The tiger (''Panthera tigris'') is the largest living

cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

and a member of the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

'' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An

apex predator An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic lev ...

, it primarily preys on

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ...

s, such as

deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...

and

wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...

. It is territorial and generally a

solitary but social Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...

predator, requiring large contiguous areas of

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

to support its requirements for

prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

and rearing of its offspring. Tiger cubs stay with their mother for about two years and then become independent, leaving their mother's

home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. He ...

to establish their own. The tiger was first

scientifically described A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...

in 1758. It once ranged widely from the

Eastern Anatolia Region The Eastern Anatolia Region ('' tr, Doğu Anadolu Bölgesi'') is a geographical region of Turkey. The most populous province in the region is Van Province. Other populous provinces are Malatya, Erzurum and Elazığ. It is bordered by the Black S ...

in the west to the Amur River basin in the east, and in the south from the foothills of the

Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...

to Bali in the Sunda Islands. Since the early 20th century, tiger populations have lost at least 93% of their historic range and have been

extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...

from

Western Western may refer to: Places *Western, Nebraska, a village in the US *Western, New York, a town in the US *Western Creek, Tasmania, a locality in Australia *Western Junction, Tasmania, a locality in Australia *Western world, countries that id ...

and

Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...

, the islands of

Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mos ...

and Bali, and in large areas of

Southeast The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...

and

South Asia South Asia is the southern subregion of Asia, which is defined in both geographical Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth descr ...

and China. What remains of the range where tigers still roam free is fragmented, stretching in spots from

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...

temperate forest A temperate forest is a forest found between the tropical and boreal regions, located in the temperate zone. It is the second largest biome on our planet, covering 25% of the world's forest area, only behind the boreal forest, which covers abou ...

s to subtropical and

tropical forest Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds. Some tropical fore ...

s on the

Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...

Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...

and a single Indonesian island, Sumatra. The tiger is listed as

Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...

on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...

. As of 2015, the global wild tiger population was estimated to number between 3,062 and 3,948 mature individuals, with most populations living in small isolated pockets.

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

currently hosts the largest tiger population. Major reasons for population decline are

habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

, habitat fragmentation and

poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...

. Tigers are also victims of

human–wildlife conflict Human–wildlife conflict (HWC) refers to the negative interactions between human and wild animals, with undesirable consequences both for people and their resources, on the one hand, and wildlife and their habitats on the other ( IUCN 2020). HW ...

, due to encroachment in countries with a high human population density. The tiger is among the most recognisable and popular of the world's

charismatic megafauna Charismatic megafauna are animal species that are large—in the relevant category that they represent—with symbolic value or widespread popular appeal, and are often used by environmental activists to gain public support for environmentalist go ...

. It featured prominently in the ancient

mythology Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narra ...

and

folklore Folklore is shared by a particular group of people; it encompasses the traditions common to that culture, subculture or group. This includes oral traditions such as tales, legends, proverbs and jokes. They include material culture, ranging ...

of cultures throughout its historic range and continues to be depicted in modern films and literature, appearing on many flags, coats of arms and as

mascot A mascot is any human, animal, or object thought to bring luck, or anything used to represent a group with a common public identity, such as a school, professional sports team, society, military unit, or brand name. Mascots are also used as fi ...

s for sporting teams. The tiger is the

national animal This is a list of countries that have officially designated one or more animals as their national animals. National animal {, class="wikitable sortable" ! Country ! Name of animal ! Scientific name (Latin name) ! class="unsortable", Picture ...

Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

and

South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korean Peninsula and sharing a land border with North Korea. Its western border is formed by the Yellow Sea, while its eas ...

Etymology

The

Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English ...

''tigre'' and

Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...

''tigras'' derive from Old French ''tigre'', from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''tigris''. This was a borrowing of

Classical Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...

τίγρις 'tigris', a foreign borrowing of unknown origin meaning 'tiger' and the river

Tigris The Tigris () is the easternmost of the two great rivers that define Mesopotamia, the other being the Euphrates. The river flows south from the mountains of the Armenian Highlands through the Syrian and Arabian Deserts, and empties into the ...

. The origin may have been the

Persian Persian may refer to: * People and things from Iran, historically called ''Persia'' in the English language ** Persians, the majority ethnic group in Iran, not to be conflated with the Iranic peoples ** Persian language, an Iranian language of the ...

word ''tigra'' ('pointed or sharp') and the Avestan word ''tigrhi'' ('arrow'), perhaps referring to the speed of the tiger's leap, although these words are not known to have any meanings associated with tigers. The generic name ''Panthera'' is derived from the

word ''panthera'' and the

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...

word πάνθηρ ('panther').

Taxonomy

In 1758,

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...

described the tiger in his work '' Systema Naturae'' and gave it the scientific name ''Felis tigris''. In 1929, the British taxonomist Reginald Innes Pocock subordinated the species under the genus '' Panthera'' using the scientific name ''Panthera tigris''.

Subspecies

Following Linnaeus's first descriptions of the species, several tiger specimens were described and proposed as subspecies. The validity of several tiger subspecies was questioned in 1999. Most putative subspecies described in the 19th and 20th centuries were distinguished on basis of fur length and colouration, striping patterns and body size, hence characteristics that vary widely within populations. Morphologically, tigers from different regions vary little, and gene flow between populations in those regions is considered to have been possible during the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...

. Therefore, it was proposed to recognize only two tiger subspecies as valid, namely '' P. t. tigris'' in mainland Asia, and '' P. t. sondaica'' in the

Greater Sunda Islands The Greater Sunda Islands ( Indonesian and Malay: ''Kepulauan Sunda Besar'') are four tropical islands situated within Indonesian Archipelago, in the Pacific Ocean. The islands, Borneo, Java, Sulawesi and Sumatra, are internationally recognised ...

. Results of craniological analysis of 111 tiger skulls from Southeast Asian range countries indicate that Sumatran tiger skulls differ from Indochinese and Javan tiger skulls, whereas Bali tiger skulls are similar in size to Javan tiger skulls. The authors proposed to classify the Sumatran and Javan tigers as distinct species, ''P. sumatrae'' and ''P. sondaica'', with the Bali tiger as subspecies ''P. sondaica balica''. In 2015, morphological, ecological, and molecular traits of all putative tiger subspecies were analysed in a combined approach. Results support distinction of the two evolutionary groups continental and Sunda tigers. The authors proposed recognition of only two subspecies, namely ''P. t. tigris'' comprising the Bengal, Malayan, Indochinese, South Chinese, Siberian and Caspian tiger populations, and ''P. t. sondaica'' comprising the Javan, Bali and Sumatran tiger populations. The authors also noted that this reclassification will affect tiger conservation management. The nominate subspecies ''P. t. tigris'' constitutes two clades: * a northern clade composed of the Siberian and Caspian tiger populations * a southern clade composed of all other mainland populations. One conservation specialist welcomed this proposal as it would make captive breeding programmes and future rewilding of zoo-born tigers easier. One

geneticist A geneticist is a biologist or physician who studies genetics, the science of genes, heredity, and variation of organisms. A geneticist can be employed as a scientist or a lecturer. Geneticists may perform general research on genetic processes ...

was sceptical of this study and maintained that the currently recognised nine subspecies can be distinguished genetically. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group revised felid taxonomy and recognized the tiger populations in continental Asia as ''P. t. tigris'', and those in the Sunda Islands as ''P. t. sondaica''. This two-subspecies view has been largely rejected by researchers. Results of a 2018

whole-genome sequencing Whole genome sequencing (WGS), also known as full genome sequencing, complete genome sequencing, or entire genome sequencing, is the process of determining the entirety, or nearly the entirety, of the DNA sequence of an organism's genome at a ...

of 32 specimens support six monophyletic tiger clades corresponding with the living subspecies and indicate that the most recent

common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. All living beings are in fact descendants of a unique ancestor commonly referred to as the last universal comm ...

lived about 110,000 years ago. The following tables are based on the classification of the species ''Panthera tigris'' provided in ''

Mammal Species of the World ''Mammal Species of the World: A Taxonomic and Geographic Reference'' is a standard reference work in mammalogy giving descriptions and bibliographic data for the known species of mammals. It is now in its third edition, published in late 2005, ...

''. It also reflects the classification used by the Cat Classification Task Force in 2017:

Evolution

The tiger's closest living relatives were previously thought to be the ''Panthera'' species lion, leopard and jaguar. Results of

genetic analysis Genetic analysis is the overall process of studying and researching in fields of science that involve genetics and molecular biology. There are a number of applications that are developed from this research, and these are also considered parts of ...

indicate that about 2.88 million years ago, the tiger and the

snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...

lineages diverged from the other ''Panthera'' species, and that both may be more closely related to each other than to the lion, leopard and jaguar. The geographic origin of the ''Panthera'' is most likely northern Central Asia. The tiger–snow leopard lineage dispersed in Southeast Asia during the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

. ''

Panthera zdanskyi ''Panthera zdanskyi'', also known as the Longdan tiger, is an extinct pantherine species that is seen as a close relative of the modern tiger. Fossils were excavated in northwestern China's Gansu province. Etymology ''Panthera zdanskyi'' was fi ...

'' is considered to be a

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of the modern tiger. It lived at the beginning of the

about two million years ago, its

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

remains were excavated in Gansu of northwestern China. It was smaller and more " primitive", but functionally and ecologically similar to the modern tiger. It is disputed as to whether it had the striping pattern. Northwestern China is thought to be the origin of the tiger lineage. Tigers grew in size, possibly in response to

adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...

s of prey species like

and

bovid The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, ...

s, which may have occurred in Southeast Asia during the Early Pleistocene. ''

Panthera tigris trinilensis ''Panthera tigris trinilensis'', known as the Trinil tiger, is an extinct tiger subspecies dating from about 1.2 million years ago that was found at the locality of Trinil, Java, Indonesia. The fossil remains are now stored in the Dubois Colle ...

