List of extinct animals of the Netherlands
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This list of extinct animals of the Netherlands includes the animal
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and subspecies once lived in the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
but have disappeared since human habitation. This list features the mammals,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s,
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
,
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estim ...
, butterflies,
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threa ...
, bees, pond damselflies,
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...
, grasshoppers and
Crickets Crickets are orthopteran insects which are related to bush crickets, and, more distantly, to grasshoppers. In older literature, such as Imms,Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 8 ...
that have disappeared from the Netherlands. There have been no known extinctions of reptiles or amphibians in the Netherlands. Most animals on this list of extinct animals in the Netherlands survive in other places in the world. However, some of them are now globally extinct, like the great auk (''Pinguinus impennis''), the European wild horse (''Equus ferus'') and the aurochs (''Bos primigenius primigenius''). One skeleton of the great auk was excavated in a Roman settlement near
Velsen Velsen () is a municipality in the Netherlands, in the province of North Holland. It is located on both sides of the North Sea Canal. On the north side of the North Sea Canal there is a major steel plant, Tata Steel IJmuiden, formerly known a ...
. Bones were also found near
Rotterdam Rotterdam ( , , , lit. ''The Dam on the River Rotte'') is the second largest city and municipality in the Netherlands. It is in the province of South Holland, part of the North Sea mouth of the Rhine–Meuse–Scheldt delta, via the ''"Ne ...
. In the Netherlands there are no bone finds of the aurochs after the Roman period (400 AD). '' Phengaris alcon arenaria'', an endemic Dutch subspecies of the Alcon blue butterfly became extinct at the end of the 1970s. Fossilized remains of the gray whale (''Eschrichtuis robustus''), have been found dated to 340 BC, demonstrating that this species once roamed the
North Sea The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea, epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the ...
, although it is no longer found there. A lower jaw of a northern lynx (''Lynx lynx lynx'') was found at the remains of a Roman settlement near Valkenburg in the Netherlands. During excavations of sites dated to the Roman period (around 400 AD) on the
Rhine ), Surselva, Graubünden, Switzerland , source1_coordinates= , source1_elevation = , source2 = Rein Posteriur/Hinterrhein , source2_location = Paradies Glacier, Graubünden, Switzerland , source2_coordinates= , so ...
delta there were findings of important breeding sites of the
Dalmatian pelican The Dalmatian pelican (''Pelecanus crispus'') is the largest member of the pelican family, and perhaps the world's largest freshwater bird, although rivaled in weight and length by the largest swans. They are elegant soaring birds, with wingspa ...
('' Pelecanus crispus''). According to the hunting rights of the bishops of
Utrecht Utrecht ( , , ) is the fourth-largest city and a municipality of the Netherlands, capital and most populous city of the province of Utrecht. It is located in the eastern corner of the Randstad conurbation, in the very centre of mainland Net ...
we know that
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is ...
s (''Ursus arctos arctos'') were still found in the Netherlands as late as the eleventh century. According to a hunting licence from Drenthe, elk (''Alces alces alces'') were also known to be in this country until 1025. The
North Atlantic right whale The North Atlantic right whale (''Eubalaena glacialis'') is a baleen whale, one of three right whale species belonging to the genus '' Eubalaena'', all of which were formerly classified as a single species. Because of their docile nature, their s ...
(''Eubalaena glacialis''), which once appeared from the Bay of Biscay to
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the ...
, have disappeared from the waters around the Netherlands. It is suspected that the last whales were killed at the end of the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
. However, there was an alleged sighting off
Texel Texel (; Texels dialect: ) is a municipality and an island with a population of 13,643 in North Holland, Netherlands. It is the largest and most populated island of the West Frisian Islands in the Wadden Sea. The island is situated north of Den ...
in 2005.


