Maastrichtian dialect
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The Maastrichtian () is, in the ICS
geologic timescale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochr ...
, the latest age (uppermost stage) of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
Epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided ...
or
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
Series, the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period or
System A system is a group of interacting or interrelated elements that act according to a set of rules to form a unified whole. A system, surrounded and influenced by its environment, is described by its boundaries, structure and purpose and express ...
, and of the
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretac ...
(part of the
Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning o ...
and
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pala ...
). The
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
(formerly known as the Cretaceous–
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
extinction event) occurred at the end of this age. In this
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. I ...
, many commonly recognized groups such as non-avian
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s,
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appea ...
s and
mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on ...
s, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous.


Stratigraphic definitions


Definition

The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying
rock strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
of the
Chalk Group The Chalk Group (often just called the Chalk) is the lithostratigraphic unit (a certain number of rock strata) which contains the Upper Cretaceous limestone succession in southern and eastern England. The same or similar rock sequences occur acro ...
close to the Dutch city of
Maastricht Maastricht ( , , ; li, Mestreech ; french: Maestricht ; es, Mastrique ) is a city and a municipality in the southeastern Netherlands. It is the capital and largest city of the province of Limburg. Maastricht is located on both sides of the ...
. These strata are now classified as the Maastricht Formation - both
formation Formation may refer to: Linguistics * Back-formation, the process of creating a new lexeme by removing or affixes * Word formation, the creation of a new word by adding affixes Mathematics and science * Cave formation or speleothem, a secondar ...
and stage derive their names from the city. The Maastricht Formation is known for its fossils from this age, most notably those of the giant
sea reptile Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the ...
''
Mosasaurus ''Mosasaurus'' (; "lizard of the Meuse River") is the type genus (defining example) of the mosasaurs, an extinct group of aquatic squamate reptiles. It lived from about 82 to 66 million years ago during the Campanian and Maastrichtian sta ...
'', which in turn derives its name from the nearby river Maas (''mosa'' being Latin for the river Maas). The base of the Maastrichtian Stage is at the first appearance of
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefis ...
species '' Pachydiscus neubergicus''. At the original type locality near Maastricht, the stratigraphic record was later found to be incomplete. A reference profile for the base was then appointed in a section along the Ardour river called ''Grande Carrière'', close to the village of
Tercis-les-Bains Tercis-les-Bains (; oc, Tèrcis) is a commune of the Landes department in Nouvelle-Aquitaine in southwestern France. Population See also *Communes of the Landes department The following is a list of the 327 communes of the Landes departm ...
in southwestern
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
. The top of the Maastrichtian Stage is defined to be at the
iridium Iridium is a chemical element with the symbol Ir and atomic number 77. A very hard, brittle, silvery-white transition metal of the platinum group, it is considered the second-densest naturally occurring metal (after osmium) with a density o ...
anomaly at the
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end ...
(K–Pg boundary), which is also characterised by the
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the Endling, last individual of the species, although the Functional ext ...
of many groups of life.


Subdivision

The Maastrichtian is commonly subdivided into two substages (Upper and Lower) and three ammonite biozones. The biozones are (from young to old): *zone of '' Anapachydiscus terminus'' *zone of ''Anapachydiscus fresvillensis'' *zone of '' Pachydiscus neubergicus'' till ''Pachydiscus epiplectus'' The Maastrichtian is roughly coeval with the
Lancian The Lancian was a North American faunal stage of the Late Cretaceous. It was the final stage of the Cretaceous period in North America, lasting from approximately 70.6 to 66 million years ago. Geology Terrestrial sedimentary strata from the Judit ...
North American Land Mammal Age The North American land mammal ages (NALMA) establishes a geologic timescale for North American fauna beginning during the Late Cretaceous and continuing through to the present. These periods are referred to as ages or intervals (or stages when ...
.


