Parchment
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Parchment is a
writing material Writing material refers to the materials that provide the surfaces on which humans use writing instruments to inscribe writings. The same materials can also be used for symbolic or representational drawings. Building material on which writings or ...
made from specially prepared untanned skins of animals—primarily sheep, calves, and goats. It has been used as a writing medium for over two millennia. Vellum is a finer quality parchment made from the skins of young animals such as lambs and young calves. It may be called animal membrane by libraries and museums that wish to avoid distinguishing between ''parchment'' and the more-restricted term ''vellum'' (see below).


Parchment and vellum

Today the term ''parchment'' is often used in non-technical contexts to refer to any animal skin, particularly
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
,
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticated ...
or
cow Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
, that has been scraped or dried under tension. The term originally referred only to the skin of sheep and, occasionally, goats. The equivalent material made from calfskin, which was of finer quality, was known as '' vellum'' (from the
Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intellig ...
or , and ultimately from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
, meaning a calf); while the finest of all was ''uterine vellum'', taken from a calf
foetus A fetus or foetus (; plural fetuses, feti, foetuses, or foeti) is the unborn offspring that develops from an animal embryo. Following embryonic development the fetal stage of development takes place. In human prenatal development, fetal develo ...
or
stillborn Stillbirth is typically defined as fetal death at or after 20 or 28 weeks of pregnancy, depending on the source. It results in a baby born without signs of life. A stillbirth can result in the feeling of guilt or grief in the mother. The ter ...
calf. Some authorities have sought to observe these distinctions strictly: for example,
lexicographer Lexicography is the study of lexicons, and is divided into two separate academic disciplines. It is the art of compiling dictionaries. * Practical lexicography is the art or craft of compiling, writing and editing dictionaries. * Theoretica ...
Samuel Johnson in 1755, and master
calligrapher Calligraphy (from el, link=y, καλλιγραφία) is a visual art related to writing. It is the design and execution of lettering with a pen, ink brush, or other writing instrument. Contemporary calligraphic practice can be defined as "t ...
Edward Johnston Edward Johnston, CBE (11 February 1872 – 26 November 1944) was a British craftsman who is regarded, with Rudolf Koch, as the father of modern calligraphy, in the particular form of the broad-edged pen as a writing tool. He is most fa ...
in 1906. However, when old books and documents are encountered it may be difficult, without scientific analysis, to determine the precise animal origin of a skin either in terms of its species, or in terms of the animal's age. In practice, therefore, there has long been considerable blurring of the boundaries between the different terms. In 1519, William Horman wrote in his ''Vulgaria'': "That that we upon, and is made of , is called , , , ." In
Shakespeare William Shakespeare ( 26 April 1564 – 23 April 1616) was an English playwright, poet and actor. He is widely regarded as the greatest writer in the English language and the world's pre-eminent dramatist. He is often called England's natio ...
's ''
Hamlet ''The Tragedy of Hamlet, Prince of Denmark'', often shortened to ''Hamlet'' (), is a tragedy written by William Shakespeare sometime between 1599 and 1601. It is Shakespeare's longest play, with 29,551 words. Set in Denmark, the play depicts ...
'' (written c. 1599–1602) the following exchange occurs: Lee Ustick, writing in 1936, commented that: It is for these reasons that many modern
conservators In certain areas of England, Conservators are statutory bodies which manage areas of countryside for the use of the public. Establishment, Role and Powers Conservators are bodies corporate generally established, and granted their powers, by a ...
, librarians and archivists prefer to use either the broader term ''parchment'', or the neutral term ''animal membrane''.