'' lived about and is known from fossils excavated near Trinil in

. The Wanhsien, Ngandong, Trinil, and Japanese tigers became extinct in prehistoric times. Tigers reached India and northern Asia in the late Pleistocene, reaching eastern Beringia, Japan, and

Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh ...

. Some fossil skulls are morphologically distinct from lion skulls, which could indicate tiger presence in Alaska during the last glacial period, about 100,000 years ago. Fossil teeth and bones found in Borneo were attributed to the Bornean tiger and date to about 13,745 to 3,000 years ago. It may have accessed Borneo, when the sea level was low during a glaciation period, and may have survived until about 200 years ago. In the

Ille Cave Dewil Valley, located in the northernmost part of Palawan, an island province of the Philippines that is located in the Mimaropa region, is an archaeological site composed of thousands of artifacts and features. According to the University of the ...

on the island of Palawan, two articulated

phalanx bone The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bon ...

s were found amidst an assemblage of other animal bones and stone tools. They were smaller than mainland tiger fossils, possibly due to

insular dwarfism Insular dwarfism, a form of phyletic dwarfism, is the process and condition of large animals evolving or having a reduced body size when their population's range is limited to a small environment, primarily islands. This natural process is disti ...

. It has been speculated that the tiger parts were either imported from elsewhere, or that the tiger colonised Palawan from Borneo before the Holocene. Fossil remains of tigers were also excavated in Sri Lanka, China, Japan and

Sarawak Sarawak (; ) is a state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the Malaysian state of Sabah to the northeast, ...

dating to the

Late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...

and Early

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...

. Results of a

phylogeographic Phylogeography is the study of the historical processes that may be responsible for the past to present geographic distributions of genealogical lineages. This is accomplished by considering the geographic distribution of individuals in light of ge ...

study indicate that all living tigers had a common ancestor 108,000 to 72,000 years ago. The potential tiger range during the late Pleistocene and Holocene was predicted applying

ecological niche modelling Species distribution modelling (SDM), also known as environmental (or ecological) niche modelling (ENM), habitat modelling, predictive habitat distribution modelling, and range mapping uses computer algorithms to predict the distribution of a sp ...

based on more than 500 tiger locality records combined with

bioclimatic Green building (also known as green construction or sustainable building) refers to both a structure and the application of processes that are environmentally responsible and resource-efficient throughout a building's life-cycle: from plannin ...

data. The resulting model shows a contiguous tiger range at the Last Glacial Maximum, indicating gene flow between tiger populations in mainland Asia. The Caspian tiger population was likely connected to the Bengal tiger population through corridors below elevations of in the

Hindu Kush The Hindu Kush is an mountain range in Central and South Asia to the west of the Himalayas. It stretches from central and western Afghanistan, Quote: "The Hindu Kush mountains run along the Afghan border with the North-West Frontier Province ...

. The tiger populations on the Sunda Islands and mainland Asia were possibly separated during

interglacial period An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene in ...

s. The tiger's full

genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding g ...

sequence was published in 2013. It was found to have similar repeat composition to other cat genomes and an appreciably conserved

synteny In genetics, the term synteny refers to two related concepts: * In classical genetics, ''synteny'' describes the physical co-localization of genetic loci on the same chromosome within an individual or species. * In current biology, ''synteny'' mo ...

Hybrids

Captive tigers were bred with lions to create hybrids called liger and tigon. They share physical and behavioural qualities of both parent species. Breeding hybrids is now discouraged due to the emphasis on conservation. The liger is a cross between a male lion and a tigress. Ligers are typically between in length, and weigh between or more. Because the lion sire passes on a growth-promoting gene, but the corresponding growth-inhibiting gene from the female tiger is absent, ligers grow far larger than either parent species. The less common tigon is a cross between a lioness and a male tiger. Because the male tiger does not pass on a growth-promoting gene and the lioness passes on a growth inhibiting gene, tigons are around the same size as their parents. Some females are fertile and have occasionally given birth to

litigon A litigon (/ˌlaɪˈtaɪɡən/) is a rare, second-generation hybrid from a female tigon (a hybrid between a male tiger and a female lion) and a male lion, specifically an Asiatic lion. Description Litigons inherit and share characteristics wit ...

s when mated to a male

Asiatic lion The Asiatic lion is a population of '' Panthera leo leo'' that today survives in the wild only in India. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of Gujar ...

Description

The tiger has a muscular body with strong forelimbs, a large head and a tail that is about half the length of its body. Its

pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...

colouration varies between shades of orange with a white underside and distinctive vertical black stripes; the patterns of which are unique in each individual. Stripes are likely advantageous for

camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...

in vegetation such as long grass with strong vertical patterns of light and shade. The tiger is one of only a few striped cat species; it is not known why spotted patterns and rosettes are the more common camouflage pattern among felids. The orange colour may also aid in camouflage as the tiger's prey are

dichromat Dichromacy (from Greek ''di'', meaning "two" and ''chromo'', meaning "color") is the state of having two types of functioning photoreceptors, called cone cells, in the eyes. Organisms with dichromacy are called dichromats. Dichromats requir ...

s, and thus may perceive the cat as green and blended in with the vegetation. A tiger's coat pattern is still visible when it is shaved. This is not due to skin pigmentation, but to the stubble and hair follicles embedded in the skin. It has a mane-like heavy growth of fur around the neck and jaws and long whiskers, especially in males. The pupils are circular with yellow irises. The small, rounded ears have a prominent white spot on the back, surrounded by black. These spots are thought to play an important role in intraspecific communication. The tiger's

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...

is similar to a lion's skull, with the frontal region usually less depressed or flattened, and a slightly longer

postorbital The ''postorbital'' is one of the bones in vertebrate skulls which forms a portion of the dermal skull roof and, sometimes, a ring about the orbit. Generally, it is located behind the postfrontal and posteriorly to the orbital fenestra. In some ...

region. The lion skull shows broader nasal openings. Due to the variation in skull sizes of the two species, the structure of the lower jaw is a reliable indicator for their identification. The tiger has fairly stout teeth; its somewhat curved

canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the surn ...

are the longest among living felids with a

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...

height of up to .

Size

There is notable

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...

between male and female tigers, with the latter being consistently smaller. The size difference between them is proportionally greater in the large tiger subspecies, with males weighing up to 1.7 times more than females. Males also have wider forepaw pads, enabling sex to be identified from tracks. It has been

hypothesis A hypothesis (plural hypotheses) is a proposed explanation for a phenomenon. For a hypothesis to be a scientific hypothesis, the scientific method requires that one can test it. Scientists generally base scientific hypotheses on previous obse ...

ed that body size of different tiger populations may be correlated with climate and be explained by

thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...

and Bergmann's rule, or by distribution and size of available prey species. Generally, males vary in total length from and weigh between with skull length ranging from . Females vary in total length from , weigh with skull length ranging from . In either sex, the tail represents about of the total length. The Bengal and Siberian tigers are amongst the tallest cats in shoulder height. They are also ranked among the biggest cats that have ever existed reaching weights of more than . The tigers of the Sunda islands are smaller and less heavy than tigers in mainland Asia, rarely exceeding in weight.

Colour variations

There are three other colour variants – white, golden and nearly stripeless snow white – that are now virtually non-existent in the wild due to the reduction of wild tiger populations, but continue in captive populations. The

white tiger The white tiger or bleached tiger is a leucistic pigmentation variant of the Mainland tiger. It is reported in the wild from time to time in the Indian states of Madhya Pradesh, Assam, West Bengal, Bihar, Odisha, in the Sunderbans region and ...

has white fur and

sepia Sepia may refer to: Biology * ''Sepia'' (genus), a genus of cuttlefish Color * Sepia (color), a reddish-brown color * Sepia tone, a photography technique Music * ''Sepia'', a 2001 album by Coco Mbassi * ''Sepia'' (album) by Yu Takahashi * " ...

-brown stripes. The

golden tiger A golden tiger, sometimes called a golden tabby tiger, is a tiger with a colour variation caused by a recessive gene. Like white tigers and black tigers The Black Tigers () was an elite suicide commando unit of the Liberation Tigers of Tam ...

has a pale golden pelage with a blond tone and reddish-brown stripes. The snow white tiger is a morph with extremely faint stripes and a pale reddish-brown ringed tail. Both snow white and golden tigers are homozygous for

CORIN Corin is a given name in English deriving from the Latin Quirinus, a Roman god. The meaning is unclear but is probably associated with "spear". The name is that of a character in William Shakespeare's ''As You Like It''. It is also used as a fami ...

gene mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, mitos ...

s. The white tiger lacks

pheomelanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the a ...

(which creates the orange colour), and has dark sepia-brown stripes and blue eyes. This altered pigmentation is caused by a

mutant In biology, and especially in genetics, a mutant is an organism or a new genetic character arising or resulting from an instance of mutation, which is generally an alteration of the DNA sequence of the genome or chromosome of an organism. It ...

gene that is inherited as an autosomal recessive trait, which is determined by a white

locus Locus (plural loci) is Latin for "place". It may refer to: Entertainment * Locus (comics), a Marvel Comics mutant villainess, a member of the Mutant Liberation Front * ''Locus'' (magazine), science fiction and fantasy magazine ** ''Locus Award' ...

. It is not an

albino Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...

, as the dark pigments are scarcely affected. The mutation changes a single

amino acid Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although hundreds of amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha a ...

in the transporter protein SLC45A2. Both parents need to have the

allele An allele (, ; ; modern formation from Greek ἄλλος ''állos'', "other") is a variation of the same sequence of nucleotides at the same place on a long DNA molecule, as described in leading textbooks on genetics and evolution. ::"The chro ...

for whiteness to have white cubs. Between the early and mid 20th century, white tigers were recorded and shot in the Indian states of

Odisha Odisha (English: , ), formerly Orissa ( the official name until 2011), is an Indian state located in Eastern India. It is the 8th largest state by area, and the 11th largest by population. The state has the third largest population of ...

