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The Yamnaya culture or the Yamna culture (russian: Ямная культура, ua, Ямна культура lit. 'culture of pits'), also known as the Pit Grave culture or Ochre Grave culture, was a late Copper Age to early
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second prin ...
archaeological culture of the region between the Southern Bug,
Dniester The Dniester, ; rus, Дне́стр, links=1, Dnéstr, ˈdⁿʲestr; ro, Nistru; grc, Τύρᾱς, Tyrās, ; la, Tyrās, la, Danaster, label=none, ) ( ,) is a transboundary river in Eastern Europe. It runs first through Ukraine and th ...
, and
Ural Ural may refer to: *Ural (region), in Russia and Kazakhstan *Ural Mountains, in Russia and Kazakhstan *Ural (river), in Russia and Kazakhstan * Ual (tool), a mortar tool used by the Bodo people of India *Ural Federal District, in Russia *Ural econ ...
rivers (the Pontic steppe), dating to 3300–2600 BCE. It was discovered by Vasily Gorodtsov following his archaelogical excavations near
Siversky Donets The Seversky Donets () or Siverskyi Donets (), usually simply called the Donets, is a river on the south of the East European Plain. It originates in the Central Russian Upland, north of Belgorod, flows south-east through Ukraine (Kharkiv, Done ...
in 1901—1903. Its name derives from its characteristic burial tradition: (
romanization Romanization or romanisation, in linguistics, is the conversion of text from a different writing system to the Roman (Latin) script, or a system for doing so. Methods of romanization include transliteration, for representing written text, a ...
: ) is a Russian adjective that means 'related to pits ()', as these people used to bury their dead in tumuli (
kurgan A kurgan is a type of tumulus constructed over a grave, often characterized by containing a single human body along with grave vessels, weapons and horses. Originally in use on the Pontic–Caspian steppe, kurgans spread into much of Central As ...
s) containing simple pit chambers. The people of the Yamnaya culture were likely the result of a
genetic admixture Genetic admixture occurs when previously diverged or isolated genetic lineages mix.⅝ Admixture results in the introduction of new genetic lineages into a population. Examples Climatic cycles facilitate genetic admixture in cold periods and gene ...
between the descendants of Eastern European Hunter-Gatherers (EHG)The Eastern European hunter-gatherers were themselves mostly descended from ancient North Eurasians, related to the palaeolithic
Mal'ta–Buret' culture The Mal'ta–Buret' culture is an archaeological culture of 24,000 to 15,000 BP / 22'050 to 13'050 BC in the Upper Paleolithic on the upper Angara River in the area west of Lake Baikal in the Irkutsk Oblast, Siberia, Russian Federation. The ...
.
and people related to hunter-gatherers from the Caucasus (CHG), an ancestral component which is often named "Steppe ancestry", with additional admixture of up to 18% from
Early European Farmers Early European Farmers (EEF), First European Farmers (FEF), Neolithic European Farmers, Ancient Aegean Farmers, or Anatolian Neolithic Farmers (ANF) are names used to describe a distinct group of early Neolithic farmers who brought agriculture to E ...
. Their material culture was very similar to the
Afanasevo culture The Afanasievo culture, or Afanasevo culture (Afanasevan culture) (russian: Афанасьевская культура ''Afanas'yevskaya'' kul'tura), is the earliest known archaeological culture of south Siberia, occupying the Minusinsk Basin a ...
, and the populations of both cultures are genetically indistinguishable. They lived primarily as nomads, with a chiefdom system and wheeled carts and wagons that allowed them to manage large herds. The people of the Yamnaya culture are also closely connected to Final Neolithic cultures, which later spread throughout
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
and
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
, especially the
Corded Ware The Corded Ware culture comprises a broad archaeological horizon of Europe between ca. 3000 BC – 2350 BC, thus from the late Neolithic, through the Chalcolithic, Copper Age, and ending in the early Bronze Age. Corded Ware culture en ...
people and the
Bell Beaker culture The Bell Beaker culture, also known as the Bell Beaker complex or Bell Beaker phenomenon, is an archaeological culture named after the inverted-bell beaker drinking vessel used at the very beginning of the European Bronze Age. Arising from ar ...
, as well as the peoples of the
Sintashta Sintashta (russian: Синташта́) is an archaeological site in Chelyabinsk Oblast, Russia. It is the remains of a fortified settlement dating to the Bronze Age, ''c''. 2800–1600 BC, and is the type site of the Sintashta culture. The site h ...
, Andronovo, and Srubnaya cultures. Back migration from Corded Ware also contributed to Sintashta and Andronovo. In these groups, several aspects of the Yamnaya culture are present.Yamnayan cultural aspects, for example, were horse-riding, burial styles, and to some extent the pastoralist economy. Genetic studies have also indicated that these populations derived large parts of their ancestry from the steppes. The Yamnaya culture is identified with the late
Proto-Indo-Europeans The Proto-Indo-Europeans are a hypothetical prehistoric population of Eurasia who spoke Proto-Indo-European (PIE), the ancestor of the Indo-European languages according to linguistic reconstruction. Knowledge of them comes chiefly from ...
, and the Pontic-Caspian steppe is the strongest candidate for the ''
Urheimat In historical linguistics, the homeland or ''Urheimat'' (, from German '' ur-'' "original" and ''Heimat'', home) of a proto-language is the region in which it was spoken before splitting into different daughter languages. A proto-language is the r ...
'' (original homeland) of the
Proto-Indo-European language Proto-Indo-European (PIE) is the reconstructed common ancestor of the Indo-European language family. Its proposed features have been derived by linguistic reconstruction from documented Indo-European languages. No direct record of Proto-Indo-E ...
.


