Luanchuanraptor
   HOME

TheInfoList



OR:

''Luanchuanraptor'' (meaning "
Luanchuan Luanchuan County () is a county under the jurisdiction of the prefecture-level city of Luoyang, in the west of Henan province, China. It has an area of and a population of 318,000. The county contains significant deposits of molybdenum and tungs ...
thief") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaurs from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
of China. The genus is based on a partial skeleton from the Qiupa Formation in
Luanchuan Luanchuan County () is a county under the jurisdiction of the prefecture-level city of Luoyang, in the west of Henan province, China. It has an area of and a population of 318,000. The county contains significant deposits of molybdenum and tungs ...
,
Henan Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...
. They were medium-sized dromaeosaurids, the first Asian dromaeosaurid taxa described from outside the Gobi Desert or northeastern China.


Discovery and naming

''Luanchuanraptor'' is known from a partial skeleton of an immature individual found at the Qiupa Formation of the
Henan Province Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...
,
Central China Central China () is a geographical and a loosely defined cultural region that includes the provinces of Henan, Hubei and Hunan. Jiangxi is sometimes also regarded to be part of this region. Central China is now officially part of South Centra ...
. The fossils were cataloged as 41HIII-0100 and described by Lü and colleagues in 2007. The remains represent the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
for the genus and species ''Luanchuanraptor henanensis'' and they are housed at the
Henan Geological Museum The Henan Museum (), located in Zhengzhou, Henan Province, China, is a history and archaeology museum. It has a collection of more than 130,000 pieces of cultural relics through the ages. In addition to its collection of human history the museum ...
. It consists of the left
frontal Front may refer to: Arts, entertainment, and media Films * ''The Front'' (1943 film), a 1943 Soviet drama film * ''The Front'', 1976 film Music * The Front (band), an American rock band signed to Columbia Records and active in the 1980s and e ...
, 4
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, t ...
(9 were identified but 5 were excluded), 4 cervical vertebrae, 6
dorsal vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebrae and they are intermediate in size between the cervical ...
, 17 caudal vertebrae, 4
ribs The rib cage, as an enclosure that comprises the ribs, vertebral column and sternum in the thorax of most vertebrates, protects vital organs such as the heart, lungs and great vessels. The sternum, together known as the thoracic cage, is a semi- ...
, 4
chevrons Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * '' Eulithis testata'', a moth * Chevron (geology), a fold in rock l ...
, a right humerus, left
scapulocoracoid The scapulocoracoid is the unit of the pectoral girdle that contains the coracoid and scapula. The coracoid itself is a beak-shaped bone that is commonly found in most vertebrates with a few exceptions. The scapula is commonly known as the ''shoulde ...
, the first
phalanx The phalanx ( grc, φάλαγξ; plural phalanxes or phalanges, , ) was a rectangular mass military formation, usually composed entirely of heavy infantry armed with spears, pikes, sarissas, or similar pole weapons. The term is particularly ...
from right manus, an isolated manual
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropod Sauropoda (), whose members are known as sauropods (; ...
, right ilium, left pubis, ischium, the sacrum and the shaft of the left
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
. These elements come from a moderately sized dromaeosaurid. Its specific name, ''Luanchuanraptor'', is a reference to the
Luanchuan County Luanchuan County () is a county under the jurisdiction of the prefecture-level city of Luoyang, in the west of Henan province, China. It has an area of and a population of 318,000. The county contains significant deposits of molybdenum and tungs ...
in which the remains were found, and the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''raptor'', meaning thief or seizer. The specific name, ''henanensis'', is a clear referral to the Henan Province.


Description

''Luanchuanraptor'' were moderate-sized dromaeosaurs, estimated at long with weights from .Genus List for Holtz 2012
/ref> However, the individual 41HIII-0100 preserves an unfused frontal, meaning that it was not a fully-grown animal at the time of death and therefore, they reached slightly larger sizes. They can be recognised from other dromaeosaur taxa in having a very stiff tail tip as indicated by the short neural spines of the caudal vertebrae, connected anterior chevrons, some posterior caudals with depressions near the neural spine, stocky proximal and posterior end of the chevrons, and a large opening on the
coracoid A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is prese ...
.


Classification

Lü and colleagues assigned ''Luanchuanraptor'' to the
Dromaeosauridae Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
based on the recurved, serrated and laterally flattened teeth, the continuous parapophyses on dorsal vertebrae, and the elongate caudal prezygapophyses. The genus has been considered to pertain to the
Averaptora Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the aviala ...
by Agnolín and Novas, but this suggestion is not widely followed. The recently performed phylogenetic analysis for the Dromaeosauridae by Hartman et al. 2019 recovers ''Luanchuanraptor'' as a
velociraptorine Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably ''Nuthetes'' from the United Kingdom, and possibly '' Deinonychus'' from North America. However, several indeterminate velociraptori ...
being the sister taxon of '' Adasaurus''. Below is the obtained result for the
Eudromaeosauria Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were relatively small to medium-sized, feathered hypercarnivores (with diets consisting almost entirely of other terrestrial vertebrates) ...
:


Paleoenvironment

The remains of ''Luanchuanraptor'' were found in the Late Cretaceous Qiupa Formation of China. It is considered to be Late
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval ...
in age about 72 million to 66 million years ago based on stratigraphic correlations. Most of the environments present consisted on shallow lakes with braided, delta rivers. The fauna present on the formation seems to be different from other Asiatic formations, with the well reported occurrences of mammals and lizards like '' Yubaatar'' or '' Funiusaurus''. Dinosaurian fauna was present as well, here ''Luanchuanraptor'' shared their habitat with other theropods such as '' Qiupalong'', ''
Qiupanykus ''Qiupanykus'' (meaning "Qiupa claw" after the Qiupa Formation) is a genus of alvarezsaurid coelurosaur theropod from the Late Cretaceous Qiupa Formation of southern China. Fossil eggs believed to be those of an oviraptorid found in association ...
'' or ''
Yulong Yulong may refer to: * Yulong Naxi Autonomous County (), of Lijiang, Yunnan * Yulong Snow Mountain (), or Jade Dragon Snow Mountain * Yulong River (), in Guangxi * Yulongsi Formation, in Yunnan * Yulong, Xingyang (), town in Xingyang City, Henan ...
''. Large tyrannosaurids have been reported from the formation under the species ''"
Tyrannosaurus ''Tyrannosaurus'' is a genus of large theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' live ...
" luanchuanensis''.Translated paper
/ref>


See also

*
Timeline of dromaeosaurid research This timeline of dromaeosaurid research is a chronological listing of events in the history of paleontology focused on the dromaeosaurids, a group of sickle-clawed, bird-like theropod dinosaurs including animals like ''Velociraptor''. Since the ...
*
Glossary of dinosaur anatomy This glossary explains technical terms commonly employed in the description of dinosaur body fossils. Besides dinosaur-specific terms, it covers terms with wider usage, when these are of central importance in the study of dinosaurs or when their ...


References

{{Taxonbar, from=Q2029421 Late Cretaceous dinosaurs of Asia Eudromaeosaurs Fossil taxa described in 2007