, Bihar,

Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...

and in the area of

Rewa, Madhya Pradesh Rewa is a city in north-eastern part of Madhya Pradesh state in India. It is the administrative center of Rewa District and Rewa Division. The city lies about northeast of the state capital Bhopal and north of the city of Jabalpur. The ma ...

. The local maharaja started breeding tigers in the early 1950s and kept a white male tiger together with its normal-coloured daughter; they had white cubs. To preserve this recessive trait, only a few white individuals were used in captive breeding, which led to a high degree of

inbreeding Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders and o ...

Inbreeding depression Inbreeding depression is the reduced biological fitness which has the potential to result from inbreeding (the breeding of related individuals). Biological fitness refers to an organism's ability to survive and perpetuate its genetic material. ...

is the main reason for many health problems of captive white tigers, including

strabismus Strabismus is a vision disorder in which the eyes do not properly align with each other when looking at an object. The eye that is focused on an object can alternate. The condition may be present occasionally or constantly. If present during a ...

, stillbirth,

deformities A deformity, dysmorphism, or dysmorphic feature is a major abnormality of an organism that makes a part of the body appear or function differently than how it is supposed to. Causes Deformity can be caused by a variety of factors: *Arthritis an ...

and premature death. Other physical defects include

cleft palate A cleft lip contains an opening in the upper lip that may extend into the nose. The opening may be on one side, both sides, or in the middle. A cleft palate occurs when the palate (the roof of the mouth) contains an opening into the nose. The ...

and scoliosis. The Tiger

Species Survival Plan The American Species Survival Plan or SSP program was developed in 1981 by the (American) Association of Zoos and Aquariums to help ensure the survival of selected species in zoos and aquariums, most of which are threatened or endangered in the ...

has condemned the breeding of white tigers, alleging they are of mixed ancestry and of unknown lineage. The

gene In biology, the word gene (from , ; "... Wilhelm Johannsen coined the word gene to describe the Mendelian units of heredity..." meaning ''generation'' or ''birth'' or ''gender'') can have several different meanings. The Mendelian gene is a b ...

s responsible for white colouration are represented by 0.001% of the population. The disproportionate growth in numbers of white tigers points to

among

homozygous Zygosity (the noun, zygote, is from the Greek "yoked," from "yoke") () is the degree to which both copies of a chromosome or gene have the same genetic sequence. In other words, it is the degree of similarity of the alleles in an organism. Mo ...

recessive In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and t ...

individuals. This would lead to

inbreeding depression Inbreeding depression is the reduced biological fitness which has the potential to result from inbreeding (the breeding of related individuals). Biological fitness refers to an organism's ability to survive and perpetuate its genetic material. ...

and loss of

genetic variability Genetic variability is either the presence of, or the generation of, genetic differences. It is defined as "the formation of individuals differing in genotype, or the presence of genotypically different individuals, in contrast to environmentally i ...

. There are also records of pseudomelanic or

black tiger A black tiger is a rare colour variant of the tiger, and is not a distinct species or geographic subspecies. Description There are reports and one painting (now lost) of pure black non-striped tigers (true melanistic tigers). Most black mammal ...

s which have thick stripes that merge. In

Simlipal National Park Simlipal is a tiger reserve in the Mayurbhanj district in the Indian state of Odisha covering . It is part of the Mayurbhanj Elephant Reserve, which includes three protected areas — Similipal Tiger Reserve, Hadgarh Wildlife Sanctuary with a ...

, 37% of the tiger population has this condition, which has been linked to isolation and inbreeding.

Distribution and habitat

The tiger historically ranged from eastern

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula in ...

and

Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...

to the coast of the

Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, i ...

, and from South Asia across Southeast Asia to the Indonesian islands of Sumatra,

and Bali. Since the end of the last glacial period, it was probably restricted by periods of deep snow lasting longer than six months. Currently, it occurs in less than 6% of its historical range, as it has been extirpated from

Southwest The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...

and Central Asia, large parts of Southeast and East Asia. It now mainly occurs in the

, the

Indochinese Peninsula Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...

, Sumatra and the

Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...

. In China and Myanmar, breeding populations appear to rely on immigration from neighbouring countries while its status in the

Korean Peninsula Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...

is unknown. The tiger is essentially associated with forest habitats. Tiger populations thrive where populations of wild

cervid Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindee ...

s and suids are stable. Records in Central Asia indicate that it occurred foremost in

Tugay Tugay is a form of riparian forest or woodland associated with fluvial and floodplain areas in arid climates. These wetlands are subject to periodic inundation, and largely dependent on floods and groundwater rather than directly from rainfall ...

riverine forests along the

Atrek The Atrek ( fa, اترک, ku, Etrek, tk, Etrek derýasy), also known as the Attruck, Atrak, and Etrek, is a fast-moving river which begins in the mountains of north-eastern Iran (), and flows westward draining into the south-eastern corner of th ...

Amu Darya The Amu Darya, tk, Amyderýa/ uz, Amudaryo// tg, Амударё, Amudaryo ps, , tr, Ceyhun / Amu Derya grc, Ὦξος, Ôxos (also called the Amu, Amo River and historically known by its Latin name or Greek ) is a major river in Central Asi ...

, Syr Darya,

Hari Hari ( sa, हरि) is among the primary epithets of the Hindu preserver deity Vishnu, meaning 'the one who takes away' (sins). It refers to the one who removes darkness and illusion, the one who removes all obstacles to spiritual progress ...

, Chu and Ili Rivers and their tributaries. In the Caucasus, it inhabited hilly and lowland forests. Historical records in

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

are known only from the southern coast of the

Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...

and adjacent

Alborz Mountains The Alborz ( fa, البرز) range, also spelled as Alburz, Elburz or Elborz, is a mountain range in northern Iran that stretches from the border of Azerbaijan along the western and entire southern coast of the Caspian Sea and finally runs no ...

. In the

Amur The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's tenth longest river, forming the border between the Russian Far East and Northeastern China ( Inner Manchuria). The Amur proper is long, ...

Ussuri The Ussuri or Wusuli (russian: Уссури; ) is a river that runs through Khabarovsk and Primorsky Krais, Russia and the southeast region of Northeast China. It rises in the Sikhote-Alin mountain range, flowing north and forming part of the ...

region, it inhabits

Korean pine ''Pinus koraiensis'' is a species of pine known commonly as the Korean pine. It is native to eastern Asia: Korea, northeastern China, Mongolia, the temperate rainforests of the Russian Far East, and central Japan. In the north of its range, it g ...

and

temperate broadleaf and mixed forest Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions. These ...

s, where

riparian forest A riparian forest or riparian woodland is a forested or wooded area of land adjacent to a body of water such as a river, stream, pond, lake, marshland, estuary, canal Canals or artificial waterways are waterways or engineered chann ...

s provide food and water, and serve as dispersal corridors for both tiger and ungulates. On the Indian subcontinent, it inhabits mainly

tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discon ...

, moist

evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zo ...

tropical dry forest The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...

s and the

swamp forest Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are found ...

s of the

Sundarbans Sundarbans (pronounced ) is a mangrove area in the delta formed by the confluence of the Padma, Brahmaputra and Meghna Rivers in the Bay of Bengal. It spans the area from the Baleswar River in Bangladesh's division of Khulna to the Hooghly R ...

. In the

Eastern Himalaya ] The Eastern Himalayas extend from eastern Nepal across Northeast India, Bhutan, the Tibet Autonomous Region to Yunnan in China and northern Myanmar. The climate of this region is influenced by the monsoon of South Asia from June to September. It ...

s, tigers were documented in

up to an elevation of in Bhutan and of in the

Mishmi Hills The Mishmi Hills are located at the northeastern tip of India, in northeastern Arunachal Pradesh. On the Chinese side, they form the southern parts of Nyingchi Prefecture in the Tibet Autonomous Region. These hills occur at the junction of No ...

. In Myanmar, tigers are distributed across the country and among every province. The country is home to two tiger populations, Bengal and Indochinese tigers. In 1996, the composition of the two populations was 60% Bengal tigers and 40% Indochinese tigers. The natural ecological divide for these two populations is assumed to be the Irrawaddy River, but there is no scientific evidence for that hypothesis. DNA studies are needed to confirm it. Today, the presence of tigers was confirmed in the

Hukawng Valley The Hukawng Valley ( my, ဟူးကောင်းချိုင့်ဝှမ်း; also spelt Hukaung Valley) is an isolated valley in Myanmar, roughly in area. It is located in Tanaing Township in the Myitkyina District of Kachin State ...

, Htamanthi Wildlife Sanctuary, and in two small areas in the

Tanintharyi Region Tanintharyi Region ( my, တနင်္သာရီတိုင်းဒေသကြီး, ; Mon: or ; ms, Tanah Sari; formerly Tenasserim Division and subsequently Tanintharyi Division, th, ตะนาวศรี, RTGS: ''Tanao Si'', ; ...

. The

Tenasserim Hills The Tenasserim Hills or Tenasserim Range ( my, တနင်္သာရီ တောင်တန်း, ; th, ทิวเขาตะนาวศรี, , ; ms, Banjaran Tanah Seri/Banjaran Tenang Sari) is the geographical name of a r ...

is an important area, but forests are harvested there. In 2015, tigers were recorded by camera traps for the first time in the hill forests of

Kayin State Kayin State ( my, ကရင်ပြည်နယ်, ; kjp, ဖၠုံခါန်ႋကၞင့်, italics=no; ksw, ကညီကီၢ်စဲၣ်, ), also known by the endonyms Kawthoolei and Karen State, is a state of Myanmar. The ...

