Pine nuts
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Pine nuts, also called piñón (), pinoli (), pignoli or chilgoza (), are the edible
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s of
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
s (family
Pinaceae The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly kn ...
, genus ''Pinus''). According to the Food and Agriculture Organization, only 29 species provide edible nuts, while 20 are traded locally or internationally owing to their seed size being large enough to be worth harvesting; in other pines, the seeds are also edible, but are too small to be of notable value as a human food.


Species and geographic spread

In Asia, two species in particular are widely harvested: Korean pine (''Pinus koraiensis'') in northeast Asia (the most important species in international trade) and chilgoza pine (''Pinus gerardiana'') in the western
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
. Four other species, Siberian pine (''Pinus sibirica''),
Siberian dwarf pine ''Pinus pumila'', commonly known as the Siberian dwarf pine, dwarf Siberian pine, dwarf stone pine, Japanese stone pine, or creeping pine, is a tree in the family Pinaceae native to northeastern Asia and the Japanese isles. It shares the commo ...
(''Pinus pumila''), Chinese white pine (''Pinus armandii'') and lacebark pine (''Pinus bungeana''), are also used to a lesser extent. Russia is the largest producer of ''Pinus sibirica'' nuts in the world, followed by
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
, which produces over 10,000 tonnes of forest-grown nuts annually. The majority of the harvest is exported to China.
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is borde ...
is an important source of pine nuts, behind China and Korea. Pine nuts produced in Europe mostly come from the stone pine (''Pinus pinea''), which has been cultivated for its nuts for over 5,000 years. Pine nuts have been harvested from wild trees for far longer. The Swiss pine (''Pinus cembra'') is also used, to a very small extent. In North America, the main species are three of the
pinyon pine The pinyon or piñon pine group grows in southwestern North America, especially in New Mexico, Arizona, and Utah. The trees yield edible nuts, which are a staple food of Native Americans, and widely eaten as a snack and as an ingredient in ...
s:
Colorado pinyon ''Pinus edulis'', the Colorado pinyon, two-needle piñon, pinyon pine, or simply piñon, is a pine in the pinyon pine group whose ancestor was a member of the Madro-Tertiary Geoflora (a group of drought resistant trees) and is native to the Unite ...
(''Pinus edulis''), single-leaf pinyon (''Pinus monophylla''), and Mexican pinyon (''Pinus cembroides''). The other eight pinyon species are used to a small extent, as are gray pine (''Pinus sabineana''),
Coulter pine The Coulter pine or big-cone pine, ''Pinus coulteri'', is a native of the coastal mountains of Southern California in the United States and northern Baja California in Mexico. Isolated groves are found as far north as Clearlake, California on the ...
(''Pinus coulteri''),
Torrey pine The Torrey pine (''Pinus torreyana'') is a rare pine species in California, United States. It is a critically endangered species growing only in coastal San Diego County, and on Santa Rosa Island, offshore from Santa Barbara in Santa Barbara ...
(''Pinus torreyana''),
sugar pine ''Pinus lambertiana'' (commonly known as the sugar pine or sugar cone pine) is the tallest and most massive pine tree, and has the longest cones of any conifer. The species name ''lambertiana'' was given by the Scottish botanist David Douglas, ...
(''Pinus lambertiana'') and
Parry pinyon ''Pinus quadrifolia'', the Parry pinyon, is a pine in the pinyon pine group native to southernmost California in the United States and northern Baja California in Mexico, from 33° 30' N south to 30° 30' N. The Parry pinyon has a lifespan of aro ...
(''Pinus quadrifolia''). Here, the nuts themselves are known by the Spanish name for the pinyon pine, ''piñón'' (plural: ''piñones''). In the United States, pine nuts are mainly harvested by Native American and Hispano communities, particularly in the Western United States and Southwestern United States, by the
Uto-Aztecan Uto-Aztecan, Uto-Aztekan or (rarely in English) Uto-Nahuatl is a family of indigenous languages of the Americas, consisting of over thirty languages. Uto-Aztecan languages are found almost entirely in the Western United States and Mexico. The na ...
Shoshone The Shoshone or Shoshoni ( or ) are a Native American tribe with four large cultural/linguistic divisions: * Eastern Shoshone: Wyoming * Northern Shoshone: southern Idaho * Western Shoshone: Nevada, northern Utah * Goshute: western Utah, easte ...
, Paiute,
Navajo The Navajo (; British English: Navaho; nv, Diné or ') are a Native Americans in the United States, Native American people of the Southwestern United States. With more than 399,494 enrolled tribal members , the Navajo Nation is the largest fe ...
,
Pueblo In the Southwestern United States, Pueblo (capitalized) refers to the Native tribes of Puebloans having fixed-location communities with permanent buildings which also are called pueblos (lowercased). The Spanish explorers of northern New Spain ...
,
Hopi The Hopi are a Native American ethnic group who primarily live on the Hopi Reservation in northeastern Arizona, United States. As of the 2010 census, there are 19,338 Hopi in the country. The Hopi Tribe is a sovereign nation within the United ...
, Washoe, and
Hispanos of New Mexico The Hispanos of New Mexico, also known as Neomexicanos ( es, Neomexicano) or Nuevomexicanos, are Hispanic residents originating in the historical region of Santa Fe de Nuevo México, today the US state of New Mexico (''Nuevo México''), south ...
. Certain treaties negotiated by tribes and laws in
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
guarantee Native Americans' right to harvest pine nuts, and the state of
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
protects the use of the word ''piñon'' for use with pine nuts from certain species of indigenous New Mexican pines.


