Mauer 1
   HOME

TheInfoList



OR:

The Mauer 1
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
is the oldest-known specimen of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely rela ...
'' in
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwee ...
. It was found in 1907 in a sand quarry in the community Mauer, around south-east of
Heidelberg Heidelberg (; Palatine German: ') is a city in the German state of Baden-Württemberg, situated on the river Neckar in south-west Germany. As of the 2016 census, its population was 159,914, of which roughly a quarter consisted of students ...
. The Mauer 1 mandible is the type specimen of the species ''
Homo heidelbergensis ''Homo heidelbergensis'' (also ''H. sapiens heidelbergensis''), sometimes called Heidelbergs, is an extinct species or subspecies of archaic human which existed during the Middle Pleistocene. It was subsumed as a subspecies of '' H. erectus'' i ...
''. Some European researchers have classified the find as ''Homo erectus heidelbergensis'', regarding it as a subspecies of ''
Homo erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor ...
''. In 2010 the mandible's age was for the first time exactly determined to be 609,000 ± 40,000 years.Günther A. Wagner et al.: ''Radiometric dating of the type-site for ''Homo heidelbergensis'' at Mauer, Germany.'' In: ''
Proceedings of the National Academy of Sciences ''Proceedings of the National Academy of Sciences of the United States of America'' (often abbreviated ''PNAS'' or ''PNAS USA'') is a peer-reviewed multidisciplinary scientific journal. It is the official journal of the National Academy of S ...
.'' vol. 107, no. 46, 2010, pp. 19726–19730 .
Previously, specialist literature had referred to an age of either 600,000 or 500,000 years on the basis of less accurate dating methods.


Discovery

On October 21, 1907, Daniel Hartmann, a worker at a sand mine in the Grafenrain open-field system of the Mauer community unearthed a mandible at a depth of , which he recognized as of human origin.Schoetensack, p. 23. He was aware of the likelihood of finds, as for 20 years the Heidelberg scholar
Otto Schoetensack Otto Karl Friedrich Schoetensack (; 12 July 1850 in Stendal – 23 December 1912 in Ospedaletti) was a German industrialist and later professor of anthropology, having retired from the chemical firm which he had founded. During a 1908 archeologic ...
had asked that the workers at the sand mine be encouraged to look out for fossils, after the well-preserved skull of a
straight-tusked elephant The straight-tusked elephant (''Palaeoloxodon antiquus'') is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene (781,000–30,000 years before present). Recovered individuals have re ...
had come to light there in 1887. Schoetensack had the workers taught the characteristics of human bones based on recent examples on his regular visits to the sand mine in search for "traces of mankind". As it was dug out, the mandible was flung in the air and only discovered after it had broken into two parts. A piece of the left side of the mandible broke off in the process and was never found. A thick cemented crust of coarse sand stuck on and around the canines and molars—a characteristic of many of the Mauer fossils. The cementing had been caused by carbonation of calcium. A chunk of
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms w ...
, probably
Muschelkalk The Muschelkalk (German for "shell-bearing limestone"; french: calcaire coquillier) is a sequence of sedimentary rock strata (a lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 million ...
, long and about wide was firmly stuck to the sand crust on top of the
bicuspid The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mout ...
s and the two frontal
molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
on the left side of the mandible. The contractor at the sand mine immediately reported the discovery to Schoetensack, who examined and documented the site and the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
. He presented the results of his studies in autumn the following year in a
monograph A monograph is a specialist work of writing (in contrast to reference works) or exhibition on a single subject or an aspect of a subject, often by a single author or artist, and usually on a scholarly subject. In library cataloging, ''monogra ...
titled: "The lower jaw of ''Homo heidelbergensis'' from the sands of Mauer near Heidelberg". On November 19, 1907, Schoetensack stated in a legal document that mine contractor Josef Rösch had given the specimen to the
Heidelberg University } Heidelberg University, officially the Ruprecht Karl University of Heidelberg, (german: Ruprecht-Karls-Universität Heidelberg; la, Universitas Ruperto Carola Heidelbergensis) is a public research university in Heidelberg, Baden-Württemberg, ...
as a gift. The mandible remains in the university's Geological-Palaeontological Institute to this day as "the most valuable object in the natural history collections of the University of Heidelberg". Further finds from the Mauer sand mine are the ''Hornstein artefacts'', unearthed in 1924 by Karl Friedrich Hormuth, which scholars interpreted as tools of ''Homo heidelbergensis''. In 1933 Wilhelm Freudenberg discovered a
frontal bone The frontal bone is a bone in the human skull. The bone consists of two portions.'' Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, ...
fragment which has also been associated with ''Homo heidelbergensis''.