History

The history of parchment is inseparable from the written word, once it had descended from clay tablets. A subsidiary use was found by scientists, most notably Thomas Graham, who employed parchment for the separation of
aqueous solutions An aqueous solution is a solution in which the solvent is water. It is mostly shown in chemical equations by appending (aq) to the relevant chemical formula. For example, a solution of table salt, or sodium chloride (NaCl), in water would be rep ...
he termed dialysis; in this respect parchment is seen to be akin to sausage casings made from the
intestines The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The GI tract contains all the major organs of the digestive system, in humans and ...
. The word parchment evolved (via the Latin and the French ) from the name of the city of
Pergamon Pergamon or Pergamum ( or ; grc-gre, Πέργαμον), also referred to by its modern Greek form Pergamos (), was a rich and powerful ancient Greek city in Mysia. It is located from the modern coastline of the Aegean Sea on a promontory on th ...
, which was a thriving center of parchment production during the
Hellenistic period In Classical antiquity, the Hellenistic period covers the time in Mediterranean history after Classical Greece, between the death of Alexander the Great in 323 BC and the emergence of the Roman Empire, as signified by the Battle of Actium in ...
. The city so dominated the trade that a legend later arose which said that parchment had been invented in
Pergamon Pergamon or Pergamum ( or ; grc-gre, Πέργαμον), also referred to by its modern Greek form Pergamos (), was a rich and powerful ancient Greek city in Mysia. It is located from the modern coastline of the Aegean Sea on a promontory on th ...
to replace the use of
papyrus Papyrus ( ) is a material similar to thick paper that was used in ancient times as a writing surface. It was made from the pith of the papyrus plant, '' Cyperus papyrus'', a wetland sedge. ''Papyrus'' (plural: ''papyri'') can also refer to a ...
which had become monopolized by the rival city of
Alexandria Alexandria ( or ; ar, ٱلْإِسْكَنْدَرِيَّةُ ; grc-gre, Αλεξάνδρεια, Alexándria) is the second largest city in Egypt, and the largest city on the Mediterranean coast. Founded in by Alexander the Great, Alexandri ...
. This account, originating in the writings of
Pliny the Elder Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic ' ...
('' Natural History'', Book XII, 69–70), is false because parchment had been in use in
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The ...
and elsewhere long before the rise of Pergamon.
Herodotus Herodotus ( ; grc, , }; BC) was an ancient Greek historian and geographer A geographer is a physical scientist, social scientist or humanist whose area of study is geography, the study of Earth's natural environment and human society ...
mentions writing on skins as common in his time, the 5th century BC; and in his ''Histories'' (v.58) he states that the Ionians of Asia Minor had been accustomed to give the name of ''skins'' () to books; this word was adapted by Hellenized Jews to describe scrolls. In the 2nd century BC, a great library was set up in Pergamon that rivaled the famous Library of Alexandria. As prices rose for papyrus and the reed used for making it was over-harvested towards local extinction in the two nomes of the Nile delta that produced it, Pergamon adapted by increasing use of parchment. Writing on prepared animal skins had a long history, however. David Diringer noted that "the first mention of Egyptian documents written on leather goes back to the
Fourth Dynasty The Fourth Dynasty of ancient Egypt (notated Dynasty IV) is characterized as a "golden age" of the Old Kingdom of Egypt. Dynasty IV lasted from to 2494 BC. It was a time of peace and prosperity as well as one during which trade with other ...
(c. 2550–2450 BC), but the earliest of such documents extant are: a fragmentary roll of leather of the
Sixth Dynasty The Sixth Dynasty of ancient Egypt (notated Dynasty VI), along with the Third, Fourth and Fifth Dynasty, constitutes the Old Kingdom of Dynastic Egypt. Pharaohs Known pharaohs of the Sixth Dynasty are listed in the table below. Manetho acc ...
(c. 24th century BC), unrolled by Dr. H. Ibscher, and preserved in the
Cairo Museum The Museum of Egyptian Antiquities, known commonly as the Egyptian Museum or the Cairo Museum, in Cairo, Egypt, is home to an extensive collection of ancient Egyptian antiquities. It has 120,000 items, with a representative amount on display ...
; a roll of the
Twelfth Dynasty The Twelfth Dynasty of ancient Egypt (Dynasty XII) is considered to be the apex of the Middle Kingdom by Egyptologists. It often is combined with the Eleventh, Thirteenth, and Fourteenth dynasties under the group title, Middle Kingdom. Some ...
(c. 1990–1777 BC) now in Berlin; the mathematical text now in the
British Museum The British Museum is a public museum dedicated to human history, art and culture located in the Bloomsbury area of London. Its permanent collection of eight million works is among the largest and most comprehensive in existence. It docum ...
(MS. 10250); and a document of the reign of
Ramses II Ramesses II ( egy, rꜥ-ms-sw ''Rīʿa-məsī-sū'', , meaning "Ra is the one who bore him"; ), commonly known as Ramesses the Great, was the third pharaoh of the Nineteenth Dynasty of Egypt. Along with Thutmose III he is often regarded as t ...
(early thirteenth century BC)." Though the
Assyria Assyria ( Neo-Assyrian cuneiform: , romanized: ''māt Aššur''; syc, ܐܬܘܪ, ʾāthor) was a major ancient Mesopotamian civilization which existed as a city-state at times controlling regional territories in the indigenous lands of the ...
ns and the