Species list

For those seeking to grow edible landscapes, these are the more commonly used species. *Old World **''Pinus armandii'' – Chinese white pine **''Pinus bungeana'' – lacebark pine **''Pinus cembra'' – Swiss pine **''Pinus gerardiana'' – Chilgoza pine **''Pinus koraiensis'' – Korean pine **''Pinus pinea'' – Mediterranean stone pine **''Pinus pumila'' –
Siberian dwarf pine ''Pinus pumila'', commonly known as the Siberian dwarf pine, dwarf Siberian pine, dwarf stone pine, Japanese stone pine, or creeping pine, is a tree in the family Pinaceae native to northeastern Asia and the Japanese isles. It shares the commo ...
**''Pinus sibirica'' – Siberian pine *New World **''Pinus albicaulis'' –
Whitebark pine ''Pinus albicaulis'', known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine ...
**''Pinus cembroides'' – Mexican pinyon **''Pinus coulteri'' –
Coulter pine The Coulter pine or big-cone pine, ''Pinus coulteri'', is a native of the coastal mountains of Southern California in the United States and northern Baja California in Mexico. Isolated groves are found as far north as Clearlake, California on the ...
**''Pinus culminicola'' –
Potosi pinyon ''Pinus culminicola'', commonly known as Potosí pinyon or Potosí Piñón, is a pine in the pinyon pine group, native and endemic to northeast Mexico. The range is highly localised, confined to a small area of high summits in the northern Sierra ...
**''Pinus edulis'' –
Two-needle piñon ''Pinus edulis'', the Colorado pinyon, two-needle piñon, pinyon pine, or simply piñon, is a pine in the pinyon pine group whose ancestor was a member of the Madro-Tertiary Geoflora (a group of drought resistant trees) and is native to the Unite ...
or Colorado pinyon (when grown in Colorado) **''Pinus johannis'' – Johann's pinyon (includes P. discolor – Border pinyon) **''Pinus monophylla'' – Single-leaf pinyon **''Pinus orizabensis'' – Orizaba pinyon **''Pinus quadrifolia'' – Four-leaved pinyon or Parry pinyon **''Pinus remota'' – Papershell pinyon or Texas pinyon **''Pinus sabiniana'' – California foothill pine