Origins

Yamnaya culture was defined by Vasily Gorodtsov in order to differentiate it from catacomb and
srubnaya culture The Srubnaya culture (russian: Срубная культура, Srubnaya kul'tura, ua, Зрубна культура, Zrubna kul'tura), also known as Timber-grave culture, was a Late Bronze Age 1850–1450 BC cultureParpola, Asko, (2012)"Format ...
s that existed in the area but considered to be of later period. Owing to the historical distance of Yamnaya culture, and reliance on archaeological findings, debate continues as to its origin. In 1996
Pavel Dolukhanov Pavel Dolukhanov (January 1, 1937, Leningrad, USSR – December 6, 2009, Newcastle, UK) was a doctor of Geographical Sciences, Professor, Emeritus Professor (2002), Russian and British paleogeographer and archaeologist at the Institute of History ...
suggested that the emergence of the Pit-Grave culture represents a social development of various local Bronze Age cultures, representing "an expression of social stratification and the emergence of chiefdom-type nomadic social structures", which in turn intensified inter-group contacts between essentially heterogeneous social groups. According to , "The origin of the Yamnaya culture is still a topic of debate," with proposals for its origins pointing to both Khvalynsk and Sredny Stog. The Khvalynsk culture (4700–3800 BC) (middle Volga) and the Don-based Repin culture (c. 3950–3300 BC) in the eastern Pontic-Caspian steppe, and the closely related
Sredny Stog culture The Sredny Stog culture (, romanized: ''Serednʹostohivsʹka kulʹtura'') is a pre-Kurgan archaeological culture from the 5th millennium BC. It is named after the Russian term for the Dnieper river islet of today's Seredny Stih, Ukraine, where ...
(c. 4500–3500 BC) in the western Pontic-Caspian steppe, preceded the Yamnaya culture (3300–2500 BC). Further efforts to pinpoint the location came from Anthony (2007), suggesting from his research that the Yamnaya culture (3300–2600 BC) originated in the
Don Don, don or DON and variants may refer to: Places *County Donegal, Ireland, Chapman code DON *Don (river), a river in European Russia *Don River (disambiguation), several other rivers with the name *Don, Benin, a town in Benin *Don, Dang, a vill ...
Volga The Volga (; russian: Во́лга, a=Ru-Волга.ogg, p=ˈvoɫɡə) is the longest river in Europe. Situated in Russia, it flows through Central Russia to Southern Russia and into the Caspian Sea. The Volga has a length of , and a catchm ...
area at c. 3400 BC, preceded by the middle Volga-based Khvalynsk culture and the Don-based
Repin culture The Repin culture was a 4th millennium BCE Eneolithic archaeological culture located in the Pontic–Caspian steppe and East European forest steppe. It developed from preceding local Neolithic cultures, and later developed into the Yamnaya culture ...
(–3300 BC), arguing that late pottery from these two cultures can barely be distinguished from early Yamnaya pottery. Earlier continuity from eneolithic but largely hunter-gatherer Samara culture and influences from the more agricultural Dnieper–Donets II are apparent. He argues that the early Yamnaya horizon spread quickly across the
Pontic–Caspian steppe The Pontic–Caspian steppe, formed by the Caspian steppe and the Pontic steppe, is the steppeland stretching from the northern shores of the Black Sea (the Pontus Euxinus of antiquity) to the northern area around the Caspian Sea. It extend ...
s between and 3200 BC: Alternatively, Parpola (2015) relates both the Corded ware culture and the Yamnaya culture to the late Tripolye culture. He hypothesizes that "the Tripolye culture was taken over by PIE speakers by c. 4000 BC," and that in its final phase the Tripolye culture expanded to the steppes, morphing into various regional cultures which fused with the late Sredny Stog pastoralist cultures, which, he suggests, gave rise to the Yamnaya culture. The Yamnaya (Pit-grave) culture was succeeded in its western range by the