Babylonians Babylonia (; Akkadian: , ''māt Akkadī'') was an ancient Akkadian-speaking state and cultural area based in the city of Babylon in central-southern Mesopotamia (present-day Iraq and parts of Syria). It emerged as an Amorite-ruled state c. 1 ...
impressed their
cuneiform Cuneiform is a logo-syllabic script that was used to write several languages of the Ancient Middle East. The script was in active use from the early Bronze Age until the beginning of the Common Era. It is named for the characteristic wedge-sh ...
on clay tablets, they also wrote on parchment from the 6th century BC onward. Rabbinic literature traditionally maintains that the institution of employing parchment made of animal hides for the writing of ritual objects such as the
Torah The Torah (; hbo, ''Tōrā'', "Instruction", "Teaching" or "Law") is the compilation of the first five books of the Hebrew Bible, namely the books of Genesis, Exodus, Leviticus, Numbers and Deuteronomy. In that sense, Torah means the ...
,
mezuzah A ''mezuzah'' ( he, מְזוּזָה "doorpost"; plural: ''mezuzot'') is a piece of parchment, known as a '' klaf'', contained in a decorative case and inscribed with specific Hebrew verses from the Torah ( and ). These verses consist of the ...
, and tefillin is Sinaitic in origin, with special designations for different types of parchment such as gevil and
klaf Klaf or Qelaf ( he, קְלָף) is the designation given a particular piece of skin. The Talmudic definition includes both the form of the skin and the way it is processed, in particular, that it must be tanned. Since the innovative ruling of ' ...
. Early Islamic texts are also found on parchment. In the later
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
, especially the 15th century, parchment was largely replaced by
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distrib ...
for most uses except luxury manuscripts, some of which were also on paper. New techniques in paper milling allowed it to be much cheaper than parchment; it was made of textile rags and of very high quality. With the advent of
printing Printing is a process for mass reproducing text and images using a master form or template. The earliest non-paper products involving printing include cylinder seals and objects such as the Cyrus Cylinder and the Cylinders of Nabonidus. The ...
in the later fifteenth century, the demands of printers far exceeded the supply of animal skins for parchment. There was a short period during the introduction of printing where parchment and paper were used at the same time, with parchment (in fact vellum) the more expensive luxury option, preferred by rich and conservative customers. Although most copies of the
Gutenberg Bible The Gutenberg Bible (also known as the 42-line Bible, the Mazarin Bible or the B42) was the earliest major book printed using mass-produced movable metal type in Europe. It marked the start of the " Gutenberg Revolution" and the age of printed ...
are on paper, some were printed on parchment; 12 of the 48 surviving copies, with most incomplete. In 1490, Johannes Trithemius preferred the older methods, because "handwriting placed on parchment will be able to endure a thousand years. But how long will printing last, which is dependent on paper? For if ... it lasts for two hundred years that is a long time." In fact high quality paper from this period has survived 500 years or more very well, if kept in reasonable library conditions. The heyday of parchment use was during the medieval period, but there has been a growing revival of its use among artists since the late 20th century. Although parchment never stopped being used (primarily for governmental documents and diplomas) it had ceased to be a primary choice for artist's supports by the end of 15th century
Renaissance The Renaissance ( , ) , from , with the same meanings. is a period in European history The history of Europe is traditionally divided into four time periods: prehistoric Europe (prior to about 800 BC), classical antiquity (800 BC to AD ...
. This was partly due to its expense and partly due to its unusual working properties. Parchment consists mostly of collagen. When the water in paint media touches parchment's surface, the collagen melts slightly, forming a raised bed for the paint, a quality highly prized by some artists. Parchment is also extremely affected by its environment and changes in humidity, which can cause buckling. Books with parchment pages were bound with strong wooden boards and clamped tightly shut by metal (often brass) clasps or leather straps; this acted to keep the pages pressed flat despite humidity changes. Such metal fittings continued to be found on books as decorative features even after the use of paper made them unnecessary. Some contemporary artists prize the changeability of parchment, noting that the material seems alive and like an active participant in making artwork. To support the needs of the revival of use by artists, a revival in the art of preparing individual skins is also underway. Hand-prepared skins are usually preferred by artists because they are more uniform in surface and have fewer oily spots – which can cause long-term cracking of paint – than mass-produced parchment, which is usually made for lamp shades, furniture, or other interior design purposes. The radiocarbon dating techniques that are used on papyrus can be applied to parchment as well. They do not date the age of the writing but the preparation of the parchment itself. While it is feasibly possible also to radiocarbon date certain kinds of ink, it is extremely difficult to do due to the fact that they are generally present on the text only in trace amounts, and it is hard to get a carbon sample of them without the carbon in the parchment contaminating it.