Mammals

* ''Alces alces alces'' – European elk (1025) * ''Barbastellus barbastellus'' – barbastelle bat * ''Bos primigenius primigenius'' – Eurasian aurochs (400 AD) * ''Equus ferus ferus'' –
tarpan The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The las ...
* ''Lynx lynx lynx'' – Northern lynx (Roman period) * ''Mustela lutreola'' –
European mink The European mink (''Mustela lutreola''), also known as the Russian mink and Eurasian mink, is a semiaquatic species of mustelid native to Europe. It is similar in colour to the American mink, but is slightly smaller and has a less specialized ...
(1887) * ''Rhinolophus ferrumequinum'' –
greater horseshoe bat The greater horseshoe bat (''Rhinolophus ferrumequinum'') is an insectivorous bat of the genus '' Rhinolophus''. Its distribution covers Europe, Northern Africa, Central Asia and Eastern Asia. It is the largest of the horseshoe bats in Europe an ...
(1974) * ''Rhinolophus hipposideros'' – lesser horseshoe bat (1983) * ''Ursus arctos arctos'' –
Eurasian brown bear The Eurasian brown bear (''Ursus arctos arctos'') is one of the most common subspecies of the brown bear, and is found in much of Eurasia. It is also called the European brown bear, common brown bear, common bear, and colloquially by many othe ...
(eleventh century)


Birds

* ''Pinguinus impennis'' – great auk – globally extinct * ''Tringa glareola'' –
wood sandpiper The wood sandpiper (''Tringa glareola'') is a small wader. This Eurasian species is the smallest of the shanks, which are mid-sized long-legged waders of the family Scolopacidae. The genus name ''Tringa'' is the New Latin name given to the gree ...
(1939) – does not nest in the Netherlands any more, but they can be found during the migration season.


Fish

* ''Alosa alosa'' –
allis shad The allis shad (''Alosa alosa'') is a widespread Northeast Atlantic species of fish in the herring family, Clupeidae. It is an anadromous fish which migrates into fresh water to spawn. It is found in the eastern Atlantic Ocean, the western Bal ...
(1993) * ''Alosa fallax'' –
twaite shad The twait shad or twaite shad (''Alosa fallax'') is a species of fish in the family Clupeidae. It is found in the eastern Atlantic Ocean and the Mediterranean Sea and is an anadromous fish which lives in the sea but migrates into fresh water to s ...
(1970) * ''Coregonus oxyrinchus'' – houting (1940) * ''Hippocampus ramulosus'' – common seahorse * ''Thymallus thymallus'' – grayling * '' Salmo trutta fario'' – brown trout * ''Syngnathus typhle'' – deepnosed pipefish


Molluscs

* '' Pisidium tenuilineatum'' * '' Rissoa membranacea'' * '' Spermodea lamellata'' * ''Unio crassus'' – thick shelled river mussel (1968)


Insects


Butterflies

* ''
Aporia crataegi ''Aporia crataegi'', the black-veined white, is a large butterfly of the family Pieridae. A. crataegi is widespread and common. Its range extends from northwest Africa in the west to Transcaucasia and across the Palearctic to Siberia and Japan ...
'' * ''Argynnis paphia'' * ''Boloria euphrosyne'' * ''Brenthis ino'' * ''Coenonympha hero'' * ''Cupido minimus minimus'' * ''Euphydryas aurinia aurinia'' * ''Lycaena hippothoe hippothoe'' * ''Melitaea diamina'' * ''Nymphalis antiopa'' * '' Phengaris alcon arenaria'' (1979) * ''Phengaris arion'' * ''Phengaris nausithous'' * ''Phengaris teleius'' * ''Plebeius idas idas'' * '' Polyommatus semiargus semiargus'' * ''Thymelicus acteon acteon'' * ''Spialia sertorius sertorius''


Damselflies

* ''
Nehalennia speciosa ''Nehalennia speciosa'' (pygmy damselfly, sedgeling or sedgling) is a species of damselfly in the family Coenagrionidae. It is found in Austria, Belarus, the Czech Republic, Denmark, Estonia, Finland, Germany, Italy, Japan, North Korea, Latvia, L ...
''