Description of the fossil

The anatomical analysis of the lower jaw of Mauer in its 1908 original
species description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have ...
by Otto Schoetensack was based largely on the expertise of Breslau professor
Hermann Klaatsch Hermann Klaatsch (10 March 1863 – 5 January 1916) was a German physician, anatomist, physical anthropologist, evolutionist, and professor at the University of Heidelberg from 1890, and at the University of Breslau (Wrocław) until 1916.
, which was only hinted at in a brief acknowledgement in the preface. In his original species description Schoetensack wrote that the "nature of our object" reveals itself "at first sight" since "a certain disproportion between the jaw and the teeth" is obvious: "The teeth are too small for the bone. The available space would allow for a far greater flexibility of development." And further, on the find: "It shows a combination of features, that has been previously found neither on a recent nor on a fossilized human mandible. Even the scholar should not be blamed if he would only reluctantly accept it as human: Entirely missing is the one feature, which is regarded as particularly human, namely an outer projection of the chin portion, yet this deficiency is found to be combined with extremely strange dimensions of the mandibular body. The actual proof that we are dealing with human remains here only lies within the nature of the
dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
. The completely preserved teeth bear the stamp ''human'' as evidence: The canines show no trace of a stronger expression in relation to the other groups of teeth. They suggest a moderate and harmonious co-evolution, as it is the case in recent humans." The characteristics of the lower jaw are therefore the lack of a chin on the one hand and on the other it is the considerable size of the lower jaw bone, on which, behind the
wisdom tooth A third molar, commonly called wisdom tooth, is one of the three molars per quadrant of the human dentition. It is the most posterior of the three. The age at which wisdom teeth come through ( erupt) is variable, but this generally occurs betw ...
a fourth
premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth ...
would easily have had space to develop. Since the third molar (the wisdom tooth) is present and its
dentin Dentin () (American English) or dentine ( or ) (British English) ( la, substantia eburnea) is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by e ...
exposed—although only in a few places—the age of death is estimated to be about 20 to 30 years. Schoetensack concluded a relationship to modern man (''Homo sapiens'') from the similarity of the dentition and put the lower jaw in the genus ''Homo''—a view that is still being held unanimously by today's palaeo-anthropologists. He derived the authority to define a new species with the type-epithet heidelbergensis from the fact that the lower jaw—in contrast to modern humans—is missing its chin. With the subtitle of his original description—"A contribution to the
paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
of the human species"—Schoetensack explicitly takes a clear position on the part of
Darwinism Darwinism is a theory of biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others, stating that all species of organisms arise and develop through the natural selection of small, inherited variations tha ...
"in the great debate on the origin of man, namely, that humans have evolved from the animal kingdom and are not the product of a singular act of creation." As to the lower jaw of Mauer's precise position in the ancestral chain of modern man Schoetensack expressed only cautious statements. Reluctantly he wrote in his study that "it seems possible that ''Homo heidelbergensis'' belongs in the ancestral series of the European man" and—after meticulous and detailed comparison with other European
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s he stated equally vague: "We must therefore denote the mandible of ''Homo heidelbergensis'' as pre-
neandertal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While the ...
oid." The classification of the lower jaw of Mauer in the time before the
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
s proved to be accurate. Schoetensack—like many of his colleagues around the beginning of the 20th century—was wrong with his assessment of kinship proximity of the lower jaw of Mauer with the apes (
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ...
s): "The mandible of ''Homo heidelbergensis'' reveals the original state that defines mankind's and the ape's common ancestor." In 1924, the hitherto oldest fossil of the big pool of hominid variants—the
Taung Child The Taung Child (or Taung Baby) is the fossilised skull of a young '' Australopithecus africanus''. It was discovered in 1924 by quarrymen working for the Northern Lime Company in Taung, South Africa. Raymond Dart described it as a new specie ...
was discovered in what is now
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun ...
. It is around two million years older than the lower jaw of Mauer and, despite its advanced age, still does not represent the common base of humans and apes.