Manufacture

Parchment is prepared from pelt – i.e. wet, unhaired, and limed skin – by drying at ordinary temperatures under tension, most commonly on a wooden frame known as a stretching frame.


Skinning, soaking, and dehairing

After a carcass is skinned, the hide is soaked in water for about a day. This removes blood and grime and prepares the skin for a dehairing liquor. The dehairing liquor was originally made of rotted, or fermented, vegetable matter, like beer or other liquors, but by the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
a dehairing bath included
lime Lime commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a color between yellow and green Lime may also refer to: Botany ...
. Today, the lime solution is occasionally sharpened by the use of sodium sulfide. The liquor bath would have been in wooden or stone vats and the hides stirred with a long wooden pole to avoid human contact with the alkaline solution. Sometimes the skins would stay in the dehairing bath for eight or more days depending how concentrated and how warm the solution was kept – dehairing could take up to twice as long in winter. The vat was stirred two or three times a day to ensure the solution's deep and uniform penetration. Replacing the lime water bath also sped the process up. However, if the skins were soaked in the liquor too long, they would be weakened and not able to stand the stretching required for parchment.


Stretching

After soaking in water to make the skins workable, the skins were placed on a stretching frame. A simple frame with nails would work well in stretching the pelts. The skins could be attached by wrapping small, smooth rocks in the skins with rope or leather strips. Both sides would be left open to the air so they could be scraped with a sharp, semi-lunar knife to remove the last of the hair and get the skin to the right thickness. The skins, which were made almost entirely of collagen, would form a natural glue while drying and once taken off the frame they would keep their form. The stretching aligned the fibres to be more nearly parallel to the surface.