. In Thailand, it lives in deciduous and evergreen forests. In Laos, 14 tigers were documented in semi-evergreen and evergreen forest interspersed with grassland in Nam Et-Phou Louey National Protected Area during surveys from 2013 to 2017. In Sumatra, tiger populations range from lowland

peat swamp forest Peat swamp forests are tropical moist forests where waterlogged soil prevents dead leaves and wood from fully decomposing. Over time, this creates a thick layer of acidic peat. Large areas of these forests are being logged at high rates. Peat ...

s to rugged montane forests.

Behaviour and ecology

Social and daily activities

When not subject to human disturbance, the tiger is mainly diurnal. It does not often climb trees but cases have been recorded. It is a strong swimmer and often bathes in ponds, lakes and rivers, thus keeping cool in the heat of the day. Individuals can cross rivers up to wide and can swim up to in a day. During the 1980s, a tiger was observed frequently hunting prey through deep lake water in

Ranthambhore National Park Ranthambore National Park is a national park in Rajasthan, India, with an area of . It is bounded to the north by the Banas River and to the south by the Chambal River. It is named after the historic Ranthambore Fort, which lies within the pa ...

. The tiger is a long-ranging species, and individuals disperse over distances of up to to reach tiger populations in other areas. Radio-collared tigers in

Chitwan National Park , iucn_category = II , location = Central Terai of Nepal , established = 1973 , nearest_city = Bharatpur , map = Nepal Bagmati Province#Nepal#India#South Asia , relief = 1 , label = Chitwan National Park , label_position = top , coordina ...

started dispersing from their natal areas earliest at the age of 19 months. Four females dispersed between , and 10 males between . None of them crossed open cultivated areas that were more than wide, but moved through forested habitat. Adult tigers lead largely solitary lives. They establish and maintain

territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...

but have much wider home ranges within which they roam. Resident adults of either sex generally confine their movements to their home ranges, within which they satisfy their needs and those of their growing cubs. Individuals sharing the same area are aware of each other's movements and activities. The size of the home range mainly depends on prey abundance, geographic area and sex of the individual. In India, home ranges appear to be while in Manchuria, they range from . In Nepal, defended territories are recorded to be for males and for females. Young female tigers establish their first territories close to their mother's. The overlap between the female and her mother's territory reduces with time. Males, however, migrate further than their female counterparts and set out at a younger age to mark out their own area. A young male acquires territory either by seeking out an area devoid of other male tigers, or by living as a transient in another male's territory until he is older and strong enough to challenge the resident male. Young males seeking to establish themselves thereby comprise the highest mortality rate (30–35% per year) amongst adult tigers. Flying Princess (cropped)

To identify his territory, the male marks trees by spraying urine,

anal gland Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involvi ...

secretions, marking trails with feces and marking trees or the ground with their claws. Females also use these "scrapes", urine and fecal markings. Scent markings of this type allow an individual to pick up information on another's identity, sex and reproductive status. Females in

oestrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...

will signal their availability by scent marking more frequently and increasing their vocalisations. Although for the most part avoiding each other, tigers are not always territorial and relationships between individuals can be complex. An adult of either sex will sometimes share its kill with others, even with unrelated tigers.

George Schaller George Beals Schaller (born 1933) is a German-born American mammalogist, biologist, conservationist and author. Schaller is recognized by many as the world's preeminent field biologist, studying wildlife throughout Africa, Asia and South Ameri ...

observed a male share a kill with two females and four cubs. Unlike male lions, male tigers allow females and cubs to feed on the kill before the male is finished with it; all involved generally seem to behave amicably, in contrast to the competitive behaviour shown by a lion pride. Stephen Mills described a social feeding event in Ranthambore National Park: Male tigers are generally less tolerant of other males within their territories than females are of other females. Territory disputes are usually solved by intimidation rather than outright violence. Several such incidents have been observed in which the subordinate tiger yielded by rolling onto its back and showing its belly in a submissive posture. Once dominance has been established, a male may tolerate a subordinate within his range, as long as they do not live in too close quarters. The most serious disputes tend to occur between two males competing for a female in oestrus, sometimes fighting to the death. Tiger's Flehmen Response - Kanha National Park

Tiger's Flehmen Response - Kanha National Park

Facial expressions include the "defense threat", where an individual bares its teeth, flattens its ears and its pupils enlarge. Both males and females show a

flehmen response The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in which ...

, a characteristic grimace, when sniffing urine markings, but flehmen is more often associated with males detecting the markings made by tigresses in oestrus. Tigers roar to signal their presence to other individuals over long distances. This vocalization is forced through an open mouth as it closes and can be heard away. They may roar three or four times in a row, and other tigers may respond in kind. When tense, tigers will moan, a sound similar to a roar but softer and made when the mouth is at least partially closed. Moaning can be heard away. For aggressive encounters, tigers growl,

snarl A snarl is a sound, often a growl or vicious utterance, often accompanied by a facial expression, where the upper lip is raised, and the nostrils widen, generally indicating hate, anger or pain. In addition to humans, other mammals including m ...

and hiss. During an attack, an explosive "coughing roar" or "coughing snarl" is emitted though an open mouth and exposed teeth. Chuffing—soft, low-frequency snorting similar to

purring A purr is a tonal fluttering sound made by some species of felids and two species of genets. It varies in loudness and tone among species and in the same animal. Felids are a family of mammals that belong to the order Carnivora and are info ...

in smaller cats—is heard in more friendly situations. Other vocalizations include grunts, woofs and miaows.

Hunting and diet

In the wild, tigers mostly feed on large and medium-sized mammals, particularly

s weighing . Range-wide the most selected prey are sambar deer,

Manchurian wapiti The Manchurian wapiti (''Cervus canadensis xanthopygus'') is a subspecies of the wapiti native to East Asia. Description The Manchurian wapiti's coat is reddish brown during summer, and brownish gray in winter. It has dark hairs on the neck and ...

barasingha The barasingha (''Rucervus duvaucelii''), also known as the swamp deer, is a deer species distributed in the Indian subcontinent. Populations in northern and central India are fragmented, and two isolated populations occur in southwestern Nepal. ...

and

. Tigers are capable of taking down larger prey like adult

gaur The gaur (''Bos gaurus''; ), also known as the Indian bison, is a bovine native to South Asia and Southeast Asia, and has been listed as Vulnerable on the IUCN Red List since 1986. The global population was estimated at a maximum of 21,000 m ...

and

wild water buffalo The wild water buffalo (''Bubalus arnee''), also called Asian buffalo, Asiatic buffalo and wild buffalo, is a large Bovinae, bovine native to the Indian subcontinent and Southeast Asia. It has been listed as ''Endangered species, Endangered'' i ...

but will also opportunistically eat much smaller prey, such as monkeys,

peafowl Peafowl is a common name for three bird species in the genera '' Pavo'' and '' Afropavo'' within the tribe Pavonini of the family Phasianidae, the pheasants and their allies. Male peafowl are referred to as peacocks, and female peafowl are ref ...

and other ground-based birds, hares, porcupines, and fish. They also prey on other predators, including dogs, leopards, pythons, bears, and crocodiles. Tigers generally do not prey on fully grown adult

Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...

s and

Indian rhinoceros } The Indian rhinoceros (''Rhinoceros unicornis''), also called the Indian rhino, greater one-horned rhinoceros or great Indian rhinoceros, is a rhinoceros species native to the Indian subcontinent. It is listed as Vulnerable on the IUCN Red Li ...

but incidents have been reported. More often, it is the more vulnerable small calves that are taken. When in close proximity to humans, tigers will also sometimes prey on such domestic livestock as cattle, horses, and donkeys. Although almost exclusively carnivorous, tigers will occasionally eat vegetation for

dietary fibre Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by th ...

such as fruit of the slow match tree. Tigers are thought to be mainly nocturnal predators, but in areas where humans are absent, remote-controlled, hidden

camera trap A camera trap is a camera that is automatically triggered by a change in some activity in its vicinity, like presence of an animal or a human being. It is typically equipped with a motion sensor – usually a passive infrared (PIR) senso ...

s recorded them hunting in daylight. They generally hunt alone and ambush their prey as most other cats do, overpowering them from any angle, using their body size and strength to knock the prey off balance. Successful hunts usually require the tiger to almost simultaneously leap onto its quarry, knock it over, and grab the throat or nape with its teeth. Despite their large size, tigers can reach speeds of about but only in short bursts; consequently, tigers must be close to their prey before they break cover. If the prey senses the tiger's presence before this, the tiger usually abandons the hunt rather than give chase or battle pre-alerted prey. Horizontal leaps of up to have been reported, although leaps of around half this distance are more typical. One in 2 to 20 hunts, including stalking near potential prey, ends in a successful kill. When hunting larger animals, tigers prefer to bite the throat and use their powerful forelimbs to hold onto the prey, often simultaneously wrestling it to the ground. The tiger remains latched onto the neck until its target dies of

strangulation Strangling is compression of the neck that may lead to unconsciousness or death by causing an increasingly hypoxic state in the brain. Fatal strangling typically occurs in cases of violence, accidents, and is one of two main ways that hangin ...

. By this method, gaurs and water buffaloes weighing over a ton have been killed by tigers weighing about a sixth as much. Although they can kill healthy adults, tigers often select the calves or infirm of very large species. Healthy adult prey of this type can be dangerous to tackle, as long, strong horns, legs and tusks are all potentially fatal to the tiger. No other extant land predator routinely takes on prey this large on its own. With small prey such as monkeys and hares, the tiger bites the

nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...

, often breaking the

spinal cord The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the sp ...

, piercing the

windpipe The trachea, also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all air-breathing animals with lungs. The trachea extends from the l ...