Catacomb culture The Catacomb culture (russian: Катакомбная культура, Katakombnaya kul'tura, uk, Катакомбна культура, Katakombna kul'tura) was a Bronze Age culture which flourished on the Pontic steppe in 2500–1950 BC.Par ...
(2800–2200 BC); in the east, by the
Poltavka culture Poltavka culture () was an early to middle Bronze Age archaeological culture which flourished on the Volga- Ural steppe and the forest steppe in 2800—2200 BCE. The Poltavka culture emerged as an eastern outgrowth of the Yamnaya culture, ...
(2700–2100 BC) at the middle Volga. These two cultures were followed by the
Srubnaya culture The Srubnaya culture (russian: Срубная культура, Srubnaya kul'tura, ua, Зрубна культура, Zrubna kul'tura), also known as Timber-grave culture, was a Late Bronze Age 1850–1450 BC cultureParpola, Asko, (2012)"Format ...
(18th–12th century BC). File:Sredny Stog culture.jpg, Sredny Stog culture (c.4500-3500 BC) File:Usatovo culture.jpg, Usatovo culture (c. 3500–3000 BC) File:Khvalynsk culture.jpg, Khvalynsk culture (c. 4900–3500 BC) File:Early Yamna.jpg, Early Yamnaya culture (3400 BC), according to


Characteristics

The Yamnaya culture was
semi-nomadic A nomad is a member of a community without fixed habitation who regularly moves to and from the same areas. Such groups include hunter-gatherers, pastoral nomads (owning livestock), tinkers and trader nomads. In the twentieth century, the p ...
, with some
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
practiced near rivers, and a few fortified sites, the largest of which is Mikhaylovka. Characteristic for the culture are the burials in pit graves under
kurgan A kurgan is a type of tumulus constructed over a grave, often characterized by containing a single human body along with grave vessels, weapons and horses. Originally in use on the Pontic–Caspian steppe, kurgans spread into much of Central As ...
s (
tumuli A tumulus (plural tumuli) is a mound of earth and stones raised over a grave or graves. Tumuli are also known as barrows, burial mounds or ''kurgans'', and may be found throughout much of the world. A cairn, which is a mound of stones built ...
), often accompanied with animal offerings. Some graves contain large anthropomorphic stelae, with carved human heads, arms, hands, belts, and weapons. The dead bodies were placed in a
supine position The supine position ( or ) means lying horizontally with the face and torso facing up, as opposed to the prone position, which is face down. When used in surgical procedures, it grants access to the peritoneal, thoracic and pericardium, pericardi ...
with bent knees and covered in ochre. Some kurgans contained "stratified sequences of graves". It has been argued that kurgan burials were rare, and reserved for special adults, who were predominantly, but not necessarily, male. Status and gender are marked by grave goods and position, and in some areas, elite individuals are buried with complete wooden wagons. Grave goods are more common in eastern Yamnaya burials, which are also characterized by a higher proportion of male burials and more male-centred rituals than western areas. In the northern Pontic steppes were excavated the oldest wheels in the world, which may tentatively be associated with the Indo-Europeans. The Yamnaya culture had and used two-wheeled carts and four-wheeled wagons, which are thought to have been oxen-drawn at this time, and there is evidence that they rode horses. Metallurgists and other craftsmen are given a special status in Yamnaya society, and metal objects are sometimes found in large quantities in elite graves. New metalworking technologies and weapon designs are used. Anthony speculates that the Yamnaya ate a diet consisting of meat, milk, yogurt, cheese, and soups made from seeds and wild vegetables, and probably consumed mead. Mallory and Adams suggest that Yamnaya society may have had a tripartite structure of three differentiated social classes, although the evidence available does not demonstrate the existence of specific classes such as priests, warriors, and farmers.