Dating

Otto Schoetensack had the site at the bottom of the sand mine marked with a commemorative stone on which a horizontal line represented the level of the find.Schoetensack, S. 4. Whether his wish had been granted—namely that the stone may remain in its place, even if the sand mine will one day be re-filled—is unknown. In fact, the part of the mine, in which the lower jaw came to light, was filled with overburden in the 1930s, later re-natured as arable land and declared a nature reserve in 1982. The actual site is therefore not accessible for research at the present time. Absolute dating of the strata using contemporary scientific methods proved to be not possible. Alternatively scientists have repeatedly tried to deduce the age of the fossil's layer, using
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostra ...
methods. Schoetensack had described the merely ten-centimeter-thick layer of the find as "layer of gravel (
scree Scree is a collection of broken rock fragments at the base of a cliff or other steep rocky mass that has accumulated through periodic rockfall. Landforms associated with these materials are often called talus deposits. Talus deposits typically ...
), slightly cemented due to carbonation of calcium, with very thin layers of
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay pa ...
(Letten), that faintly reacts with
hydrochloric acid Hydrochloric acid, also known as muriatic acid, is an aqueous solution of hydrogen chloride. It is a colorless solution with a distinctive pungent smell. It is classified as a strong acid. It is a component of the gastric acid in the dige ...
". Above and below of the finding's strata, sands and other materials had accumulated in various definable layers. These had been deposited on the edge of a former
Neckar The Neckar () is a river in Germany, mainly flowing through the southwestern state of Baden-Württemberg, with a short section through Hesse. The Neckar is a major right tributary of the Rhine. Rising in the Schwarzwald-Baar-Kreis near Schwen ...
River arc over the course of millennia. In the preface to his study he states: "The age of these sands is commonly agreed to as paleo- diluvial (altdiluvial) by reference to embedded
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
remains; although certain species suggest a significant relationship with a more recent section of the
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
, namely the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58commemorative publication complained that there "still existed no satisfactory exact data for the determination of the geological age of the lower jaw of ''Homo heidelbergensis''". After all, in 1995 the age of the sands of Mauer 1 could approximately be determined on the basis of small fossils. Additionally, attempts were made to perform an absolute dating in yet accessible, adjacent sand mines. However, to date researchers could not agree on which of the several possible layers that belong to the Cromer-warm period is identical with the layer of the Grafenrain mine. So it happens that the commune of Mauer dates the fossil to an age "of more than 600,000 years" on its website, whereas the memorial stone called an age of 500,000 years. A range from 474,000 to 621,000 years is currently considered to be the established age of "layer 4", in which the actual fossil either originates from the lower (around 600,000) or the upper (around 500,000) spectrum. In November 2010, the final dating of sand grains via
infrared Infrared (IR), sometimes called infrared light, is electromagnetic radiation (EMR) with wavelengths longer than those of Light, visible light. It is therefore invisible to the human eye. IR is generally understood to encompass wavelengths from ...
-radio fluorescence (IR-RF) and the dating of the teeth via a combined electron-spin and
uranium–lead dating Uranium–lead dating, abbreviated U–Pb dating, is one of the oldest and most refined of the radiometric dating schemes. It can be used to date rocks that formed and crystallised from about 1 million years to over 4.5 billion years ago with routi ...
was published in the ''
Proceedings of the National Academy of Sciences ''Proceedings of the National Academy of Sciences of the United States of America'' (often abbreviated ''PNAS'' or ''PNAS USA'') is a peer-reviewed multidisciplinary scientific journal. It is the official journal of the National Academy of S ...
'' wherein the age of the fossil was concluded as to be of 609,000 ± 40,000 years.


