Mycelium
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Mycelium (plural mycelia) is a root-like structure of a
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
consisting of a mass of branching, thread-like hyphae. Fungal colonies composed of mycelium are found in and on
soil Soil, also commonly referred to as earth or dirt Dirt is an unclean matter, especially when in contact with a person's clothes, skin, or possessions. In such cases, they are said to become dirty. Common types of dirt include: * Debri ...
and many other substrates. A typical single spore germinates into a monokaryotic mycelium, which cannot reproduce sexually; when two compatible monokaryotic mycelia join and form a dikaryotic mycelium, that mycelium may form
fruiting bodies The sporocarp (also known as fruiting body, fruit body or fruitbody) of fungi is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne. The fruitbody is part of the sexual phase of a fungal life cyc ...
such as
mushroom A mushroom or toadstool is the fleshy, spore-bearing fruiting body of a fungus, typically produced above ground, on soil, or on its food source. ''Toadstool'' generally denotes one poisonous to humans. The standard for the name "mushroom" is ...
s. A mycelium may be minute, forming a colony that is too small to see, or may grow to span thousands of acres as in ''
Armillaria ''Armillaria'' is a genus of fungi that includes the ''Armillaria mellea, A. mellea'' species known as honey fungi that live on trees and woody shrubs. It includes about 10 species formerly categorized summarily as ''A. mellea''. ''Armill ...
''. Through the mycelium, a fungus absorbs nutrients from its environment. It does this in a two-stage process. First, the hyphae secrete
enzyme Enzymes () are proteins that act as biological catalysts by accelerating chemical reactions. The molecules upon which enzymes may act are called substrates, and the enzyme converts the substrates into different molecules known as products ...
s onto or into the food source, which break down biological polymers into smaller units such as
monomers In chemistry, a monomer ( ; ''mono-'', "one" + '' -mer'', "part") is a molecule that can react together with other monomer molecules to form a larger polymer chain or three-dimensional network in a process called polymerization. Classification Mo ...
. These monomers are then absorbed into the mycelium by facilitated diffusion and
active transport In cellular biology, ''active transport'' is the movement of molecules or ions across a cell membrane from a region of lower concentration to a region of higher concentration—against the concentration gradient. Active transport requires cellul ...
. Mycelia are vital in terrestrial and aquatic
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
s for their role in the
decomposition Decomposition or rot is the process by which dead organic substances are broken down into simpler organic or inorganic matter such as carbon dioxide, water, simple sugars and mineral salts. The process is a part of the nutrient cycle and is e ...
of plant material. They contribute to the organic fraction of soil, and their growth releases
carbon dioxide Carbon dioxide ( chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is trans ...
back into the atmosphere (see
carbon cycle The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of the Earth. Carbon is the main component of biological compounds as well as a major componen ...
).
Ectomycorrhizal extramatrical mycelium Ectomycorrhizal extramatrical mycelium (also known as extraradical mycelium) is the collection of filamentous fungal hyphae emanating from ectomycorrhizas. It may be composed of fine, hydrophilic hypha which branches frequently to explore and expl ...
, as well as the mycelium of
arbuscular mycorrhizal fungi An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'', a.k.a. ''endomycorrhiza'') is a type of mycorrhiza in which the symbiont fungus (''AM fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. ...
, increase the efficiency of water and nutrient absorption of most plants and confers resistance to some plant pathogens. Mycelium is an important food source for many soil invertebrates. They are vital to
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
and are important to almost all species of
plants Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude ...
, many species co-evolving with the
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
. Mycelium is a primary factor in a plant's health, nutrient intake, and growth, with mycelium being a major factor to plant fitness. Networks of mycelia can transport water and spikes of electrical potential. "Mycelium", like "fungus", can be considered a mass noun, a word that can be either singular or plural. The term "mycelia", though, like "fungi", is often used as the preferred plural form.
Sclerotia A sclerotium (; (), is a compact mass of hardened fungal mycelium containing food reserves. One role of sclerotia is to survive environmental extremes. In some higher fungi such as ergot, sclerotia become detached and remain dormant until favor ...
are compact or hard masses of mycelium.


Uses

One of the primary roles of fungi in an ecosystem is to
decompose Decomposition or rot is the process by which dead organic substances are broken down into simpler organic or inorganic matter such as carbon dioxide, water, simple sugars and mineral salts. The process is a part of the nutrient cycle and is ...
organic compounds.
Petroleum Petroleum, also known as crude oil, or simply oil, is a naturally occurring yellowish-black liquid mixture of mainly hydrocarbons, and is found in geological formations. The name ''petroleum'' covers both naturally occurring unprocessed crud ...
products and some pesticides (typical soil contaminants) are organic molecules (i.e., they are built on a carbon structure), and thereby show a potential carbon source for fungi. Hence, fungi have the potential to eradicate such pollutants from their environment unless the chemicals prove toxic to the fungus. This biological degradation is a process known as
bioremediation Bioremediation broadly refers to any process wherein a biological system (typically bacteria, microalgae, fungi, and plants), living or dead, is employed for removing environmental pollutants from air, water, soil, flue gasses, industrial effluent ...
. Mycelial mats have been suggested as having potential as biological filters, removing chemicals and microorganisms from soil and water. The use of fungal mycelium to accomplish this has been termed mycofiltration. Knowledge of the relationship between mycorrhizal fungi and plants suggests new ways to improve crop yields. When spread on logging roads, mycelium can act as a binder, holding disturbed new soil in place preventing washouts until
woody plants A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until sp ...
can establish roots. Alternatives to polystyrene and plastic packaging can be produced by growing mycelium in agricultural waste. Mycelium has also been used as a material in furniture, bricks, and
artificial leather Artificial leather, also called synthetic leather, is a material intended to substitute for leather in upholstery, clothing, footwear, and other uses where a leather-like finish is desired but the actual material is cost prohibitive or unsuitab ...
. Fungi are essential for converting biomass into compost, as they decompose feedstock components such as lignin, which many other composting microorganisms cannot. Turning a backyard compost pile will commonly expose visible networks of mycelia that have formed on the decaying organic material within. Compost is an essential soil amendment and
fertilizer A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...
for
organic farming Organic farming, also known as ecological farming or biological farming,Labelling, article 30 o''Regulation (EU) 2018/848 of the European Parliament and of the Council of 30 May 2018 on organic production and labelling of organic products and re ...
and gardening. Composting can divert a substantial fraction of
municipal solid waste Municipal solid waste (MSW), commonly known as trash or garbage in the United States and rubbish in Britain, is a waste type consisting of everyday items that are discarded by the public. "Garbage" can also refer specifically to food waste ...
from landfills.


See also

* *


References


External links


Mycelium
at anbg.gov.au {{Fungus structure Fungal morphology and anatomy