Pallas's fish-eagle
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Pallas's fish eagle (''Haliaeetus leucoryphus''), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish
sea eagle A sea eagle or fish eagle (also called erne or ern, mostly in reference to the white-tailed eagle) is any of the birds of prey in the genus ''Haliaeetus'' in the bird of prey family Accipitridae. Taxonomy and evolution The genus ''Haliaeetus'' ...
. It breeds in the east
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sib ...
in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, China,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
and
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
. It is listed as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
. It is partially migratory, with
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
n birds wintering among the southern Asian birds in northern India, and also further west to the
Persian Gulf The Persian Gulf ( fa, خلیج فارس, translit=xalij-e fârs, lit=Gulf of Fars, ), sometimes called the ( ar, اَلْخَلِيْجُ ٱلْعَرَبِيُّ, Al-Khalīj al-ˁArabī), is a mediterranean sea in Western Asia. The bo ...
.


Description

The Pallas's fish eagle has a light sandy-brown hood and a whitish face. The wings are darker brown and the back rufous. The long, slender wings (particularly slender for a sea eagle) are rather dark brown underneath. The tail is black with a wide, distinctive white stripe. Juveniles are overall darker, cooler brown with no band on the tail but with several pale areas on the wing, including the underwing coverts and inner primaries. This results in underwings that have a white band in young fish eagles. It takes until the 4th year or so to obtain adult plumage. Among related species,, it mainly overlaps in range with the quite dissimilar much shorter winged and slightly smaller
grey-headed fish eagle The grey-headed fish eagle (''Haliaeetus ichthyaetus'') is a fish-eating bird of prey from South East Asia.Robson, C. (2000). ''A Field Guide to the Birds of South-East Asia''. UK: New Holland Publishers. It is a large stocky raptor with adults h ...
and scarcely with the larger, bulkier and much broader winged
white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diurnal raptors ...
, which also seldom resembles the coloring of the Pallas's. This fairly large species measures in length with a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
of . Females are generally reported to weigh , with this sample of nine averaging , and are slightly larger than males at , in ten that weighed an average of . However, in some cases Pallas's fish eagles have been reported to weigh as much as and span as much as much as .Steele, M. (2017). ''Where in the World are Pallas's Fish Eagles? Migration and Ecology of Haliaeetus leucoryphus in Asia''. University of Arkansas. Thus, their size falls just slightly under the large northern sea eagles (i.e.
bald Hair loss, also known as alopecia or baldness, refers to a loss of hair from part of the head or body. Typically at least the head is involved. The severity of hair loss can vary from a small area to the entire body. Inflammation or scarrin ...
, white-tailed and Steller's) and broadly similar to slightly larger than the sea eagles of more tropical central distribution.


Behaviour and ecology


Diet

Its diet consists primarily of large freshwater
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
. However, Pallas's fish eagle seems to have a broadly opportunistic diet much like better known sea eagles. They also regularly prey upon
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, especially
water bird A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from seabi ...
s. Other prey reportedly can include
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s, including
leporid Leporidae is the family of rabbits and hares, containing over 60 species of extant mammals in all. The Latin word ''Leporidae'' means "those that resemble ''lepus''" (hare). Together with the pikas, the Leporidae constitute the mammalian order ...
s and
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s,
frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s,
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...
s, including
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
s and
terrapin Terrapins are one of several small species of turtle (order Testudines) living in fresh or brackish water. Terrapins do not form a taxonomic unit and may not be closely related. Many belong to the families Geoemydidae and Emydidae. The name ...
s,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s (i.e.
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
larvae) and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
.Orta, J., D. A. Christie, G. M. Kirwan, and C.J. Sharpe (2020). ''Pallas's Fish-Eagle (Haliaeetus leucoryphus)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. Their prey in
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...
were reported to consist of
mrigal carp The mrigal carp (''Cirrhinus cirrhosus''), ( bn, মৃগেল, translit=mrigél) also known as the white carp, is a species of ray-finned fish in the carp family. Native to streams and rivers in India, the only surviving wild population is in ...
,
rohu The rohu, rui, ruhi or roho labeo (''Labeo rohita'') is a species of fish of the carp family, found in rivers in South Asia. It is a large omnivore and extensively used in aquaculture. Description The rohu is a large, silver-colored fish o ...
, catla,
ilish The ilish (''Tenualosa ilisha'') ( bn, ইলিশ, translit=iliś; also known as the ilishi, hilsa, hilsa herring or hilsa shad, is a species of fish related to the herring, in the family Clupeidae. It is a very popular and sought-after food ...
, ''
Chitala chitala ''Chitala chitala'' (Assamese: চিতল ''sitawl'', Bengali: চিতল, ''chitol'') is a knifefish from Bangladesh, India, Nepal, and Pakistan, found in the Brahmaputra, Indus, Ganges and Mahanadi River basins. It is sometimes known as ...
'', ''
Mystus cavasius ''Mystus cavasius'', the Gangetic mystus, is a species of catfish of the family Bagridae. In the wild it is found in Indian Subcontinent countries such as, India, Bangladesh, Pakistan, Sri Lanka, Nepal, and Myanmar. Reports of this species ...
'', ''
Ompok bimaculatus ''Ompok bimaculatus'', known as butter catfish, is a species of sheatfishes native to Asian countries such as Bangladesh, India, Pakistan, and Sri Lanka, but recently identified in Myanmar. Known in Thailand Thailand ( ), historicall ...
'' and '' Wallago attu''. Known water bird prey can include
pheasant-tailed jacana The pheasant-tailed jacana (''Hydrophasianus chirurgus'') is a jacana in the monotypic genus ''Hydrophasianus''. Like all other jacanas, they have elongated toes and nails that enable them to walk on floating vegetation in shallow lakes, their p ...
,
lesser whistling duck The lesser whistling duck (''Dendrocygna javanica''), also known as Indian whistling duck or lesser whistling teal, is a species of whistling duck that breeds in the Indian subcontinent and Southeast Asia. They are nocturnal feeders that during ...
,
ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from Greek '' aithuia'' an unidentified seabir ...
,
common pochard The common pochard (; ''Aythya ferina'') is a medium-sized diving duck. The scientific name is derived from Greek '' aithuia'', an unidentified seabird mentioned by authors including Hesychius and Aristotle, and Latin ''ferina'', "wild game", ...
and
little grebe The little grebe (''Tachybaptus ruficollis''), also known as dabchick, is a member of the grebe family of water birds. The genus name is from Ancient Greek ''takhus'' "fast" and ''bapto'' "to sink under". The specific ''ruficollis'' is from Lati ...
. Pallas's fish eagles are known to attack quite large (i.e. about their own size or somewhat larger) adult water birds including
greylag geese The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus ''Anser (bird), Anser''. It has mottled and barred grey and white plumage and an orange beak and ...
,
bar-headed geese The bar-headed goose (''Anser indicus'') is a goose that breeds in Central Asia in colonies of thousands near mountain lakes and winters in South Asia, as far south as peninsular India. It lays three to eight eggs at a time in a ground nest. It ...
and
Demoiselle crane The demoiselle crane (''Grus virgo'') is a species of crane found in central Eurosiberia, ranging from the Black Sea to Mongolia and North Eastern China. There is also a small breeding population in Turkey. These cranes are migratory birds. Bir ...
by assaulting them on the surface of the water and then flying off with the kill. Since the greylag goose species is slightly heavier than the eagle, this is one of the greatest weight-lifting feats ever recorded for a flying bird. Another case of lifting a great load was recorded at the
Yamuna The Yamuna ( Hindustani: ), also spelt Jumna, is the second-largest tributary river of the Ganges by discharge and the longest tributary in India. Originating from the Yamunotri Glacier at a height of about on the southwestern slopes of B ...
River in north-central India, where an eagle captured a huge
carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
and flew with the struggling fish very low over the water, before dropping it in response to gunfire. The carp was found to have weighed , which is probably roughly twice the weight of the eagle carrying it. How they interact with other raptors is not well understood. However, observations of wintering Pallas's fish eagles in Bharatpur suggest they may dominate several other eagle species, select the highest perches and have the highest daily food intake, including over similarly sized eagles such as
eastern imperial eagle The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South ...
s and
steppe eagle The steppe eagle (''Aquila nipalensis'') is a large bird of prey. Like all eagles, it belongs to the family Accipitridae. The steppe eagle's well-feathered legs illustrate it to be a member of the subfamily Aquilinae, also known as the "booted ...
s.