, or severing the

jugular vein The jugular veins are veins that take deoxygenated blood from the head back to the heart via the superior vena cava. The internal jugular vein descends next to the internal carotid artery and continues posteriorly to the sternocleidomastoid mu ...

common carotid artery In anatomy, the left and right common carotid arteries (carotids) (Entry "carotid"
in
and break the backs of

sloth bear The sloth bear (''Melursus ursinus'') is a myrmecophagous bear species native to the Indian subcontinent. It feeds on fruits, ants and termites. It is listed as vulnerable on the IUCN Red List, mainly because of habitat loss and degradation. ...

s. After killing their prey, tigers sometimes drag it to conceal it in vegetation, grasping with their mouths at the site of the killing bite. This, too, can require great physical strength. In one case, after it had killed an adult gaur, a tiger was observed to drag the massive carcass over a distance of . When 13 men simultaneously tried to drag the same carcass later, they were unable to move it. An adult tiger can go for up to two weeks without eating, then gorge on of flesh at one time. In captivity, adult tigers are fed of meat a day.

Enemies and competitors

Tigers usually prefer to eat self-killed prey, but eat carrion in times of scarcity and also steal prey from other large carnivores. Although predators typically avoid one another, if a prize is under dispute or a serious competitor is encountered, displays of aggression are common. If these fail, the conflicts may turn violent; tigers may kill or even prey on competitors such as leopards,

dhole The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It ...

s, striped hyenas,

wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...

, bears,

pythons The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 42 species are currently recognized. Distribution ...

, and

mugger crocodile The mugger crocodile (''Crocodylus palustris'') is a medium-sized broad- snouted crocodile, also known as mugger and marsh crocodile. It is native to freshwater habitats from southern Iran to the Indian subcontinent, where it inhabits marshes ...

s on occasion. Crocodiles, bears, and large packs of dholes may win conflicts with tigers, and crocodiles and bears can even kill them. The considerably smaller leopard avoids competition from tigers by hunting at different times of the day and hunting different prey. Ecology.info In India's

Nagarhole National Park Nagarahole National Park is a national park located in Kodagu district and Mysore district in Karnataka, India. This park was declared the 37th Tiger Reserve of India in 1999. It is part of the Nilgiri Biosphere Reserve. The Western Ghats N ...

, most prey selected by leopards were from against a preference for heavier prey by tigers. The average prey weight in the two respective big cats in India was against . With relatively abundant prey, tigers and leopards were seen to successfully coexist without competitive exclusion or interspecies dominance hierarchies that may be more common to the African savanna, where the leopard lives beside the lion.

Golden jackal The golden jackal (''Canis aureus''), also called common jackal, is a wolf-like canid that is native to Southeast Europe, Southwest Asia, South Asia, and regions of Southeast Asia. The golden jackal's coat varies in color from a pale creamy ...

s may scavenge on tiger kills. Tigers appear to inhabit the deep parts of a forest while smaller predators like leopards and dholes are pushed closer to the fringes.

Reproduction and life cycle

The tiger

mates Mates is an English surname, and may refer to: * Mates (born 1964), British newsreader and journalist * Michael Mates (born 1934), British politician * Frederick S. Mates, founded the Mates Investment Fund in 1967 that crashed in the bear market ...

all year round, but most cubs are born between March and June, with a second peak in September. Gestation ranges from 93 to 114 days, with an average of 103 to 105 days. A female is only receptive for three to six days. Mating is frequent and noisy during that time. The female gives birth in a sheltered location such as in tall grass, in a dense thicket, cave or rocky crevice. The father generally takes no part in rearing. Litters consist of two or three cubs, rarely as many as six. Cubs weigh from each at birth, and are born with eyes closed. They open their eyes when they are six to 14 days old. Their milk teeth break through at the age of about two weeks. They start to eat meat at the age of eight weeks. At around this time, females usually shift them to a new den. They make short ventures with their mother, although they do not travel with her as she roams her territory until they are older. Females lactate for five to six months. Around the time they are weaned, they start to accompany their mother on territorial walks and are taught how to hunt. A dominant cub emerges in most litters, usually a male. The dominant cub is more active than its siblings and takes the lead in their play, eventually leaving its mother and becoming independent earlier. The cubs start hunting on their own earliest at the age of 11 months, and become independent around 18 to 20 months of age. They separate from their mother at the age of two to two and a half years, but continue to grow until the age of five years. Young females reach sexual maturity at three to four years, whereas males at four to five years. Unrelated wandering male tigers often kill cubs to make the female receptive, since the tigress may give birth to another litter within five months if the cubs of the previous litter are lost. The mortality rate of tiger cubs is about 50% in the first two years. Few other predators attack tiger cubs due to the diligence and ferocity of the mother. Apart from humans and other tigers, common causes of cub mortality are starvation, freezing, and accidents. Generation length of the tiger is about eight years. The oldest recorded captive tiger lived for 26 years. Occasionally, male tigers participate in raising cubs, usually their own, but this is extremely rare and not always well understood. In May 2015, Amur tigers were photographed by camera traps in the Sikhote-Alin Nature Reserve. The photos show a male Amur tiger pass by, followed by a female and three cubs within the span of about two minutes. In Ranthambore, a male Bengal tiger raised and defended two orphaned female cubs after their mother had died of illness. The cubs remained under his care, he supplied them with food, protected them from his rival and sister, and apparently also trained them.

Conservation

In the 1990s, a new approach to tiger conservation was developed: Tiger Conservation Units (TCUs), which are blocks of habitat that have the potential to host tiger populations in 15 habitat types within five

bioregion A bioregion is an ecologically and geographically defined area that is smaller than a biogeographic realm, but larger than an ecoregion or an ecosystem, in the World Wide Fund for Nature classification scheme. There is also an attempt to use the ...

s. Altogether 143 TCUs were identified and prioritized based on size and integrity of habitat, poaching pressure and population status. They range in size from . In 2016, an estimate of a global wild tiger population of approximately 3,890 individuals was presented during the Third Asia Ministerial Conference on Tiger Conservation. The WWF subsequently declared that the world's count of wild tigers had risen for the first time in a century. Major threats to the tiger include

, habitat fragmentation and

for fur and body parts, which have simultaneously greatly reduced tiger populations in the wild. In India, only 11% of the historical tiger habitat remains due to habitat fragmentation. Demand for tiger parts for use in

traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of acti ...

has also been cited as a major threat to tiger populations. Some estimates suggest that there are fewer than 2,500 mature breeding individuals, with no subpopulation containing more than 250 mature breeding individuals. India is home to the world's largest population of wild tigers. A 2014 census estimated a population of 2,226, a 30% increase since 2011. On International Tiger Day 2019, the 'Tiger Estimation Report 2018' was released by Prime Minister Narendra Modi. The report estimates a population of 2967 tigers in India with 25% increase since 2014. Modi said "India is one of the safest habitats for tigers as it has achieved the target of doubling the tiger population from 1411 in 2011 to 2967 in 2019". As of 2022, India accounts for 75 percent of global tiger population. In 1973, India's ''

Project Tiger Project Tiger is a tiger conservation programme launched in April 1973 by the Government of India during Prime Minister Indira Gandhi's tenure. The project aims at ensuring a viable population of the Bengal tiger in its natural habitats, protect ...

'', started by Indira Gandhi, established numerous tiger reserves. The project was credited with tripling the number of wild Bengal tigers from some 1,200 in 1973 to over 3,500 in the 1990s, but a 2007 census showed that numbers had dropped back to about 1,400 tigers because of poaching. Following the report, the Indian government pledged $153 million to the initiative, set up measures to combat poaching, promised funds to relocate up to 200,000 villagers in order to reduce human-tiger interactions, and set up eight new

tiger reserves The tiger reserves of India were set up in 1973 and are governed by Project Tiger, which is administrated by the National Tiger Conservation Authority. Until 2018, 50 protected areas have been designated tiger reserves. In 2022, 53rd tiger reserve ...

. India also reintroduced tigers to the

Sariska Tiger Reserve Sariska Tiger Reserve is a tiger reserve in Alwar district, Rajasthan, India. It stretches over an area of comprising scrub-thorn arid forests, dry deciduous forests, grasslands, and rocky hills. This area was a hunting preserve of the Alwa ...

and by 2009 it was claimed that poaching had been effectively countered at

Ranthambore National Park Ranthambore National Park is a national park in Rajasthan, India, with an area of . It is bounded to the north by the Banas River and to the south by the Chambal River. It is named after the historic Ranthambore Fort, which lies within the par ...

. In the 1940s, the Siberian tiger was on the brink of extinction with only about 40 animals remaining in the wild in Russia. As a result, anti-poaching controls were put in place by the

Soviet Union The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a List of former transcontinental countries#Since 1700, transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, ...

and a network of protected zones (

zapovednik Zapovednik (russian: заповедник, plural , from the Russian , 'sacred, prohibited from disturbance, committed o protect committed o heritage; ) is an established term on the territory of the former Soviet Union for a protected area which ...

s) were instituted, leading to a rise in the population to several hundred. Poaching again became a problem in the 1990s, when the

economy of Russia The economy of Russia has gradually transformed from a planned economy into a mixed market-oriented economy. —Rosefielde, Steven, and Natalia Vennikova. “Fiscal Federalism in Russia: A Critique of the OECD Proposals.” Cambridge Journa ...

collapsed. The major obstacle in preserving the species is the enormous territory individual tigers require, up to needed by a single female and more for a single male. Current conservation efforts are led by local governments and NGO's in concert with international organisations, such as the

World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...

and the

Wildlife Conservation Society The Wildlife Conservation Society (WCS) is a non-governmental organization headquartered at the Bronx Zoo in New York City, that aims to conserve the world's largest wild places in 14 priority regions. Founded in 1895 as the New York Zoological ...