Pollination and seed development

The pine nut (seed) species will take a time that depends on the exact species (e.g., 36 months for a stone pine seed) to complete its maturity; to reach full maturity, the environmental conditions must be favorable for the tree and its cone. For some American species, development begins in early spring with pollination. A tiny cone, about the size of a small marble, will form from mid-spring to the end of summer; the premature cone will then become and remain dormant (with a cessation of growth) until the following spring. The cone will then commence growth until it reaches maturity near the end of summer. The mature piñon pine cone is ready to harvest ten days before the green cone begins to open. A cone is harvested by placing it in a
burlap bag A burlap sack or gunny sack, also known as a gunny shoe, hessian sack or tow sack, is an inexpensive bag, traditionally made of fibres which are also known as "tow," such as hessian fabric (burlap) formed from jute, hemp or other natural fibr ...
and exposing it to a heat source such as the sun to begin the drying process. It takes about 20 days until the cone fully opens. Once it is fully open and dry, the seed can be easily extracted in various ways. The most common and practical extraction method used is the repeated striking of the burlap bag containing the cone(s) against a rough surface to cause the cone(s) to shatter, leaving just the job of separating by hand the seed from the residue within the bag. Another option for harvesting is to wait until the cone opens on the tree (as it naturally will) and harvest the cone from the piñon pine, followed by the extracting process mentioned above. Fallen seed can also be gathered beneath the trees.


Ecology and status

Because pine nuts are an important food source for many animals, overharvesting of pine nuts threatens local ecosystems, an effect occurring during the early 21st century with increased
culinary Culinary arts are the cuisine arts of food preparation, cooking and presentation of food, usually in the form of meals. People working in this field – especially in establishments such as restaurants – are commonly called chefs ...
uses for pine nuts. In the United States, millions of hectares of productive pinyon pine woods have been destroyed due to conversion of lands, and in China and Russia, destructive harvesting techniques (such as breaking off whole branches to harvest the cones) and removal of trees for timber have led to losses in production capacity.


Elevation and pinecone production

The elevation of the pinyon pine is an important determinant of the quantity of pine cone production, and therefore, will largely determine the number of pine nuts the tree will yield. American pinyon pine cone production is most commonly found at an elevation between , and ideally at . This is due to higher temperatures at elevations lower than during the spring, which dry up humidity and moisture content (particularly snow packs) that provide for the tree throughout the spring and summer, causing little nourishment for pine cone maturity. Although there are several other environmental factors that determine the conditions of the ecosystem (such as clouds and rain), without sufficient water, the trees tend to abort cones. High humidity encourages cone development. There are certain topographical areas found in lower elevations, such as shaded canyons, where the humidity remains constant throughout the spring and summer, allowing pine cones to fully mature and produce seed. At elevations above , the temperature substantially drops, drastically affecting the state of the dormant cone. During the winter, frequent dramatic changes in temperature, along with drying, gusty winds, make the cones susceptible to
freeze-drying Freeze drying, also known as lyophilization or cryodesiccation, is a low temperature dehydration process that involves freezing the product and lowering pressure, removing the ice by sublimation. This is in contrast to dehydration by most conve ...
that damages them permanently; in this case, growth is stunted and the seeds deteriorate.


Physical characteristics

When first extracted from the pine cone, they are covered with a hard shell (seed coat), thin in some species, thick in others. The nutrition is stored in the embryo (
sporophyte A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote ...
) in the centre. Although a nut in the culinary sense, in the
botanical Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
sense pine nuts are seeds; being a
gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμν ...
, they lack a
carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils' ...
(fruit) outside. The shell must be removed before the pine nut can be eaten. Unshelled pine nuts have a long shelf life if kept dry and refrigerated (); shelled nuts (and unshelled nuts in warm conditions) deteriorate rapidly, becoming rancid within a few weeks or even days in warm humid conditions. Pine nuts are commercially available in shelled form, but due to poor storage, can have poor flavor and may be already rancid at the time of purchase. Consequently, pine nuts are often frozen to preserve their flavor. European pine nuts may be distinguished from Asian ones by their greater length in comparison to girth; Asian pine nuts are stubbier, shaped somewhat like long kernels of corn. The American piñon nuts are known for their large size and ease of shelling. In the United States, ''P. edulis'', the hard shell of New Mexico and Colorado, became a sought-after species due to the
trading post A trading post, trading station, or trading house, also known as a factory, is an establishment or settlement where goods and services could be traded. Typically the location of the trading post would allow people from one geographic area to tr ...
system, and the Navajo people who used the nuts as a means of commerce. The Italian pine nut (''P. pinea'') was brought to the United States by immigrants, and became a favored treat along the East Coast in the early 1930s, when bumper crops of American pine nuts were readily available at low prices.