Taxonomy

''Aquila leucorypha'' was the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
proposed by
Peter Simon Pallas Peter Simon Pallas FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son of Professor of Surgery ...
in 1771, who first described a Pallas's fish eagle that he encountered during his travels in eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
. In the 19th century, it was also placed in the genera '' Falco'' and ''Haliaeetus'' by different authors. This species is the hardest-to-place sea-eagle. Among the
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, it has no close living relatives.
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
cytochrome ''b''
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called ...
data is unable to reliably suggest a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
place for it among the sea-eagles. However, some information can be drawn from the molecular data, and especially from morphology and
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
: This species retains the ancestral dark eye,
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pla ...
, and talons of the first sea-eagles, shared with the older
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
lineage. It is peculiar insofar as it has a black band at the end of the tail in adult birds, similar to juvenile
Madagascar fish eagle The Madagascar fish eagle (''Haliaeetus vociferoides'') or Madagascar sea-eagle (to distinguish it from the ''Ichthyophaga'' fishing-eagles), is a large bird of prey in the family Accipitridae which also includes many other diurnal raptors such ...
s (which look like a smaller, darker version of this bird, but are not very closely related). Its distribution indicates that this species evolved fairly independently of other sea-eagle lineages, but the molecular data tentatively suggests it is possibly closer to the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
species. It diverged from its common ancestor with other species soon after the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
and the tropical lineages split. Dependent on the interpretation of a possible Early
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but t ...
sea-eagle fossil from
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...
, this happened either at the very start or the end of the Oligocene, somewhere between 34 and 25 mya.


Distribution and habitat

The Pallas's fish eagle occurs in Russia, Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan, Mongolia, China,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
and
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
.


Conservation

The Pallas's fish eagle is listed as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
. The global population is estimated at less than 2,500 individuals. Besides direct persecution, humans contribute to the decline of this species through habitat degradation,
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, th ...
, and draining or
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in t ...
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
s. In India, the spread of
water hyacinth ''Pontederia crassipes'' (formerly ''Eichhornia crassipes''), commonly known as common water hyacinth is an aquatic plant native to South America, naturalized throughout the world, and often invasive outside its native range.prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
difficult for the Pallas's fish eagle. Its large range is deceptive, as Pallas's fish eagle is rare and isolated throughout its territory and may not breed in large areas of it.


References


External links

* * {{Taxonbar, from=Q271462
Pallas's fish eagle Pallas's fish eagle (''Haliaeetus leucoryphus''), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish sea eagle. It breeds in the east Palearctic in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, C ...
Birds of prey of Asia Birds of Bangladesh Birds of Myanmar
Pallas's fish eagle Pallas's fish eagle (''Haliaeetus leucoryphus''), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish sea eagle. It breeds in the east Palearctic in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, C ...