. The competitive exclusion of wolves by tigers has been used by Russian conservationists to convince hunters to tolerate the big cats. Tigers have less impact on ungulate populations than do wolves, and are effective in controlling the latter's numbers. In 2005, there were thought to be about 360 animals in Russia, though these exhibited little genetic diversity. However, in a decade later, the Siberian tiger census was estimated from 480 to 540 individuals. In China, tigers became the target of large-scale 'anti-pest' campaigns in the early 1950s, where suitable habitats were fragmented following deforestation and resettlement of people to rural areas, who hunted tigers and prey species. Though tiger hunting was prohibited in 1977, the population continued to decline and is considered extinct in southern China since 2001. Having earlier rejected the Western-led environmentalist movement, China changed its stance in the 1980s and became a party to the

CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...

treaty. By 1993 it had banned the trade in tiger parts, and this diminished the use of tiger bones in

. The

Tibetan people The Tibetan people (; ) are an East Asian ethnic group native to Tibet. Their current population is estimated to be around 6.7 million. In addition to the majority living in Tibet Autonomous Region of China, significant numbers of Tibetans l ...

's trade in tiger skins has also been a threat to tigers. The pelts were used in clothing, tiger-skin ''

chuba A chuba is a long sheepskin coat made of thick Tibetan wool worn by many of the nomadic peoples of high altitude in the cold mountains of Tibet. The traditional sherpa clothing is distinctive to solu-khumba, the basic garment of the sherpas; t ...

'' being worn as fashion. In 2006 the

14th Dalai Lama The 14th Dalai Lama (spiritual name Jetsun Jamphel Ngawang Lobsang Yeshe Tenzin Gyatso, known as Tenzin Gyatso (Tibetan: བསྟན་འཛིན་རྒྱ་མཚོ་, Wylie: ''bsTan-'dzin rgya-mtsho''); né Lhamo Thondup), known as ...

was persuaded to take up the issue. Since then there has been a change of attitude, with some Tibetans publicly burning their chubas. Wild Sumatran tiger

In 1994, the Indonesian Sumatran Tiger Conservation Strategy addressed the potential crisis that tigers faced in Sumatra. The Sumatran Tiger Project (STP) was initiated in June 1995 in and around the

Way Kambas National Park Way Kambas National Park ( id, Taman Nasional Way Kambas, abbreviated TNWK) is a national park covering 1,300 square kilometres in Lampung province, southern Sumatra, Indonesia. It consists of swamp forest and lowland rain forest, mostly of second ...

to ensure the long-term viability of wild Sumatran tigers and to accumulate data on tiger life-history characteristics vital for the management of wild populations. By August 1999, the teams of the STP had evaluated 52 sites of potential tiger habitat in Lampung Province, of which only 15 these were intact enough to contain tigers.Tilson, R. (1999). ''Sumatran Tiger Project Report No. 17 & 18: July − December 1999''. Grant number 1998-0093-059. Indonesian Sumatran Tiger Steering Committee, Jakarta. In the framework of the STP a community-based conservation programme was initiated to document the tiger-human dimension in the park to enable conservation authorities to resolve tiger-human conflicts based on a comprehensive database rather than anecdotes and opinions.Nyhus, P., Sumianto and R. Tilson (1999). ''The tiger-human dimension in southeast Sumatra'', pp. 144–145 in: Seidensticker, J., Christie, S. and Jackson, P. (eds). ''Riding the tiger: tiger conservation in human-dominated landscapes''. Cambridge University Press, Cambridge, . The

and

Panthera Corporation Panthera Corporation, or Panthera, is a charitable organization devoted to preserving wild cats and their ecosystems around the globe. Founded in 2006, Panthera is devoted to the conservation of the world’s 40 species of wild cats and the vast ec ...

formed the collaboration ''Tigers Forever'', with field sites including the world's largest tiger reserve, the

Hukaung Valley The Hukawng Valley ( my, ဟူးကောင်းချိုင့်ဝှမ်း; also spelt Hukaung Valley) is an isolated valley in Myanmar, roughly in area. It is located in Tanaing Township in the Myitkyina District of Kachin State i ...

in Myanmar. Other reserves were in the Western Ghats in India, Thailand, Laos, Cambodia, the Russian Far East covering in total about . Tigers have been studied in the wild using a variety of techniques. Tiger population have been estimated using plaster casts of their

pugmark Pugmark is the term used to refer to the footprint of most animals (especially megafauna). "Pug" means foot in Hindi (Sanskrit पद् "pad"; Greek πούς "poús"). Every individual animal species has a distinct pugmark and as such this is us ...

s, although this method was criticized as being inaccurate. More recent techniques include the use of

s and studies of DNA from tiger scat, while radio-collaring has been used to track tigers in the wild. Tiger spray has been found to be just as good, or better, as a source of DNA than scat.

Relation with humans

Tiger hunting

The tiger has been one of the most sought after game animals of Asia. Tiger hunting took place on a large scale in the early 19th and 20th centuries, being a recognised and admired sport by the British in colonial India, the

maharaja Mahārāja (; also spelled Maharajah, Maharaj) is a Sanskrit title for a "great ruler", "great king" or " high king". A few ruled states informally called empires, including ruler raja Sri Gupta, founder of the ancient Indian Gupta Empire, a ...

s and aristocratic class of the erstwhile princely states of pre-independence India. A single maharaja or English hunter could claim to kill over a hundred tigers in their hunting career. Over 80,000 tigers were slaughtered in just 50 years spanning from 1875 to 1925 in British-ruled India. Tiger hunting was done by some hunters on foot; others sat up on '' machans'' with a goat or buffalo tied out as bait; yet others on elephant-back. King

George V George V (George Frederick Ernest Albert; 3 June 1865 – 20 January 1936) was King of the United Kingdom and the British Dominions, and Emperor of India, from 6 May 1910 until his death in 1936. Born during the reign of his grandmother Qu ...

on his visit to Colonial India in 1911 killed 39 tigers in a matter of 10 days One of these is on display at the

Royal Albert Memorial Museum Royal Albert Memorial Museum & Art Gallery (RAMM) is a museum and art gallery in Exeter, Devon, the largest in the city. It holds significant and diverse collections in areas such as zoology, anthropology, fine art, local and overseas archaeolo ...

. Historically, tigers have been hunted at a large scale so their famous striped skins could be collected. The trade in tiger skins peaked in the 1960s, just before international conservation efforts took effect. By 1977, a tiger skin in an English market was considered to be worth US$4,250.

Body part use

Tiger parts are commonly used as amulets in South and Southeast Asia. In the Philippines, the fossils in Palawan were found besides stone tools. This, besides the evidence for cuts on the bones, and the use of fire, suggests that early humans had accumulated the bones, and the condition of the tiger subfossils, dated to approximately 12,000 to 9,000 years ago, differed from other fossils in the assemblage, dated to the

Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...

. The tiger subfossils showed longitudinal fracture of the

cortical bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, and ...

due to weathering, which suggests that they had

post-mortem An autopsy (post-mortem examination, obduction, necropsy, or autopsia cadaverum) is a surgical procedure that consists of a thorough examination of a corpse by dissection to determine the cause, mode, and manner of death or to evaluate any di ...

been exposed to light and air. Tiger canines were found in Ambangan sites dating to the 10th to 12th centuries in

Butuan Butuan (pronounced ), officially the City of Butuan ( ceb, Dakbayan sa Butuan; Butuanon: ''Dakbayan hong Butuan''; fil, Lungsod ng Butuan), is a 1st class highly urbanized city in the region of Caraga, Philippines. It is the ''de facto'' c ...

Mindanao Mindanao ( ) ( Jawi: مينداناو) is the second-largest island in the Philippines, after Luzon, and seventh-most populous island in the world. Located in the southern region of the archipelago, the island is part of an island group of ...

COLLECTIE TROPENMUSEUM Een groep mannen en kinderen poseert bij een pas geschoten tijger te Malingping in Bantam West-Java TMnr 10006636

Many people in China and other parts of Asia have a belief that various tiger parts have medicinal properties, including as pain killers and

aphrodisiac An aphrodisiac is a substance that increases sexual desire, sexual attraction, sexual pleasure, or sexual behavior. Substances range from a variety of plants, spices, foods, and synthetic chemicals. Natural aphrodisiacs like cannabis or cocai ...

s. There is no scientific evidence to support these beliefs. The use of tiger parts in pharmaceutical drugs in China is already banned, and the government has made some offences in connection with tiger poaching punishable by death. Furthermore, all trade in tiger parts is illegal under the

Convention on International Trade in Endangered Species of Wild Fauna and Flora CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...

and a domestic trade ban has been in place in China since 1993. However, the trading of tiger parts in Asia has become a major black market industry and governmental and conservation attempts to stop it have been ineffective to date. Almost all black marketers engaged in the trade are based in China and have either been shipped and sold within in their own country or into

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

or Japan. The Chinese subspecies was almost completely decimated by killing for commerce due to both the parts and skin trades in the 1950s through the 1970s. Contributing to the illegal trade, there are a number of tiger farms in the country specialising in breeding them for profit. It is estimated that between 5,000 and 10,000 captive-bred, semi-tame animals live in these farms today. However, many tigers for traditional medicine black market are wild ones shot or snared by

poacher Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...

s and may be caught anywhere in the tiger's remaining range (from Siberia to India to the Malay Peninsula to Sumatra). In the Asian black market, a

tiger penis In traditional Chinese medicine, a tiger penis ( Vietnamese: ''Pín hổ''; ) is said to have important therapeutic properties. However, there is no scientific proof that tiger penis can be used to treat any medical disorder. The demand for tiger ...

can be worth the equivalent of around $300 U.S. dollars. In the years of 1990 through 1992, 27 million products with tiger derivatives were found. In July 2014 at an international convention on endangered species in

Geneva , neighboring_municipalities= Carouge, Chêne-Bougeries, Cologny, Lancy, Grand-Saconnex, Pregny-Chambésy, Vernier, Veyrier , website = https://www.geneve.ch/ Geneva ( ; french: Genève ) frp, Genèva ; german: link=no, Genf ; it, Ginevr ...

, Switzerland, a Chinese representative admitted for the first time his government was aware trading in tiger skins was occurring in China.

