Marabou stork
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The marabou stork (''Leptoptilos crumenifer'') is a large wading
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the stork family Ciconiidae native to sub-Saharan Africa. It breeds in both wet and arid habitats, often near human habitation, especially landfill sites. It is sometimes called the "
undertaker A funeral director, also known as an undertaker (British English) or mortician (American English), is a professional involved in the business of funeral rites. These tasks often entail the embalming and burial or cremation of the dead, as ...
bird" due to its shape from behind: cloak-like wings and back, skinny white legs, and sometimes a large white mass of "hair".


Taxonomy

The marabou stork was formally described in 1831 by the French naturalist René Lesson. He placed it in the stork genus '' Ciconia'' and coined the binomial name ''Ciconia crumenifera''. He specified that
locality Locality may refer to: * Locality (association), an association of community regeneration organizations in England * Locality (linguistics) * Locality (settlement) * Suburbs and localities (Australia), in which a locality is a geographic subdivis ...
as Senegal. The species is now placed with the
lesser adjutant The lesser adjutant (''Leptoptilos javanicus'') is a large wading bird in the stork family Ciconiidae. Like other members of its genus, it has a bare neck and head. It is however more closely associated with wetland habitats where it is solitary ...
and the greater adjutant in the genus '' Leptoptilos'' that Lesson had introduced at the same time he described the marabou stork. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
: no subspecies are recognised. The common name marabou is thought to be derived from the Arabic word '' murābit'' meaning quiet or hermit-like. The species was originally described as ''Ciconia crumenifera''. When the species was moved into the genus ''Leptoptilos'', the ending was modified to ''crumeniferus'' and this was used by many authors until it was noted that the correct masculine ending to match the genus is ''crumenifer''.


Description

The marabou stork is a massive bird: large specimens are thought to reach a height of and a weight of . A wingspan of was accepted by Fisher and Peterson, who ranked the species as having the largest wing-spread of any living bird. Even higher measurements of up to have been reported, although no measurement over has been verified. It is often credited with the largest spread of any landbird, to rival the
Andean condor The Andean condor (''Vultur gryphus'') is a giant South American Cathartid vulture and is the only member of the genus ''Vultur''. Found in the Andes mountains and adjacent Pacific coasts of western South America, the Andean condor is the larg ...
; more typically, however, these storks measure across the wings, which is about a foot less than the average Andean condor wingspan and nearly two feet less than the average of the largest albatrosses and pelicans. Typical weight is , unusually as low as , and length (from bill to tail) is . Females are smaller than males. Bill length can range from . Unlike most storks, the three '' Leptoptilos'' species fly with the neck retracted like a heron. The marabou is unmistakable due to its size, bare head and neck, black back, and white underparts. It has a huge bill, a pink
gular sac Gular skin (throat skin), in ornithology, is an area of featherless skin on birds that joins the lower mandible of the beak (or ''bill'') to the bird's neck. Other vertebrate taxa may have a comparable anatomical structure that is referred to as e ...
at its throat (''crumenifer(us)'' means "carrier of a pouch for money"), a neck ruff, and black legs and wings. The sexes are alike, but the young bird is browner and has a smaller bill. Full maturity is not reached for up to four years.


Behavior and ecology

Like most storks, the marabou is gregarious and a colonial breeder. In the African dry season (when food is more readily available as the pools shrink), it builds a tree nest in which two or three
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
are laid. It is known to be quite ill-tempered. It also resembles other storks in that it is not very vocal, but indulges in bill-rattling
courtship display A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), ...
s. The throat sac is also used to make various noises at that time.


Breeding

The marabou stork breeds in Africa south of the Sahara. In East Africa, the birds interact with humans and breed in urban areas. In southern African countries, the birds breed mainly in less populated areas. The marabou stork breeds in colonies, starting during the dry season. The female lays two to three eggs in a small nest made of sticks; eggs hatch after an incubation period of 30 days. Their young reach sexual maturity at 4 years of age. Lifespan is 43 years in captivity and 25 years in wild.


Feeding

The marabou stork is a frequent scavenger, and the naked head and long neck are adaptations to this livelihood, as it is with the vultures with which the stork often feeds. In both cases, a feathered head would become rapidly clotted with
blood Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in the cir ...
and other substances when the bird's head was inside a large corpse, and the bare head is easier to keep clean. This large and powerful bird eats mainly carrion, scraps, and
faeces Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...
but will opportunistically eat almost any animal matter it can swallow. It occasionally eats other birds including ''
Quelea ''Quelea'' is a genus of small passerine birds that belongs to the weaver family Ploceidae, confined to Africa. These are small-sized, sparrow- or finch-like gregarious birds, with bills adapted to eating seeds. Queleas may be nomadic over vas ...
'' nestlings,
pigeon Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s,
dove Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s, pelican and cormorant chicks, and even
flamingo Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbea ...
s. During the breeding season, adults scale back on carrion and take mostly small, live prey since nestlings need this kind of food to survive. Common prey at this time may consist of fish, frogs, insects, eggs, small mammals and reptiles such as crocodile hatchlings and eggs,Hancock, Kushlan & Kahl, Storks, ''Storks, Ibises and Spoonbills of the World''. Princeton University Press (1992), and lizards and snakes. Though known to eat putrid and seemingly inedible foods, these storks may sometimes wash food in water to remove soil. When feeding on carrion, marabou frequently follow vultures, which are better equipped with hooked bills for tearing through carrion meat and may wait for the vultures to cast aside a piece, steal a piece of meat directly from the vulture or wait until the vultures are done. As with vultures, marabou storks perform an important natural function by cleaning areas via their ingestion of carrion and waste. Increasingly, marabous have become dependent on human garbage and hundreds of the huge birds can be found around African dumps or waiting for a hand out in urban areas. Marabous eating human garbage have been seen to devour virtually anything that they can swallow, including shoes and pieces of metal. Marabous conditioned to eating from human sources have been known to lash out when refused food.