Archaeogenetics

According to Jones et al. (2015) and Haak et al. (2015),
autosomal An autosome is any chromosome that is not a sex chromosome. The members of an autosome pair in a diploid cell have the same morphology, unlike those in allosomal (sex chromosome) pairs, which may have different structures. The DNA in autosom ...
tests indicate that the Yamnaya people were the result of a genetic admixture between two different hunter-gatherer populations: distinctive "
Eastern Hunter-Gatherer In archaeogenetics, the term Eastern Hunter-Gatherer (EHG), sometimes East European Hunter-Gatherer, or Eastern European Hunter-Gatherer is the name given to a distinct ancestral component that represents descent from Mesolithic hunter-gatherers o ...
s" (EHG), from Eastern Europe, with high affinity to the
Mal'ta–Buret' culture The Mal'ta–Buret' culture is an archaeological culture of 24,000 to 15,000 BP / 22'050 to 13'050 BC in the Upper Paleolithic on the upper Angara River in the area west of Lake Baikal in the Irkutsk Oblast, Siberia, Russian Federation. The ...
or other, closely related people from
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
and a population of " Caucasus hunter-gatherers" (CHG) who probably arrived from the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia (country), Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range ...
or Iran. Each of those two populations contributed about half the Yamnaya DNA. This admixture is referred to in archaeogenetics as
Western Steppe Herder In archaeogenetics, the term Western Steppe Herders (WSH), or Western Steppe Pastoralists, is the name given to a distinct ancestral component first identified in individuals from the Chalcolithic, Eneolithic steppe around the turn of the 5th mi ...
(WSH) ancestry. Admixture between EHGs and CHGs is believed to have occurred on the eastern Pontic-Caspian steppe starting around 5,000 BC, while admixture with
Early European Farmer Early European Farmers (EEF), First European Farmers (FEF), Neolithic European Farmers, Ancient Aegean Farmers, or Anatolian Neolithic Farmers (ANF) are names used to describe a distinct group of early Neolithic farmers who brought agriculture to E ...
s (EEF) happened in the southern parts of the Pontic-Caspian steppe sometime later. More recent genetic studies have found that the Yamnaya were a mixture of EHGs, CHGs, and to a lesser degree Anatolian farmers and Levantine farmers, but not EEFs from Europe due to lack of WHG DNA in the Yamnaya. This occurred in two distinct admixture events from
West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
into the Pontic-Caspian steppe. Haplogroup R1b, especially subclades of R1b-M269, is the most common
Y-DNA The Y chromosome is one of two sex chromosomes (allosomes) in therian mammals, including humans, and many other animals. The other is the X chromosome. Y is normally the sex-determining chromosome in many species, since it is the presence or abse ...
haplogroup found among both the Yamnaya and modern-day Western Europeans. Additionally, a minority are found to belong to haplogroup I2. They are found to belong to a wider variety of
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
haplogroups, including U, T, and haplogroups associated with Caucasus Hunter-Gatherers and
Early European Farmer Early European Farmers (EEF), First European Farmers (FEF), Neolithic European Farmers, Ancient Aegean Farmers, or Anatolian Neolithic Farmers (ANF) are names used to describe a distinct group of early Neolithic farmers who brought agriculture to E ...
s. It has been hypothesized that an allele associated with
lactase persistence Lactase persistence is the continued activity of the lactase enzyme in adulthood, allowing the digestion of lactose in milk. In most mammals, the activity of the enzyme is dramatically reduced after weaning. In some human populations, though, lact ...
(conferring lactose tolerance into adulthood) was brought to Europe from the steppe by Yamnaya-related migrations. "Furthermore, ancient DNA studies found that the LP mutation was absent or very rare in Europe until the end of the Bronze Age 6–29and appeared first in individuals with steppe ancestry 9,20 Thus, it was proposed that the mutation originated in Yamnaya-associated populations and arrived later in Europe by migration of these steppe herders." "the 101 sequenced individuals, the Yamnaya were most likely to have the DNA variation responsible for lactose tolerance, hinting that the steppe migrants might have eventually introduced the trait to Europe" "For example, one lineage could have a biological evolutionary advantage over the other. Allentoft et al. (2015: 171) have found a remarkably high rate of lactose tolerance among individuals connected to Yamnaya and to Corded Ware, as opposed to the majority of Late Neolithic individuals." People of the Yamnaya culture are believed to have had mostly brown eye colour, light to intermediate skin, and brown hair colour, with some variation. Some individuals are believed to have carried a mutation to the KITLG gene associated with blond hair, as several individuals with Steppe ancestry are later found to carry this mutation. The
Ancient North Eurasian In archaeogenetics, the term Ancient North Eurasian (generally abbreviated as ANE) is the name given to an ancestral component that represents a lineage ancestral to the people of the Mal'ta–Buret' culture and populations closely related to th ...
population, who contributed significant ancestry to
Western Steppe Herders In archaeogenetics, the term Western Steppe Herders (WSH), or Western Steppe Pastoralists, is the name given to a distinct ancestral component first identified in individuals from the Eneolithic steppe around the turn of the 5th millennium BCE, ...
, are believed to be the source of this mutation. A study in 2015 found that Yamnaya had the highest ever calculated genetic selection for height of any of the ancient populations tested. The geneticist David Reich has argued that the genetic data supports the likelihood that the people of the Yamnaya culture were a "single, genetically coherent group" who were responsible for spreading many Indo-European languages. Reich also argues that the genetic evidence shows that Yamnaya society was an oligarchy dominated by a small number of elite males. The genetic evidence for the extent of the role of the Yamnaya culture in the spread of Indo-European languages has however been questioned by Russian archaeologist
Leo Klejn Lev Samuilovich Kleyn (; 1 July 1927 – 7 November 2019), better known in English as Leo Klejn, was a Russian archaeologist, anthropologist and philologist. Early life Klejn was born in Vitebsk, Belarus, to two Jewish physicians, Polish-born S ...
and Balanovsky et al., who note a lack of male haplogroup continuity between the people of the Yamnaya culture and the contemporary populations of Europe.