Treatments

To make the parchment more aesthetically pleasing or more suitable for the scribes, special treatments were used. According to Reed there were a variety of these treatments. Rubbing pumice powder into the flesh side of parchment while it was still wet on the frame was used to make it smooth and to modify the surface to enable inks to penetrate more deeply. Powders and pastes of calcium compounds were also used to help remove grease so the ink would not run. To make the parchment smooth and white, thin pastes (starchgrain or staunchgrain) of lime, flour, egg whites and milk were rubbed into the skins. Meliora di Curci in her paper "The History and Technology of Parchment Making" notes that parchment was not always white. " Cennini, a 15th-century craftsman provides recipes to tint parchment a variety of colours including purple, indigo, green, red and peach." The Early medieval
Codex Argenteus The Codex Argenteus (Latin for "Silver Book/Codex") is a 6th-century illuminated manuscript, originally containing part of the 4th-century translation of the Christian Bible into the Gothic language. Traditionally ascribed to the Arian bi ...
and Codex Vercellensis, the Stockholm Codex Aureus and the
Codex Brixianus The Codex Brixianus (Brescia, Biblioteca Civica Queriniana, s.n.), designated by f, is a 6th-century Latin Gospel Book which was probably produced in Italy. Description The manuscript contains 419 folios. The text, written on purple dyed vellum ...
give a range of luxuriously produced manuscripts all on purple vellum, in imitation of Byzantine examples, like the Rossano Gospels, Sinope Gospels and the Vienna Genesis, which at least at one time are believed to have been reserved for Imperial commissions. Many techniques for parchment repair exist, to restore creased, torn, or incomplete parchments.


Reuse

Between the seventh and the ninth centuries, many earlier parchment manuscripts were scrubbed and scoured to be ready for rewriting, and often the earlier writing can still be read. These recycled parchments are known as
palimpsest In textual studies, a palimpsest () is a manuscript page, either from a scroll or a book, from which the text has been scraped or washed off so that the page can be reused for another document. Parchment was made of lamb, calf, or kid skin an ...
s. Later, more thorough techniques of scouring the surface irretrievably lost the earlier text.


Jewish parchment

The way in which parchment was processed (from hide to parchment) has undergone a tremendous evolution based on time and location. Parchment and vellum are not the sole methods of preparing animal skins for writing. In the Babylonian Talmud (
Bava Batra Bava Batra (also Baba Batra; Talmudic Aramaic: בָּבָא בַּתְרָא "The Last Gate") is the third of the three Talmudic tractates in the Talmud in the order Nezikin; it deals with a person's responsibilities and rights as the owner of pr ...
14B), Moses is described as having written the first Torah Scroll on the unsplit cow-hide called '' gevil''. Parchment is still the only medium used by traditional
religious Religion is usually defined as a social- cultural system of designated behaviors and practices, morals, beliefs, worldviews, texts, sanctified places, prophecies, ethics, or organizations, that generally relates humanity to supernatur ...
Jews Jews ( he, יְהוּדִים, , ) or Jewish people are an ethnoreligious group and nation originating from the Israelites Israelite origins and kingdom: "The first act in the long drama of Jewish history is the age of the Israelites""The ...
for Torah scrolls or tefilin and
mezuzah A ''mezuzah'' ( he, מְזוּזָה "doorpost"; plural: ''mezuzot'') is a piece of parchment, known as a '' klaf'', contained in a decorative case and inscribed with specific Hebrew verses from the Torah ( and ). These verses consist of the ...
s, and is produced by large companies in
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
. For those uses, only hides of kosher animals are permitted. Since there are many requirements for it being fit for the religious use, the liming is usually processed under supervision of a qualified
Rabbi A rabbi () is a spiritual leader or religious teacher in Judaism. One becomes a rabbi by being ordained by another rabbi – known as ''semikha'' – following a course of study of Jewish history and texts such as the Talmud. The basic form of ...
.


Additional uses of the term

In some universities, the word parchment is still used to refer to the certificate (scroll) presented at graduation ceremonies, even though the modern document is printed on paper or thin card; although doctoral graduates may be given the option of having their scroll written by a calligrapher on vellum. Heriot-Watt University still uses goatskin parchment for their degrees.