Man-eating tigers

Wild tigers that have had no prior contact with humans actively avoid interactions with them. However, tigers cause more human deaths through direct attack than any other wild mammal. Attacks are occasionally provoked, as tigers lash out after being injured while they themselves are hunted. Attacks can be provoked accidentally, as when a human surprises a tiger or inadvertently comes between a mother and her young, or as in a case in rural India when a postman startled a tiger, used to seeing him on foot, by riding a bicycle. Occasionally tigers come to view people as prey. Such attacks are most common in areas where population growth, logging, and farming have put pressure on tiger habitats and reduced their wild prey. Most man-eating tigers are old, missing teeth, and unable to capture their preferred prey. For example, the

Champawat Tiger The Champawat Tiger was a Bengal Tigress responsible for an estimated 436 deaths in Nepal and the Kumaon division of India, during the last years of the 19th century and the first years of the 20th century. Her attacks have been listed in the ...

, a tigress found in

Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...

and then India, had two broken canines. She was responsible for an estimated 430 human deaths, the most attacks known to be perpetrated by a single wild animal, by the time she was shot in 1907 by

Jim Corbett Edward James Corbett (25 July 1875 – 19 April 1955) was a British hunter, tracker, naturalist, and author who hunted a number of man-eating tigers and leopards in the Indian subcontinent. He held the rank of colonel in the British Indian ...

. According to Corbett, tiger attacks on humans are normally in daytime, when people are working outdoors and are not keeping watch. Early writings tend to describe man-eating tigers as cowardly because of their ambush tactics. The Man-Eater of Segur", from ''Nine Man-Eaters and One Rogue'', Kenneth Anderson, Allen & Unwin, 1954 Man-eaters have been a particular problem in recent decades in India and Bangladesh, especially in

Kumaon Kumaon or Kumaun may refer to: * Kumaon division, a region in Uttarakhand, India * Kumaon Kingdom, a former country in Uttarakhand, India * Kumaon, Iran, a village in Isfahan Province, Iran * , a ship of the Royal Indian Navy during WWII See also ...

, Garhwal and the

mangrove swamps of

Bengal Bengal ( ; bn, বাংলা/বঙ্গ, translit=Bānglā/Bôngô, ) is a geopolitical, cultural and historical region in South Asia, specifically in the eastern part of the Indian subcontinent at the apex of the Bay of Bengal, predom ...

, where some healthy tigers have hunted humans. Because of rapid habitat loss attributed to

climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...

, tiger attacks have increased in the Sundarbans. The Sundarbans area had 129 human deaths from tigers from 1969 to 1971. In the 10 years prior to that period, about 100 attacks per year in the Sundarbans, with a high of around 430 in some years of the 1960s. Unusually, in some years in the Sundarbans, more humans are killed by tigers than vice versa. In 1972, India's production of

honey Honey is a sweet and viscous substance made by several bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primar ...

and beeswax dropped by 50% when at least 29 people who gathered these materials were devoured. In 1986 in the Sundarbans, since tigers almost always attack from the rear, masks with human faces were worn on the back of the head, on the theory that tigers usually do not attack if seen by their prey. This decreased the number of attacks only temporarily. All other means to prevent attacks, such as providing more prey or using electrified human dummies, did not work as well.

In captivity

Ringling Bros and Barnum & Bailey Circus Gunther Gebel-Williams 1969

In Ancient Roman times, tigers were kept in menageries and

amphitheatre An amphitheatre (British English) or amphitheater (American English; both ) is an open-air venue used for entertainment, performances, and sports. The term derives from the ancient Greek ('), from ('), meaning "on both sides" or "around" and ...

s to be exhibited, trained and paraded, and were often provoked to fight

gladiators A gladiator ( la, gladiator, "swordsman", from , "sword") was an armed combatant who entertained audiences in the Roman Republic and Roman Empire in violent confrontations with other gladiators, wild animals, and condemned criminals. Some gla ...

and other exotic beasts. Since the 17th century, tigers, being rare and ferocious, were sought after to keep at European castles as symbols of their owners' power. Tigers became central

zoo A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for conservation purposes. The term ''zoological garden'' refers to zoo ...

and

circus A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicyclis ...

exhibits in the 18th century: a tiger could cost up to 4,000

francs The franc is any of various units of currency. One franc is typically divided into 100 centimes. The name is said to derive from the Latin inscription ''francorum rex'' (King of the Franks) used on early French coins and until the 18th centu ...

in France (for comparison, a professor of the Beaux-Arts at Lyons earned only 3,000 francs a year), or up to $3,500 in the United States, where a lion cost no more than $1,000. In 2007, over 4,000 captive tigers lived in China, of which 3,000 were held by about 20 larger facilities, with the rest held by some 200 smaller facilities. In 2011, 468 facilities in the USA kept 2,884 tigers. Nineteen US states banned private ownership of tigers, fifteen require a license, and sixteen states have no regulation. Genetic ancestry of 105 captive tigers from fourteen countries and regions showed that forty-nine animals belonged distinctly to five subspecies; fifty-two animals had mixed subspecies origins. Many Siberian tigers in zoos today are actually the result of crosses with Bengal tigers.

Cultural depictions

Tigers and their superlative qualities have been a source of fascination for mankind since ancient times, and they are routinely visible as important cultural and media motifs. They are also considered one of the

, and are used as the face of conservation campaigns worldwide. In a 2004 online poll conducted by cable television channel

Animal Planet Animal Planet (stylized in all lowercase since 2018) is an American multinational pay television channel owned by the Warner Bros. Discovery Networks unit of Warner Bros. Discovery. First established on June 1, 1996, the network is primarily ...

, involving more than 50,000 viewers from 73 countries, the tiger was voted the world's favourite animal with 21% of the vote, narrowly beating the dog.

Mythology and legend

Chinese mythology Chinese mythology () is mythology that has been passed down in oral form or recorded in literature in the geographic area now known as Greater China. Chinese mythology includes many varied myths from regional and cultural traditions. Much of t ...

and

culture Culture () is an umbrella term which encompasses the social behavior, institutions, and norms found in human societies, as well as the knowledge, beliefs, arts, laws, customs, capabilities, and habits of the individuals in these groups ...

, the

tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on ...

is one of the 12 animals of the

Chinese zodiac The Chinese zodiac is a traditional classification scheme based on the lunar calendar that assigns an animal and its reputed attributes to each year in a repeating twelve-year cycle. Originating from China, the zodiac and its variations remain ...

. In

Chinese art Chinese art is visual art that originated in or is practiced in China, Greater China or by Chinese artists. Art created by Chinese residing outside of China can also be considered a part of Chinese art when it is based in or draws on Chinese ...

, the tiger is depicted as an earth symbol and equal rival of the

Chinese dragon The Chinese dragon, also known as ''loong'', ''long'' or ''lung'', is a legendary creature in Chinese mythology, Chinese folklore, and Chinese culture at large. Chinese dragons have many animal-like forms such as turtles and fish, but are most ...

– the two representing matter and spirit respectively. The Southern Chinese martial art

Hung Ga Hung Ga (), Hung Kuen (), or Hung Ga Kuen () is a southern Chinese martial art belonging to the southern Shaolin styles. The hallmarks of Hung Ga are strong stances, notably the horse stance, or "si ping ma" (), and strong hand techniques, not ...

is based on the movements of the tiger and the crane. In Imperial China, a tiger was the personification of war and often represented the highest army

general A general officer is an officer of high rank in the armies, and in some nations' air forces, space forces, and marines or naval infantry. In some usages the term "general officer" refers to a rank above colonel."general, adj. and n.". OED ...

(or present day

defense secretary The United States secretary of defense (SecDef) is the head of the United States Department of Defense, the executive department of the U.S. Armed Forces, and is a high ranking member of the federal cabinet. DoDD 5100.1: Enclosure 2: a The ...

), while the emperor and empress were represented by a dragon and

phoenix Phoenix most often refers to: * Phoenix (mythology), a legendary bird from ancient Greek folklore * Phoenix, Arizona, a city in the United States Phoenix may also refer to: Mythology Greek mythological figures * Phoenix (son of Amyntor), a ...

, respectively. The

White Tiger The white tiger or bleached tiger is a leucistic pigmentation variant of the Mainland tiger. It is reported in the wild from time to time in the Indian states of Madhya Pradesh, Assam, West Bengal, Bihar, Odisha, in the Sunderbans region and ...

() is one of the

Four Symbols The Four Symbols (, literally meaning "four images"), are four mythological creatures appearing among the Chinese constellations along the ecliptic, and viewed as the guardians of the four cardinal directions. These four creatures are also ref ...

of the

Chinese constellation Traditional Chinese astronomy has a system of dividing the celestial sphere into asterisms or constellations, known as "officials" ( Chinese ''xīng guān''). The Chinese asterisms are generally smaller than the constellations of Hellenisti ...

s. It is sometimes called the White Tiger of the West (), and it represents the west and the autumn season. The tiger's tail appears in stories from countries including China and Korea, it being generally inadvisable to grasp a tiger by the tail. In Korean myth and

, the tiger is regarded as a guardian that drives away evil spirits and a sacred creature that brings good luck – the symbol of courage and absolute power. For the people who live in and around the forests of Korea, the tiger considered the symbol of the Mountain Spirit or King of mountain animals. So, Koreans also called the tigers "San Gun" (산군) means Mountain Lord. In

Buddhism Buddhism ( , ), also known as Buddha Dharma and Dharmavinaya (), is an Indian religions, Indian religion or Indian philosophy#Buddhist philosophy, philosophical tradition based on Pre-sectarian Buddhism, teachings attributed to the Buddha. ...