Threats

Fully grown marabou storks have few natural enemies, and have high annual survival rate, though lions have reportedly preyed on some individuals in ambush. A number of endoparasites have been identified in wild marabous including ''Cheilospirura'', ''Echinura'' and ''Acuaria''
nematodes The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broa ...
, ''Amoebotaenia sphenoides'' (Cestoda) and ''Dicrocoelium hospes'' (Trematoda).


Human uses

Marabou down is frequently used in the trimming of various items of clothing and hats, as well as fishing lures.''The Oxford Pocket Dictionary of Current English 2008''
(
Oxford University Press Oxford University Press (OUP) is the university press of the University of Oxford. It is the largest university press in the world, and its printing history dates back to the 1480s. Having been officially granted the legal right to print books ...
, 2008)
Turkey down and similar feathers have been used as a substitute for making 'marabou' trimming. In
tarot The tarot (, first known as '' trionfi'' and later as ''tarocchi'' or ''tarocks'') is a pack of playing cards, used from at least the mid-15th century in various parts of Europe to play card games such as Tarocchini. From their Italian roots ...
and
tarock Tarot games are card games played with tarot decks, that is, decks with numbered permanent trumps parallel to the suit cards. The games and decks which English-speakers call by the French name Tarot are called Tarocchi in the original Italian, ...
, the Marabou forms part of "the Birds", together with the Sparrow (I of trumps),
Eagle Owl The American (North and South America) horned owls and the Old World eagle-owls make up the genus ''Bubo'', at least as traditionally described. The genus name ''Bubo'' is Latin for the Eurasian eagle-owl. This genus contains 19 species that ar ...
(II of trumps), and the
Cockatoo A cockatoo is any of the 21 parrot species belonging to the family Cacatuidae, the only family in the superfamily Cacatuoidea. Along with the Psittacoidea (true parrots) and the Strigopoidea (large New Zealand parrots), they make up the orde ...
(III of trumps); sometimes "the Birds" also include the
Dodo The dodo (''Raphus cucullatus'') is an extinct flightless bird that was endemic to the island of Mauritius, which is east of Madagascar in the Indian Ocean. The dodo's closest genetic relative was the also-extinct Rodrigues solitaire. The ...
(V of trumps).


Gallery

File:Marabou stork, Leptoptilos crumeniferus edit1.jpg,
Mikumi National Park Mikumi National Park is a national park near Morogoro, Tanzania with an area of that was established in 1964. It is the fourth largest in the country. The park is crossed by Tanzania's A-7 highway. Territory Mikumi National Park borders Selou ...
, Tanzania File:The Big Game of Africa (1910) - Giant Marabou.png, Large marabou stork,
British East Africa East Africa Protectorate (also known as British East Africa) was an area in the African Great Lakes occupying roughly the same terrain as present-day Kenya from the Indian Ocean inland to the border with Uganda in the west. Controlled by Bri ...
File:Marabou Stork (Leptoptilos crumeniferus) skeleton at the Royal Veterinary College anatomy museum.JPG, Skeleton of a marabou stork File:The Childrens Museum of Indianapolis - Marabou stork skull cast.jpg, Skull cast from a ''Leptoptilos crumenifer'' displays its long beak File:Leptoptilos crumeniferus and vultures -Masai Mara -Kenya-8.jpg, A Marabou stork and
Griffon vulture The Eurasian griffon vulture (''Gyps fulvus'') is a large Old World vulture in the bird of prey family Accipitridae. It may also be known as the Griffon vulture, though it may be used for the genus as a whole. It is not to be confused with Rü ...
s (''G. fulvus'') scavenging in the Masai Mara, Kenya File:BLW Evening Cloak.jpg, 1920 cloak trimmed with marabou feathers File:Marabou stork (Leptoptilos crumenifer) spreading wings.jpg, spreading wings,
Uganda }), is a landlocked country in East Africa. The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The sou ...
File:Marabou stork (Leptoptilos crumenifer) spreading wings 2.jpg, spreading wings, Ethiopia Marabou storks (Leptoptilos crumenifer) on elephant carcass.jpg, feeding on elephant carcass


References


External links

* *
Marabou Stork
– ''The Atlas of Southern African Birds'' {{Taxonbar, from=Q512737 marabou stork Scavengers Birds of Sub-Saharan Africa marabou stork Birds of East Africa marabou stork