Nutrition

When dried for eating, pine nuts are 2% water, 13%
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...
s, 14%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
, and 68% fat (table). In a reference serving, dried pine nuts supply of
food energy Food energy is chemical energy that animals (including humans) derive from their food to sustain their metabolism, including their muscular activity. Most animals derive most of their energy from aerobic respiration, namely combining the carbohy ...
and are a rich source (20% or more of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of healthy ...
, DV) of numerous
micronutrient Micronutrients are essential dietary elements required by organisms in varying quantities throughout life to orchestrate a range of physiological functions to maintain health. Micronutrient requirements differ between organisms; for example, huma ...
s, particularly
manganese Manganese is a chemical element with the Symbol (chemistry), symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese is a transition metal with a multifaceted array of ...
(419% DV),
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ea ...
(82% DV),
magnesium Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 of the periodic ...
(71% DV),
zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...
(67% DV), copper (65% DV),
vitamin E Vitamin E is a group of eight fat soluble compounds that include four tocopherols and four tocotrienols. Vitamin E deficiency, which is rare and usually due to an underlying problem with digesting dietary fat rather than from a diet low in vi ...
(62% DV),
vitamin K Vitamin K refers to structurally similar, fat-soluble vitamers found in foods and marketed as dietary supplements. The human body requires vitamin K for post-synthesis modification of certain proteins that are required for blood coagulation ...
(51% DV), and the
B vitamins B vitamins are a class of water-soluble vitamins that play important roles in cell metabolism and synthesis of red blood cells. Though these vitamins share similar names (B1, B2, B3, etc.), they are chemically distinct compounds that often coexi ...
, thiamin and
niacin Niacin, also known as nicotinic acid, is an organic compound and a form of vitamin B3, an essential human nutrient. It can be manufactured by plants and animals from the amino acid tryptophan. Niacin is obtained in the diet from a variet ...
(29–35% DV), among others (table).


Culinary uses

Pine nuts have been eaten in Europe and Asia since the
Paleolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός '' palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...
period. They are frequently added to meat, fish, salads and vegetable dishes or baked into bread. In Italian they are called ''pinoli'' (in the US they are often called ''pignoli'', but in Italy ''pignolo'' is actually a word far more commonly used to describe a fussy, overly fastidious or extremely meticulous person) and are an essential component of Italian pesto sauce; the upsurge in the popularity of this sauce since the 1990s has increased the visibility of the nut in America, primarily on the West Coast. ''Torta della nonna'' (literally "granny's cake") is a generic Italian dish name that in most families indicates an old family recipe for any kind of cake but often is used for a tart or a
pie A pie is a baked dish which is usually made of a pastry dough casing that contains a filling of various sweet or savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts (pecan pie), brown sugar ( sugar pie), sweete ...
filled with custard, topped with pine nuts and optionally dusted with
icing sugar Powdered sugar, also called confectioners' sugar, or icing sugar, is a finely ground sugar produced by milling granulated sugar into a powdered state. It usually contains between 2% and 5% of an anti-caking agent – such as corn starch, p ...
. Pignoli cookies, an Italian American specialty confection (in Italy these would be called ''biscotti ai pinoli''), are made of almond flour formed into a dough similar to that of a
macaroon A macaroon ( ) is a small cake or biscuit, typically made from ground almonds (the original main ingredient), coconut or other nuts (or even potato), with sugar and sometimes flavourings (e.g. honey, vanilla, spices), food colouring, glacé che ...
and then topped with pine nuts. In
Catalonia Catalonia (; ca, Catalunya ; Aranese Occitan: ''Catalonha'' ; es, Cataluña ) is an autonomous community of Spain, designated as a '' nationality'' by its Statute of Autonomy. Most of the territory (except the Val d'Aran) lies on the no ...
, a sweet is made of small
marzipan Marzipan is a confection consisting primarily of sugar, honey, and almond meal (ground almonds), sometimes augmented with almond oil or extract. It is often made into sweets; common uses are chocolate-covered marzipan and small marzipan imit ...
balls covered with pine nuts, painted with egg and lightly cooked, and those are called "Panellets". Pine nuts are also featured in the ''salade landaise'' of southwestern France. The Nevada, or Great Basin, pine nut has a sweet fruity flavor and is promoted for its large size, sweet flavor and ease of peeling. Pine nuts are also widely used in Middle Eastern cuisine, reflected in a diverse range of dishes such as '' kibbeh'', ''
sambusak A samosa () or singara is a fried Indian pastry with a savory filling, including ingredients such as spiced potatoes, onions, and peas. It may take different forms, including triangular, cone, or half-moon shapes, depending on the region. Sam ...
'', and '' fatayer'', desserts such as
baklava Baklava (, or ; ota, باقلوا ) is a layered pastry dessert made of filo pastry, filled with chopped nuts, and sweetened with syrup or honey. It was one of the most popular sweet pastries of Ottoman cuisine. The pre- Ottoman origin of t ...
, and many others. Throughout Europe and Middle East the pine nuts used are traditionally from '' Pinus pinea'' (stone pine). They are easily distinguished from the Asian pine nuts by their more slender shape and more homogeneous flesh. Because of the lower price, Asian pine nuts are also often used, especially in cheaper preparations. Pine nuts contain thiamine (vitamin B1) and protein. Pine nut coffee, known as ''piñón'' (Spanish for pine nut), is a specialty found in the southwest United States, especially
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
, and is typically a dark roast coffee having a deep, nutty flavor; roasted and lightly salted pine nuts can often be found sold on the side of the road in cities across New Mexico to be used for this purpose, as well as a snack. Pine nut oil is added to foods for flavor.