, the tiger is one of the Three Senseless Creatures, symbolising anger, with the monkey representing greed and the deer lovesickness. The

Tungusic peoples Tungusic peoples are an ethno-linguistic group formed by the speakers of Tungusic languages (or Manchu–Tungus languages). They are native to Siberia and Northeast Asia. The Tungusic phylum is divided into two main branches, northern (Evenic or ...

considered the Siberian tiger a near-deity and often referred to it as "Grandfather" or "Old man". The Udege and Nanai called it "Amba". The Manchu considered the Siberian tiger as "Hu Lin," the king. In

Hinduism Hinduism () is an Indian religion or '' dharma'', a religious and universal order or way of life by which followers abide. As a religion, it is the world's third-largest, with over 1.2–1.35 billion followers, or 15–16% of the global p ...

, the god

Shiva Shiva (; sa, शिव, lit=The Auspicious One, Śiva ), also known as Mahadeva (; ɐɦaːd̪eːʋɐ, or Hara, is one of the principal deities of Hinduism. He is the Supreme Being in Shaivism, one of the major traditions within Hindu ...

wears and sits on tiger skin. The ten-armed warrior goddess

Durga Durga ( sa, दुर्गा, ) is a major Hindu goddess, worshipped as a principal aspect of the mother goddess Mahadevi. She is associated with protection, strength, motherhood, destruction, and wars. Durga's legend centres around c ...

rides the tigress (or lioness) Damon into battle. In southern India the god

Ayyappan Ayyappan an incarnation of dharma sastha, also called Manikandan, is a Hindu deity popular in Southern India, He is considered to be the epitome of dharma, truth, and righteousness and is often called upon to obliterate evil. Although devotion ...

was associated with a tiger. Dingu-Aneni is the god in North-East India is also associated with tiger. The

weretiger A werecat (also written in a hyphenated form as were-cat) is an analog to "werewolf" for a feline therianthropic creature. Etymology Ailuranthropy comes from the Greek root words ''ailouros'' meaning "cat",< and ''anthropos'', meaning "human" ...

replaces the

werewolf In folklore, a werewolf (), or occasionally lycanthrope (; ; uk, Вовкулака, Vovkulaka), is an individual that can shapeshift into a wolf (or, especially in modern film, a therianthropic hybrid wolf-like creature), either purposely ...

in shapeshifting folklore in Asia; in India they were evil sorcerers, while in Indonesia and Malaysia they were somewhat more benign. In Greco-Roman tradition, the tiger was depicted being ridden by the god

Dionysus In ancient Greek religion and myth, Dionysus (; grc, Διόνυσος ) is the god of the grape-harvest, winemaking, orchards and fruit, vegetation, fertility, insanity, ritual madness, religious ecstasy, festivity, and theatre. The Roma ...

Literature and media

In the Hindu epic

Mahabharata The ''Mahābhārata'' ( ; sa, महाभारतम्, ', ) is one of the two major Sanskrit literature, Sanskrit Indian epic poetry, epics of ancient India in Hinduism, the other being the ''Ramayana, Rāmāyaṇa''. It narrates the s ...

, the tiger is fiercer and more ruthless than the lion.

William Blake William Blake (28 November 1757 – 12 August 1827) was an English poet, painter, and printmaker. Largely unrecognised during his life, Blake is now considered a seminal figure in the history of the poetry and visual art of the Romantic Age. ...

's poem in his ''

Songs of Experience ''Songs of Innocence and of Experience'' is a collection of illustrated poems by William Blake. It appeared in two phases: a few first copies were printed and illuminated by Blake himself in 1789; five years later, he bound these poems with a ...

'' (1794), titled "

The Tyger "The Tyger" is a poem by the English poet William Blake, published in 1794 as part of his '' Songs of Experience'' collection and rising to prominence in the romantic period. The poem is one of the most anthologised in the English literary can ...

", portrays the tiger as a menacing and fearful animal. In

Rudyard Kipling Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)''The Times'', (London) 18 January 1936, p. 12. was an English novelist, short-story writer, poet, and journalist. He was born in British India, which inspired much of his work. ...

's 1894 ''

The Jungle Book ''The Jungle Book'' (1894) is a collection of stories by the English author Rudyard Kipling. Most of the characters are animals such as Shere Khan the tiger and Baloo the bear, though a principal character is the boy or "man-cub" Mowgli, w ...

'', the tiger Shere Khan is the mortal enemy of the human protagonist Mowgli.

Yann Martel Yann Martel, (born 25 June 1963) is a Canadian author who wrote the Man Booker Prize–winning novel '' Life of Pi'', an international bestseller published in more than 50 territories. It has sold more than 12 million copies worldwide and spen ...

's 2001 Booker Prize winning novel ''

Life of Pi ''Life of Pi'' is a Canadian philosophical novel by Yann Martel published in 2001. The protagonist is Piscine Molitor "Pi" Patel, an Indian boy from Pondicherry, India who explores issues of spirituality and metaphysics from an early age. He ...

'', features the title character surviving shipwreck for months on a small boat with a large Bengal tiger while avoiding being eaten. The story was adapted in

Ang Lee Ang Lee (; born October 23, 1954) is a Taiwanese filmmaker. Born in Pingtung County of southern Taiwan, Lee was educated in Taiwan and later in the United States. During his filmmaking career, he has received international critical and popula ...

's 2012 feature film of the same name. Friendly tiger characters include

Tigger Tigger is a fictional character, an anthropomorphic stuffed tiger. He was originally introduced in the 1928 story collection '' The House at Pooh Corner'', the sequel to the 1926 book ''Winnie-the-Pooh'' by A. A. Milne. Like other Pooh char ...

A. A. Milne Alan Alexander Milne (; 18 January 1882 – 31 January 1956) was an English writer best known for his books about the teddy bear Winnie-the-Pooh, as well as for children's poetry. Milne was primarily a playwright before the huge success of Winni ...

Winnie-the-Pooh Winnie-the-Pooh, also called Pooh Bear and Pooh, is a fictional anthropomorphic teddy bear created by English author A. A. Milne and English illustrator E. H. Shepard. The first collection of stories about the character was the book ''Win ...

and

Hobbes Thomas Hobbes ( ; 5/15 April 1588 – 4/14 December 1679) was an English philosopher, considered to be one of the founders of modern political philosophy. Hobbes is best known for his 1651 book ''Leviathan'', in which he expounds an influe ...

of the comic strip ''

Calvin and Hobbes ''Calvin and Hobbes'' is a daily American comic strip created by cartoonist Bill Watterson that was syndicated from November 18, 1985, to December 31, 1995. Commonly cited as "the last great newspaper comic", ''Calvin and Hobbes'' has enjoyed b ...

'', both represented as stuffed animals come to life.

Tony the Tiger Tony the Tiger is the advertising cartoon mascot for Kellogg's Frosted Flakes (also known as Frosties) breakfast cereal, appearing on its packaging and advertising. Tony has also been the mascot for related cereals such as Tony's Cinnamon K ...

is a famous mascot for Kellogg's breakfast cereal

Frosted Flakes Frosted Flakes or Frosties is a breakfast cereal, produced by the Kellogg Company and consisting of sugar-coated corn flakes. It was introduced in the United States, in 1952, as "Sugar Frosted Flakes". The word "sugar" was dropped from the name ...

, known for his catchphrase "They're Gr-r-reat!".

Heraldry and emblems

The tiger is one of the animals displayed on the Pashupati seal of the

Indus Valley civilisation The Indus Valley Civilisation (IVC), also known as the Indus Civilisation was a Bronze Age civilisation in the northwestern regions of South Asia, lasting from 3300 BCE to 1300 BCE, and in its mature form 2600 BCE to 1900& ...

. The tiger was the emblem of the

Chola Dynasty The Chola dynasty was a Tamil thalassocratic empire of southern India and one of the longest-ruling dynasties in the history of the world. The earliest datable references to the Chola are from inscriptions dated to the 3rd century BCE ...

and was depicted on coins, seals and banners. The seals of several Chola copper coins show the tiger, the Pandyan emblem fish and the Chera dynasty, Chera emblem bow, indicating that the Cholas had achieved political supremacy over the latter two dynasties. Gold coins found in Kavilayadavalli in the Nellore district of Andhra Pradesh have motifs of the tiger, bow and some indistinct marks. The tiger symbol of Chola Empire was later adopted by the Liberation Tigers of Tamil Eelam and the tiger became a symbol of the unrecognised state of Tamil Eelam and Tamil independence movement. The Bengal tiger is the

of India and Bangladesh. The Malaysian tiger is the national animal of

. The Siberian tiger is the national animal of

. The Tiger is featured on the logo of the Delhi Capitals Indian Premier League, IPL team. In European heraldry, the Tyger (heraldry), tyger, a depiction of a tiger as imagined by European artists, is among the creatures used in charges and supporters. This creature has several notable differences from real tigers, lacking stripes and having a leonine tufted tail and a head terminating in large, pointed jaws. A more realistic tiger entered the heraldic armory through the British Empire's expansion into Asia, and is referred to as the Bengal tiger to distinguish it from its older counterpart. The Bengal tiger is not a common creature in heraldry, but is used as a supporter in the arms of Bombay and emblazoned on the shield of the University of Madras.Arthur Fox-Davies, ''A Complete Guide to Heraldry'', T.C. and E.C. Jack, London, 1909, 191–192, https://archive.org/details/completeguidetoh00foxduoft.

References

External links

* * * * * {{Portal bar, Cats, Mammals, Animals, Biology, Asia Apex predators Big cats Conservation-reliant species EDGE species Extant Pleistocene first appearances Fauna of South Asia Fauna of Southeast Asia Felids of Asia Mammals described in 1758 Mammals of East Asia National symbols of India National symbols of Bangladesh National symbols of Malaysia National symbols of Singapore Panthera Species endangered by agricultural development Species endangered by deliberate extirpation efforts Species endangered by human consumption for medicinal or magical purposes Species endangered by logging Species endangered by urbanization Taxa named by Carl Linnaeus