Palaeogeography and paleoclimate

The breakup of
Pangaea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...
was nearly complete in the Maastrichtian, with Australia beginning to break away from Antarctica and Madagascar breaking away from India. However, Arabia had not yet rifted away from Africa. North America was separated from Europe by rift basins, but sea floor spreading had not yet commenced between the two continents. The Pacific Plate was rapidly growing in size as the surrounding oceanic plates were consumed by
subduction Subduction is a geological process in which the oceanic lithosphere is recycled into the Earth's mantle at convergent boundaries. Where the oceanic lithosphere of a tectonic plate converges with the less dense lithosphere of a second plate, ...
, and the Pacific-Izanagi Ridge was rapidly approaching Asia. Eruption of the
Deccan Traps The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flo ...
large igneous province began during the Maastrichtian, at around 67 million years ago. This is thought to be a consequence of India drifting over the
Réunion hotspot The Réunion hotspot is a volcanic hotspot which currently lies under the island of Réunion in the Indian Ocean. The Chagos-Laccadive Ridge and the southern part of the Mascarene Plateau are volcanic traces of the Réunion hotspot. The hotsp ...
. During the Maastrichtian, the global climate began to shift from the warm and humid climate of the Mesozoic to the colder and more arid climate of the Cenozoic. Variation of climate with latitude also became greater. This was likely caused by a major reorganization of oceanic circulation that took place at the boundary between the early and late Maastrichtian. This reorganization was triggered by the breach of tectonic barriers in the South Atlantic, permitting deep ocean water to begin circulating from the nascent North Atlantic to the south. This initiated thermohaline circulation similar to that of the modern oceans. At the same time, the
Laramide orogeny The Laramide orogeny was a time period of mountain building in western North America, which started in the Late Cretaceous, 70 to 80 million years ago, and ended 35 to 55 million years ago. The exact duration and ages of beginning and end of the ...
drained the Western Interior Seaway of North America, further contributing to global cooling.


Paleontology

Dinosaurs remained the dominant large terrestrial animals throughout the Maasastrichtian, though mammals with internal organs similar to modern mammals were also present. Both
ammonites Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefis ...
and
pterosaurs Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 6 ...
were in serious decline during the Maastrichtian.


Dinosaurs


Birds

Several archaic clades of birds, such as
Enantiornithes The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
, Ichthyornithes, Hesperornithes, and
Apsaravis ''Apsaravis'' is a Mesozoic bird genus from the Late Cretaceous. The single known species, ''Apsaravis ukhaana'', lived about 78 million years ago, in the Campanian age of the Cretaceous period. Its fossilized remains were found in the Camel's ...
persisted to the latest Maasatrichtian but became extinct during the Cretaceous-Paleogene extinction event.


Pterosaurs

Traditionally, pterosaur faunas of the Maastrichtian were assumed to be dominated by
azhdarchids Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cret ...
, with other pterosaur groups having become extinct earlier on. However, more recent findings suggest a fairly composite pterosaur diversity: at least six ("Nyctosaurus" ''lamegoi'', a Mexican humerus, a Jordan humerus and several taxa from Morocco) nyctosaurs date to this period, as do a few
pteranodontids The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head ...
, and ''
Navajodactylus ''Navajodactylus'' (meaning "Navajo finger") is an extinct genus of pterodactyloid pterosaur from Late Cretaceous (late Campanian stage) deposits of the San Juan Basin in New Mexico, United States. Discovery The holotype specimen of ''Navajodac ...
'', tentatively assigned to Azhdarchidae, lacks any synapomorphies of the group. This seems to underscore a higher diversity of terminal Cretaceous pterosaurs than previously thought.


Flora

The radiation of
angiosperms Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants ...
(flowering plants) was well under way in the Maastrichtian. From 50% to 80% of all
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
of land plants were angiosperms, though
gymnosperms The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, Î³Ï…Î¼Î½Ï ...
and ferns still covered larger areas of the land surface.


Notes


References


External links


GeoWhen Database - MaastrichtianLate Cretaceous timescale
at the website of the subcommission for stratigraphic information of the ICS
Stratigraphic chart of the Late Cretaceous
at the website of Norges Network of offshore records of geology and stratigraphy
Maastrichtian Microfossils: 60+ images of Foraminifera
{{Authority control *06 Geological ages Cretaceous geochronology Maastricht