Taste disturbances

Some raw pine nuts can cause taste disturbances, lasting from a few days to a few weeks after consumption. A bitter, metallic, unpleasant taste is reported. There are no known lasting effects, with the United States
Food and Drug Administration The United States Food and Drug Administration (FDA or US FDA) is a federal agency of the Department of Health and Human Services. The FDA is responsible for protecting and promoting public health through the control and supervision of food ...
reporting that there are "no apparent adverse clinical side effects". Raw nuts from '' Pinus armandii'', mainly in China, may be the cause of the problem. Metallic taste disturbance is typically reported 1–3 days after ingestion, being worse on day two and typically lasting up to two weeks. Cases are self-limited and resolve without treatment.


Other uses

Pine nuts have long been a dietary staple in some Native American tribes. Today, though some tribes still use pine nuts in traditional cooking, others use the hard outer shell of the pine nut as a
bead A bead is a small, decorative object that is formed in a variety of shapes and sizes of a material such as stone, bone, shell, glass, plastic, wood, or pearl and with a small hole for threading or stringing. Beads range in size from under ...
for decorative purposes in traditional regalia and jewelry. In the
Great Basin The Great Basin is the largest area of contiguous endorheic watersheds, those with no outlets, in North America. It spans nearly all of Nevada, much of Utah, and portions of California, Idaho, Oregon, Wyoming, and Baja California. It is noted fo ...
area of the US, collecting pine nuts is a protected right through state law and treaty. In northern California, pine nuts are collected from the grey pine or bull pine. Tribes burn designs into the hard shell, reflecting the same design they use in baskets; however, they are often left blank, or burned to blacken. These are more often used in women's regalia and jewelry.


See also

* Jatjuk – a Korean porridge prepared using pine nuts *
Assidat Zgougou Assidat Zgougou ( aeb, عصيدة الزقوقو εasidet ezgougou) is a Tunisian dessert often prepared to celebrate the Mūled. It was originally made out of Aleppo pine, Flour, milk, cream and nuts, that one variant of the dish is now known ...
– a Tunisian dessert made of pine nuts *
List of edible seeds An edible seed is a seed that is suitable for human or animal consumption. Of the six major plant parts, seeds are the dominant source of human calories and protein. A wide variety of plant species provide edible seeds; most are angiosperms, while ...


References


Further reading

* *


External links

* {{DEFAULTSORT:Pine Nut Edible nuts and seeds Pinus Plants used in Native American cuisine