Plant-based parchment

Vegetable (paper) parchment is made by passing a waterleaf (an unsized paper like blotters) made of pulp fibers into sulfuric acid. The sulfuric acid hydrolyses and solubilises the main natural organic polymer, cellulose, present in the pulp wood fibers. The paper web is then washed in water, which stops the hydrolysis of the cellulose and causes a kind of cellulose coating to form on the waterleaf. The final paper is dried. This coating is a natural non-porous cement, that gives to the vegetable parchment paper its resistance to grease and its semi-translucency. Other processes can be used to obtain grease-resistant paper, such as waxing the paper or using fluorine-based chemicals. Highly beating the fibers gives an even more translucent paper with the same grease resistance. Silicone and other coatings may also be applied to the parchment. A silicone-coating treatment produces a cross-linked material with high density, stability and heat resistance and low surface tension which imparts good anti-stick or release properties. Chromium salts can also be used to impart moderate anti-stick properties.


Parchment craft

Historians believe that parchment craft originated as an art form in Europe during the fifteenth or sixteenth centuries. Parchment craft at that time occurred principally in Catholic communities, where crafts persons created lace-like items such as devotional pictures and communion cards. The craft developed over time, with new techniques and refinements being added. Until the sixteenth century, parchment craft was a European art form. However, missionaries and other settlers relocated to South America, taking parchment craft with them. As before, the craft appeared largely among the Catholic communities. Often, young girls receiving their first communion received gifts of handmade parchment crafts. Although the invention of the printing press led to a reduced interest in hand made cards and items, by the eighteenth century, people were regaining interest in detailed handwork. Parchment cards became larger in size and crafters began adding wavy borders and perforations. In the nineteenth century, influenced by French romanticism, parchment crafters began adding floral themes and cherubs and hand embossing. Parchment craft today involves various techniques, including tracing a pattern with white or colored ink, embossing to create a raised effect, stippling, perforating, coloring and cutting. Parchment craft appears in hand made cards, as scrapbook embellishments, as bookmarks, lampshades, decorative small boxes, wall hangings and more.


DNA testing

An article published in 2009 by Timothy L. Stinson considered the possibilities of tracing the origin of medieval parchment manuscripts and codices through DNA analysis. The methodology would employ
polymerase chain reaction The polymerase chain reaction (PCR) is a method widely used to rapidly make millions to billions of copies (complete or partial) of a specific DNA sample, allowing scientists to take a very small sample of DNA and amplify it (or a part of it) ...
to replicate a small DNA sample to a size sufficiently large for testing. Stinson's article discusses the use of DNA testing to estimate the age of the calf at the creation of the vellum parchment. A 2006 study revealed the genetic signature of several Greek manuscripts to have "goat-related sequences". Utilizing these techniques we may be able to determine whether related library materials were made from genetically related animals (perhaps from the same herd) and locate the vellum's origination.Stinson, Timothy L. "Knowledge of the Flesh: Using DNA Analysis to Unlock Bibliographical Secrets of Medieval Parchment" The Papers of the Bibliographical Society. Volume 103, Number 4 2009 Dec pp:435-5

/ref> In 2020, it was reported that the species of several of the animals used to provide parchment for the
Dead Sea Scrolls The Dead Sea Scrolls (also the Qumran Caves Scrolls) are ancient Jewish and Hebrew religious manuscripts discovered between 1946 and 1956 at the Qumran Caves in what was then Mandatory Palestine, near Ein Feshkha in the West Bank, on the ...
could be identified, and the relationship between skins obtained from the same animal inferred. The breakthrough was made possible by the use of whole genome sequencing.


See also

* Conservation and restoration of parchment * Manuscript culture


References


Notes


Bibliography

* * * * * * *


Further reading

* * * * * * * *


External links

*
Preservation of 18th Century Parchment , "From the Stacks" at New-York Historical Society


, Bibliothèque nationale de France – Text in French, but mostly visual {{Authority control Leather goods Book design Writing media Textual scholarship