Bees

* '' Ammobates punctatus'' * '' Andrena curvungula'' * '' Andrena marginata'' * '' Andrena nitidiuscula'' * '' Andrena pandellei'' * '' Andrena schencki'' * '' Andrena thoracica'' * '' Anthidium byssinum'' * '' Anthophora aestivalis'' * '' Anthophora bimaculata'' * '' Anthophora borealis'' * '' Anthophora plagiata'' * '' Biastes truncatus'' * '' Bombus confusus'' * '' Bombus cullumanus'' * '' Bombus pomorum'' * '' Bombus subterraneus'' * '' Coelioxys alata'' * '' Dufourea minuta'' * '' Halictus eurygnathus'' * ''
Halictus quadricinctus The genus ''Halictus'' is a large assemblage of bee species in the family Halictidae. The genus is divided into 15 subgenera, some of dubious monophyly, containing over 200 species, primarily in the Northern Hemisphere (a few species occur in So ...
'' * '' Lasioglossum laeve'' * '' Lasioglossum laevigatum'' * '' Nomada argentata'' * '' Nomada furva'' * '' Nomada mutabilis'' * '' Nomada obtusifrons'' * '' Nomada piccioliana'' * '' Nomada rhenana'' * '' Nomada roberjeotiana'' * '' Osmia anthocopoides'' * '' Osmia papaveris'' * '' Osmia xanthomela'' * '' Rophites quinquespinosus'' * '' Thyreus orbatus''


Pond damselflies

* '' Holocentropus insignis'' * '' Hydroptila cornuta'' * '' Hydroptila dampfi'' * '' Ithytrichia lamellaris'' * '' Micrasemodes minimus'' * '' Oligoplectrum maculatum'' * '' Sericostoma flavicorne'' * '' Setodes viridis'' * '' Silo piceus''


Grasshoppers and crickets

* ''
Locusta migratoria The migratory locust (''Locusta migratoria'') is the most widespread locust species, and the only species in the genus ''Locusta''. It occurs throughout Africa, Asia, Australia and New Zealand. It used to be common in Europe but has now become ...
'' * '' Psophus stridulus''


Stoneflies

* '' Euleuctra geniculata'' * '' Isogenus nubecula'' * '' Isoperla grammatica'' * '' Isoptena serricornis'' * '' Leuctra fusca'' * '' Marthamea selysii'' * '' Protonemura nitida'' * '' Taeniopteryx nebulosa'' * ''
Xanthoperla apicalis ''Xanthoperla apicalis'' is a species of stoneflies in the family Chloroperlidae. Subspecies *''Xanthoperla apicalis hamulata'' (Morton, 1930) Distribution and habitat This species is present in most of Europe (Albania, Belgium, Germany, Great ...
''


Mayflies

* '' Ametropus fragilis'' * '' Choroterpes picteti'' * '' Ecdyonurus affinis'' * '' Ecdyonurus dispar'' * '' Habroleptoides modesta'' * '' Habrophlebia lauta'' * '' Heptagenia coerulans'' * '' Isonychia ignota'' * '' Oligoneuriella rhenana'' * '' Palingenia longicauda'' * '' Potamanthus luteus'' * '' Siphlonurus aestivalis'' * '' Siphlonurus alternatus'' * '' Siphlonurus lacustris''