Language

Marija Gimbutas Marija Gimbutas ( lt, Marija Gimbutienė, ; January 23, 1921 – February 2, 1994) was a Lithuanian archaeologist and anthropologist known for her research into the Neolithic and Bronze Age cultures of " Old Europe" and for her Kurgan hypothesis ...
identified the Yamnaya culture with the late
Proto-Indo-Europeans The Proto-Indo-Europeans are a hypothetical prehistoric population of Eurasia who spoke Proto-Indo-European (PIE), the ancestor of the Indo-European languages according to linguistic reconstruction. Knowledge of them comes chiefly from ...
(PIE) in her
Kurgan hypothesis The Kurgan hypothesis (also known as the Kurgan theory, Kurgan model, or steppe theory) is the most widely accepted proposal to identify the Proto-Indo-European homeland from which the Indo-European languages spread out throughout Europe and par ...
. The Pontic-Caspian steppe is the strongest candidate for the ''
Urheimat In historical linguistics, the homeland or ''Urheimat'' (, from German '' ur-'' "original" and ''Heimat'', home) of a proto-language is the region in which it was spoken before splitting into different daughter languages. A proto-language is the r ...
'' (original homeland) of the
Proto-Indo-European language Proto-Indo-European (PIE) is the reconstructed common ancestor of the Indo-European language family. Its proposed features have been derived by linguistic reconstruction from documented Indo-European languages. No direct record of Proto-Indo-E ...
, and evidence from linguistics and genetics suggests that the Yamnaya culture may be the homeland of the core Indo-European languages, excluding the Anatolian languages. According to David W. Anthony, the genetic evidence suggests that the leading clans of the Yamnaya were of EHG (Eastern European hunter-gatherer) and WHG (Western European hunter-gatherer) paternal origin and implies that the
Indo-European languages The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, Dutc ...
were the result of "a dominant language spoken by EHGs that absorbed Caucasus-like elements in phonology, morphology, and lexicon."