Relation to modern humans

The Mauer mandible is the type specimen of the species ''Homo heidelbergensis''. "The anatomy is clearly more primitive than that of Neanderthal, but the harmoniously rounded dental arch and the complete row of teeth...already typically human." Based on these circumstances—the chronological delineation from the more recent Neanderthal on the one hand, and from older fossils, denoted as ''Homo erectus'' on the other—today's researchers consider it justifiable to declare Mauer 1 as an independent
chronospecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
. According to
Chris Stringer Christopher Brian Stringer (born 1947) is a British physical anthropologist noted for his work on human evolution. Biography Growing up in a working-class family in the East End of London, Stringer's interest in anthropology began in primar ...
, ''Homo heidelbergensis'' ranks separable between earlier ''Homo erectus'' and the more recent Neanderthal and ''Homo sapiens''; it is from this point of view the last common ancestor of Neanderthal and anatomically modern man. Other researchers hold the contrary view that the evolutionary development in Africa and Europe was a gradual process from ''Homo erectus'' via the representative of the findings assigned to ''Homo heidelbergensis'' towards Neanderthal. Any form of segregation is considered arbitrary, which is why these researchers forgo the name ''Homo heidelbergensis'' altogether. They classify the Mauer 1 man as a late local (European) form of ''Homo erectus''. However, there is agreement among all paleo-anthropologists that the lower jaw of Mauer does not belong to the immediate ancestral line of modern man. He is regarded rather as a descendant of an early migration to Europe and Asia (depending on the terminology—of ''Homo erectus'' or ''Homo heidelbergensis''), whose oldest fossils outside of Africa are about 1.8 million years old. The last descendant of this first migration to Europe was Neanderthal, who became extinct about 30,000 years ago. Members of the species ''Homo sapiens'' arrived in Europe only in a second migratory wave of the genus ''Homo'' at around 40,000 to 30,000 years ago, whose descendants include modern man.


Habitat

As uncertain as the exact dating of the lower jaw of Mauer was until recently, so is the assignment of other fossils to its layer. Such contextual fossils are the only direct evidence for an accurate reconstruction of the
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
of a find. Not until 1991 two sets of core drilling were conducted in the defunct sand mine of Grafenrain and—beginning in 1995, several dozen cubic meters of sand were screened in search of fossil fragments that could shed light on accompanying species. However, mice teeth that were found proved to be not suitable for a more precise dating of the layer, since these mice lived anatomically virtually unchanged over too long a period. At least
palynological Palynology is the "study of dust" (from grc-gre, παλύνω, palynō, "strew, sprinkle" and ''-logy'') or of "particles that are strewn". A classic palynologist analyses particulate samples collected from the air, from water, or from deposit ...
findings of similar vegetation areas of the Cromer
interglacial An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene i ...
period can help with a description of the habitat.: "across alluvial forests in the river valleys, forests on the slopes and open forests on the heights; these were rather dry locations, regarding the hydro-geology of the crevasses (german: Kluftwassersystem) of the
Buntsandstein The Buntsandstein (German for ''coloured'' or ''colourful sandstone'') or Bunter sandstone is a lithostratigraphic and allostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Buntsandst ...
—and
Muschelkalk The Muschelkalk (German for "shell-bearing limestone"; french: calcaire coquillier) is a sequence of sedimentary rock strata (a lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 million ...
mountains, bare of any
Loess Loess (, ; from german: Löss ) is a clastic, predominantly silt-sized sediment that is formed by the accumulation of wind-blown dust. Ten percent of Earth's land area is covered by loess or similar deposits. Loess is a periglacial or aeoli ...
sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand ...
s." Animal fossils from different layers of the sand mine Grafenrain that were clearly identified and belonged to the same interglacial epoch as the find layer, inspired the author of a 2007 article of ''
Die Zeit ''Die Zeit'' (, "The Time") is a German national weekly newspaper published in Hamburg in Germany. The newspaper is generally considered to be among the German newspapers of record and is known for its long and extensive articles. History Th ...
'' to a vivid
diorama A diorama is a replica of a scene, typically a three-dimensional full-size or miniature model, sometimes enclosed in a glass showcase for a museum. Dioramas are often built by hobbyists as part of related hobbies such as military vehicle mode ...
: : "...among spruce, birch and oak trees romped flying squirrels, roamed roe deer, elk and wild boar. Mole and shrew tunnelled through the grounds. And: Beavers damming the waters of the juvenile
Neckar The Neckar () is a river in Germany, mainly flowing through the southwestern state of Baden-Württemberg, with a short section through Hesse. The Neckar is a major right tributary of the Rhine. Rising in the Schwarzwald-Baar-Kreis near Schwen ...
river. Hares flitted and horses galloped across open landscapes. Theoretically, nature provided steaks of elephant, the woolly rhinoceros and hippopotamus. Whether the "Heidelberger" ventured on such prey, must be doubted. Almost certainly he took to his heels in front of bear, wolf, leopard, saber-toothed cat and hyena."Urs Willmann: ''Der multiple Adam.'' In: ''
Die Zeit ''Die Zeit'' (, "The Time") is a German national weekly newspaper published in Hamburg in Germany. The newspaper is generally considered to be among the German newspapers of record and is known for its long and extensive articles. History Th ...
.'' Nr. 43 vom 18. Oktober 2007, S. 43
Volltext
/ref>