Reintroductions and rediscoveries


Mammals

* ''Castor fiber albicus'' –
European beaver The Eurasian beaver (''Castor fiber'') or European beaver is a beaver species that was once widespread in Eurasia, but was hunted to near-extinction for both its fur and castoreum. At the turn of the 20th century, only about 1,200 beavers survi ...
The last known European beaver in the Netherlands was killed in 1826. In 1988 European beavers were reintroduced in the
Biesbosch De Biesbosch National Park is one of the largest national parks of the Netherlands and one of the last extensive areas of freshwater tidal wetlands in Northwestern Europe. The Biesbosch ('forest of sedges' or 'rushwoods') consists of a large ...
, and in 1994 beavers were released in the Gelderse Poort (a wilderness area between
Arnhem Arnhem ( or ; german: Arnheim; South Guelderish: ''Èrnem'') is a city and municipality situated in the eastern part of the Netherlands about 55 km south east of Utrecht. It is the capital of the province of Gelderland, located on both ban ...
and Nijmegen). The new beavers are doing very well; their numbers are increasing and they are spreading to other parts of the Netherlands. * ''Canis lupus lupus'' –
Eurasian wolf The Eurasian wolf (''Canis lupus lupus''), also known as the common wolf,Mech, L. David (1981), ''The Wolf: The Ecology and Behaviour of an Endangered Species'', University of Minnesota Press, p. 354, is a subspecies of grey wolf native to Euro ...
ICUN lists the grey wolf as regionally extinct in the Netherlands. In March 2015, the first wolf in 100 years was sighted in the country. This was the first recorded, and second reported sighting following recent successful wolf reintroduction programs in neighboring Germany, with transient migrant wolves apparently occasionally crossing the border. Three wolve cubs were spotted on the Veluwe in June, 2019 * ''Cricetus cricetus canescens'' – European hamsters Under orders from the Dutch Government, in 1999 the Das&Boom Foundation caught all the remaining European hamsters in the Netherlands. These animals were placed in a breeding programme in
Diergaarde Blijdorp ( ''Blijdorp Zoo''), officially Rotterdam Zoo, is a zoo located in the northwestern part of Rotterdam. It is one of the oldest zoos in the Netherlands, and has been operated by the ("Royal Rotterdam Zoo Foundation"). Divided into several zoogeog ...
(Rotterdam Zoo). They were extinct in the wild, but offspring from the breeding programme have been reintroduced in a hamster reserve in
Sibbe Sibbe ( Limburgish: ''Sub'' or ''Süp'') is the official Dutch name of a village in the municipality of Valkenburg aan de Geul in the province of Limburg in the southern part of the Netherlands. History It has been proposed that the name of the vi ...
in the southern province of
Limburg Limburg or Limbourg may refer to: Regions * Limburg (Belgium), a province since 1839 in the Flanders region of Belgium * Limburg (Netherlands), a province since 1839 in the south of the Netherlands * Diocese of Limburg, Roman Catholic Diocese in ...
. In 2003 more hamsters were released in a second hamster reserve in Amby, near
Maastricht Maastricht ( , , ; li, Mestreech ; french: Maestricht ; es, Mastrique ) is a city and a municipality in the southeastern Netherlands. It is the capital and largest city of the province of Limburg. Maastricht is located on both sides of the ...
. These reintroductions were followed by four more reintroductions in Heer, Sittard, Puth and Koningsbosch. The wild hamster population has now grown to c. 600 burrows (December 2006). * ''Felis silvestris silvestris'' – European wild cat The wild cat probably became extinct in the Netherlands in Roman times. The first confirmed specimen in the Netherlands since that time was a dead cat found near Groenlanden in
Gelderland Gelderland (), also known as Guelders () in English, is a province of the Netherlands, occupying the centre-east of the country. With a total area of of which is water, it is the largest province of the Netherlands by land area, and second by ...
, while another dead animal was found in 2002 near
Vaals Vaals (; Ripuarian: ) is a town in the extreme southeastern part of the Dutch province of Limburg, which is in the southeastern part of the Netherlands. The municipality covers an area of in the foothills of the Ardennes–Eifelrange ...
in South
Limburg Limburg or Limbourg may refer to: Regions * Limburg (Belgium), a province since 1839 in the Flanders region of Belgium * Limburg (Netherlands), a province since 1839 in the south of the Netherlands * Diocese of Limburg, Roman Catholic Diocese in ...
. The first living cat was caught near Heeze,
North Brabant North Brabant ( nl, Noord-Brabant ; Brabantian: ; ), also unofficially called Brabant, is a province in the south of the Netherlands. It borders the provinces of South Holland and Gelderland to the north, Limburg to the east, Zeeland to the w ...
in 2004. In 2006 a wild cat was caught on camera near Vaals. These few sightings are not yet positive proof of the wildcat settling in the Netherlands, but the known range of the wildcat has been approaching the Dutch borders since the 1990s. * ''Lutra lutra'' –
European otter The Eurasian otter (''Lutra lutra''), also known as the European otter, Eurasian river otter, common otter, and Old World otter, is a semiaquatic mammal native to Eurasia. The most widely distributed member of the otter subfamily (Lutrinae) of th ...
The last lonely otter in the Netherlands was killed by a car on 17 September 1988 in the neighbourhood of Joure (Province of
Friesland Friesland (, ; official fry, Fryslân ), historically and traditionally known as Frisia, is a province of the Netherlands located in the country's northern part. It is situated west of Groningen, northwest of Drenthe and Overijssel, north of ...
). The first otters were reintroduced in National Park De Weerribben (Province of Overijssel) on 8 July 2002. By 2012 they had been released in other parts of the Netherlands as well. * ''Phocoena phocoena'' –
harbour porpoise The harbour porpoise (''Phocoena phocoena'') is one of eight extant species of porpoise. It is one of the smallest species of cetacean. As its name implies, it stays close to coastal areas or river estuaries, and as such, is the most familiar ...
The twentieth century saw the taming of the
Zuiderzee The Zuiderzee or Zuider Zee (; old spelling ''Zuyderzee'' or ''Zuyder Zee'') was a shallow bay of the North Sea in the northwest of the Netherlands, extending about 100 km (60 miles) inland and at most 50 km (30 miles) wide, with an o ...
as a large enclosure dam (the
Afsluitdijk The ''Afsluitdijk'' (; fry, Ofslútdyk; nds-nl, Ofsluutdiek; en, "Closure Dyke") is a major dam and causeway in the Netherlands. It was constructed between 1927 and 1932 and runs from Den Oever in North Holland province to the village of ...
) was constructed. Completed in 1932, the Zuiderzee became the IJsselmeer and large areas of water could be reclaimed for farming and housing. After this the harbour porpoise, together with the bottlenose dolphin, disappeared from the waters around the Netherlands. They came back in the 1980s.


