...
*
Moment closure In probability theory, moment closure is an approximation method used to estimate moments of a stochastic process. Introduction Typically, differential equations describing the ''i-''th moment will depend on the ''(i + 1)''-st moment. T ...


References


External links


GreenBoxes code sharing network
Greenboxes (Beta) is a repository for open-source population modeling code. Greenboxes allows users an easy way to share their code and to search for others shared code. {{DEFAULTSORT:Population Modeling Conceptual models Population models