Images of the original species description

File:Homo heidelbergensis (Erstbeschreibung) 01.jpg, Title page of the first description of ''Homo heidelbergensis'' File:Homo heidelbergensis (Erstbeschreibung) 02.jpg, Original condition with adherent material File:Homo heidelbergensis (Erstbeschreibung) 03.jpg, Condition after cleaning File:Homo heidelbergensis (Erstbeschreibung) 04.jpg, Radiography File:Homo heidelbergensis (Erstbeschreibung) 05.jpg, Inferior - and occlusal view


Bibliography

* Otto Schoetensack: ''Der Unterkiefer des Homo Heidelbergensis aus den Sanden von Mauer bei Heidelberg. Ein Beitrag zur Paläontologie des Menschen''. Verlag von Wilhelm Engelmann, Leipzig 1908
Complete text
. * Alfried Wieczorek,
Wilfried Rosendahl Wilfried Rosendahl (born 1966) is a German bioarchaeologist, geoscientist, and cultural manager, general director of the Reiss-Engelhorn-Museums in Mannheim, and honorary professor at the Institute of History at the University of Mannheim. ...
(Hrsg.): ''MenschenZeit. Geschichten vom Aufbruch der frühen Menschen''. Philipp von Zabern, Mainz 2003, (Katalog zur gleichnamigen Ausstellung der Reiss-Engelhorn-Museen in Mannheim). * Günther A. Wagner, Hermann Rieder, Ludwig Zöller, Erich Mick (Hrsg.): ''Homo heidelbergensis. Schlüsselfund der Menschheitsgeschichte''. Konrad Theiss Verlag, Stuttgart 2007, . * Katerina Harvati: ''100 years of Homo heidelbergensis – life and times of a controversial taxon.'' In: ''Mitteilungen der Gesellschaft für Urgeschichte'' 16, 2007, 85-9
PDF


See also

*
Altamura Man The Altamura Man is a fossil of the genus ''Homo'' discovered in 1993 in a karst sinkhole in the Lamalunga Cave near the city of Altamura, Italy. Remarkably well preserved but covered in a thick layer of calcite taking the shape of cave popcorn ...
*
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of t ...
*
List of human evolution fossils The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages) in the late Miocene, roug ...
*
Neanderthal 1 Feldhofer 1 or Neanderthal 1 is the scientific name of the 40,000-year-old type specimen fossil of the species ''Homo neanderthalensis'', found in August 1856 in a German cave, the Kleine Feldhofer Grotte in the Neandertal valley, east of D ...
*
Saldanha man __NOTOC__ Saldanha man also known as Saldanha cranium or Elandsfontein cranium are fossilized remains of an archaic human. It is one of the key specimens for ''Homo heidelbergensis''. It has not been dated directly, and is estimated to be roughly ...


References


External links

* {{Homo heidelbergensis, state=collapsed Homo heidelbergensis fossils 1907 archaeological discoveries Archaeological discoveries in Germany