Yamnaya-related migrations


Western Europe

Haak et al. (2015) conducted a genome-wide study of 69 ancient skeletons from Europe and Russia. They concluded that Yamnaya autosomal characteristics are very close to the
Corded Ware culture The Corded Ware culture comprises a broad archaeological horizon of Europe between ca. 3000 BC – 2350 BC, thus from the late Neolithic, through the Copper Age, and ending in the early Bronze Age. Corded Ware culture encompassed a v ...
people, with an estimated 73% ancestral contribution from the Yamnaya DNA in the DNA of Corded Ware skeletons from
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
. The same study estimated a (38.8–50.4 %) ancestral contribution of the Yamnaya in the DNA of modern Central, and Northern Europeans, and an 18.5–32.6 % contribution in modern Southern Europeans; this contribution is found to a lesser extent in
Sardinians The Sardinians, or Sards ( sc, Sardos or ; Italian and Sassarese: ''Sardi''; Gallurese: ''Saldi''), are a Romance language-speaking ethnic group native to Sardinia, from which the western Mediterranean island and autonomous region of Italy de ...
(2.4–7.1 %) and
Sicilians Sicilians or the Sicilian people are a Romance speaking people who are indigenous to the island of Sicily, the largest island in the Mediterranean Sea, as well as the largest and most populous of the autonomous regions of Italy. Origin and i ...
(5.9–11.6 %). Haak et al. also note that their results state that haplogroup R-M269 spread into Europe from the East after 3000 BC. Studies that analysed ancient human remains in
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
and
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
support the thesis that R-M269 was introduced in these places along with autosomal DNA from the Eastern European steppes. Autosomal tests also indicate that the Yamnaya are the most likely vector for "Ancient North Eurasian" admixture into Europe. "
Ancient North Eurasian In archaeogenetics, the term Ancient North Eurasian (generally abbreviated as ANE) is the name given to an ancestral component that represents a lineage ancestral to the people of the Mal'ta–Buret' culture and populations closely related to th ...
" is the name given in literature to a genetic component that represents descent from the people of the
Mal'ta–Buret' culture The Mal'ta–Buret' culture is an archaeological culture of 24,000 to 15,000 BP / 22'050 to 13'050 BC in the Upper Paleolithic on the upper Angara River in the area west of Lake Baikal in the Irkutsk Oblast, Siberia, Russian Federation. The ...
or a population closely related to them. That genetic component is visible in tests of the Yamnaya people as well as modern-day Europeans, but not of Europeans predating the Bronze Age.


Eastern Europe and Finland

In the Baltic, Jones et al. (2017) found that the
Neolithic transition The Neolithic Revolution, or the (First) Agricultural Revolution, was the wide-scale transition of many human cultures during the Neolithic period from a lifestyle of hunting and gathering to one of agriculture and settlement, making an inc ...
 – the passage from a hunter-gatherer economy to a farming-based economy – coincided with the arrival en masse of individuals with Yamnaya-like ancestry. This is different from what happened in Western and Southern Europe, where the Neolithic transition was caused by a population that came from Anatolia, with Pontic steppe ancestry being detected from only the late Neolithic onward. Per Haak et al. (2015), the Yamnaya contribution in the modern populations of
Eastern Europe Eastern Europe is a subregion of the European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russia, whic ...
ranges from 46.8% among
Russians , native_name_lang = ru , image = , caption = , population = , popplace = 118 million Russians in the Russian Federation (2002 '' Winkler Prins'' estimate) , region1 = , pop1 ...
to 42.8% in
Ukrainians Ukrainians ( uk, Українці, Ukraintsi, ) are an East Slavic ethnic group native to Ukraine. They are the seventh-largest nation in Europe. The native language of the Ukrainians is Ukrainian. The majority of Ukrainians are Eastern Ort ...
.
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...
has one of the highest Yamnaya contributions in all of Europe (50.4%).Per , adding a north-Siberian people as a fourth reference population improves residuals for northeastern European populations. This accounts for the higher than expected Yamnaya contribution and brings it down to expected levels (67.8–50.4 % in Finns, 64.9–46.8 % in Russians).


Central and South Asia

Studies also point to the strong presence of Yamnaya descent in the current nations of South Asia, especially in groups that are referred to as Indo-Aryans. According to Pathak et al. (2018), the "North-Western Indian & Pakistani" populations (PNWI) showed significant Middle-Late Bronze Age Steppe (Steppe_MLBA) ancestry along with Yamnaya Early-Middle Bronze Age (Steppe_EMBA) ancestry, but the Indo-Europeans of
Gangetic Plains The Indo-Gangetic Plain, also known as the North Indian River Plain, is a fertile plain encompassing northern regions of the Indian subcontinent, including most of northern and eastern India, around half of Pakistan, virtually all of Ba ...
and
Dravidian people The Dravidian peoples, or Dravidians, are an ethnolinguistic and cultural group living in South Asia who predominantly speak any of the Dravidian languages. There are around 250 million native speakers of Dravidian languages. Dravidian spe ...
only showed significant Yamnaya (Steppe_EMBA) ancestry and no Steppe_MLBA. The study also noted that ancient south Asian samples had significantly higher Steppe_MLBA than Steppe_EMBA (or Yamnaya). The study identified the Rors and Jats as the population in South Asia with the highest proportion of Steppe ancestry. Lazaridis et al. (2016) estimated (6.5–50.2 %) steppe-related admixture in South Asians, though the proportion of Steppe ancestry varies widely across ethnic groups. According to Narasimhan et al. (2019), the Yamnaya-related ancestry, termed Western_Steppe_EMBA, that reached central and south Asia was not the initial expansion from the steppe to the east, but a secondary expansion that involved a group possessing ~67% Western_Steppe_EMBA ancestry and ~33% ancestry from the European cline. This group included people similar to that of
Corded Ware The Corded Ware culture comprises a broad archaeological horizon of Europe between ca. 3000 BC – 2350 BC, thus from the late Neolithic, through the Chalcolithic, Copper Age, and ending in the early Bronze Age. Corded Ware culture en ...
, Srubnaya, Petrovka, and
Sintashta Sintashta (russian: Синташта́) is an archaeological site in Chelyabinsk Oblast, Russia. It is the remains of a fortified settlement dating to the Bronze Age, ''c''. 2800–1600 BC, and is the type site of the Sintashta culture. The site h ...
. Moving further east in the central steppe, it acquired ~9% ancestry from a group of people that possessed West Siberian Hunter Gatherer ancestry, thus forming the Central Steppe MLBA cluster, which is the primary source of steppe ancestry in South Asia, contributing up to 30% of the ancestry of the modern groups in the region. According to Unterländer et al. (2017), all Iron Age
Scythian The Scythians or Scyths, and sometimes also referred to as the Classical Scythians and the Pontic Scythians, were an ancient Eastern * : "In modern scholarship the name 'Sakas' is reserved for the ancient tribes of northern and eastern Centra ...
Steppe nomads can best be described as a mixture of Yamnaya-related ancestry and an East Asian-related component, which most closely corresponds to the modern North
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
n Nganasan people of the lower
Yenisey River The Yenisey (russian: Енисе́й, ''Yeniséy''; mn, Горлог мөрөн, ''Gorlog mörön''; Buryat: Горлог мүрэн, ''Gorlog müren''; Tuvan: Улуг-Хем, ''Uluğ-Hem''; Khakas: Ким суғ, ''Kim suğ''; Ket: Ӄук, ...
, to varying degrees, but generally higher among Eastern Scythians, while Bronze Age Steppe nomads of Central Asia had the East Asian-related component at only trace levels.


Artifacts

Hermitage Museum The State Hermitage Museum ( rus, Государственный Эрмитаж, r=Gosudarstvennyj Ermitaž, p=ɡəsʊˈdarstvʲɪn(ː)ɨj ɪrmʲɪˈtaʂ, links=no) is a museum of art and culture in Saint Petersburg, Russia. It is the largest ...
collections"> File:Yamna02.jpg, Corded ware pot File:Yamna03.jpg File:Yamna05.jpg File:Yamna04.jpg File:KAM56.jpg, alt=, Bronze daggers, arrowheads and bone artefacts


See also

*
Kurgan A kurgan is a type of tumulus constructed over a grave, often characterized by containing a single human body along with grave vessels, weapons and horses. Originally in use on the Pontic–Caspian steppe, kurgans spread into much of Central As ...
*
Kurgan stelae Kurgan stelae ( Mongolian: ; Russian: ; Ukrainian: "stone babas"; ky, балбал ) or Balbals ( ''balbal'', most probably from a Turkic word ' meaning "ancestor" or "grandfather") are anthropomorphic stone stelae, images cut from stone, i ...
* Butmir culture *
Vinča culture The Vinča culture (), also known as Turdaș culture or Turdaș–Vinča culture, is a Neolithic archaeological culture of Southeast Europe, dated to the period 5700–4500 BC or 5300–4700/4500 BC.. Named for its type site, Vinča-Belo Brdo, ...
*
Beaker culture The Bell Beaker culture, also known as the Bell Beaker complex or Bell Beaker phenomenon, is an archaeological culture named after the inverted-bell beaker drinking vessel used at the very beginning of the European Bronze Age. Arising from a ...
*
Baden culture The Baden culture was a Chalcolithic culture from 3520–2690 BC. It was found in Central and Southeast Europe, and is in particular known from Moravia (Czech Republic), Hungary, southern Poland, Slovakia, northern Croatia and eastern Austria. I ...
* Botai culture * Khvalynsk culture *
Mamai-Hora Mamai–Hora is the largest multicomplex burial mound of national importance in the Northern Black Sea region. It is located on the left bank of the Kakhovka Reservoir near the village of Velyka Znamianka in Vasylivka Raion, Zaporizhzhia Oblast. ...
* Samara culture *
Sintashta culture The Sintashta culture (russian: Синташтинская культура, Sintashtinskaya kul'tura), around 2050–1900 BCE, is the first phase of the Sintashta–Petrovka culture. or Sintashta–Arkaim culture,. and is a late Middle Bronze Ag ...
* ''
Yersinia pestis ''Yersinia pestis'' (''Y. pestis''; formerly '' Pasteurella pestis'') is a gram-negative, non-motile, coccobacillus bacterium without spores that is related to both ''Yersinia pseudotuberculosis'' and ''Yersinia enterocolitica''. It is a facult ...
'' *
Proto-Indo-Europeans The Proto-Indo-Europeans are a hypothetical prehistoric population of Eurasia who spoke Proto-Indo-European (PIE), the ancestor of the Indo-European languages according to linguistic reconstruction. Knowledge of them comes chiefly from ...
*
Indigenous Aryanism Indigenous Aryanism, also known as the Indigenous Aryans theory (IAT) and the Out of India theory (OIT), is the conviction that the Aryans are indigenous to the Indian subcontinent, and that the Indo-European languages radiated out from a homela ...


Notes


References


Notes


Sources

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Supplementary Information
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External links

* * {{cite news , url=https://www.pbs.org/wgbh/nova/video/first-horse-warriors/?linkId=67160903 , website=PBS.org NOVA series , date=May 15, 2019 , title=First Horse Warriors Indo-European archaeological cultures Archaeological cultures of Eastern Europe Bronze Age cultures of Europe Chalcolithic cultures of Europe Archaeological cultures in Ukraine Archaeological cultures in Russia Archaeological cultures in Moldova Archaeology of Kazakhstan 4th millennium BC Prehistoric Russia