Birds

* ''Ciconia ciconia'' –
white stork The white stork (''Ciconia ciconia'') is a large bird in the stork family, Ciconiidae. Its plumage is mainly white, with black on the bird's wings. Adults have long red legs and long pointed red beaks, and measure on average from beak tip to e ...
Once these birds were very common in the Netherlands, but their number decreased fast in the twentieth century. 1891 was the first year that no white stork bred in the Netherlands. A conservation and reintroduction program that started in 1967 resulted in 396 pairs in 2000. * ''Egretta garzetta garzetta'' – little egret This bird became extinct in the Netherlands in the nineteenth century, due to overhunting because of their feathers which were used in the hat industry. In 1979 this bird first bred again in the Oostvaardersplassen nature reserve. The second time this bird bred again in the Netherlands was in 1994. After that year it bred yearly in the Netherlands. Their numbers are still increasing. * ''Grus grus'' –
common crane The common crane (''Grus grus''), also known as the Eurasian crane, is a bird of the family Gruidae, the cranes. A medium-sized species, it is the only crane commonly found in Europe besides the demoiselle crane (''Grus virgo'') and the Siberian ...
In 2001, one common crane pair bred successfully after 250 years in the Fochteloërveen, a nature reserve on the border of the provinces of
Friesland Friesland (, ; official fry, Fryslân ), historically and traditionally known as Frisia, is a province of the Netherlands located in the country's northern part. It is situated west of Groningen, northwest of Drenthe and Overijssel, north of ...
and
Drente Drenthe () is a province of the Netherlands located in the northeastern part of the country. It is bordered by Overijssel to the south, Friesland to the west, Groningen to the north, and the German state of Lower Saxony to the east. As of Nov ...
. Before 2001 the common crane could only be found during the migration period. * ''Porzana pusilla intermedia'' – Baillon's crake This bird was considered extinct in the Netherlands after it was last sighted breeding in 1972. In early 2005 five territorial and two breeding pairs were located again in the province of
Utrecht Utrecht ( , , ) is the fourth-largest city and a municipality of the Netherlands, capital and most populous city of the province of Utrecht. It is located in the eastern corner of the Randstad conurbation, in the very centre of mainland Net ...
.


Fish

* ''Salmo salar'' – Atlantic salmon The Atlantic salmon was very common in the Netherlands in the seventeenth century, but disappeared when the rivers were tamed and closed by the Dutch to protect their land. The salmon could not reach their breeding ground in the rivers
Rhine ), Surselva, Graubünden, Switzerland , source1_coordinates= , source1_elevation = , source2 = Rein Posteriur/Hinterrhein , source2_location = Paradies Glacier, Graubünden, Switzerland , source2_coordinates= , so ...
and
Meuse The Meuse ( , , , ; wa, Moûze ) or Maas ( , ; li, Maos or ) is a major European river, rising in France and flowing through Belgium and the Netherlands before draining into the North Sea from the Rhine–Meuse–Scheldt delta. It has a t ...
. A reintroduction program resulted in salmon in the IJsselmeer and the river Rhine.


Dragonflies

* ''Coenagrion armatum'' – Norfolk damselfly In 1956 this damselfly was thought to be extinct in the Netherlands, but was rediscovered in the National Park De Weerribben on 9 May 1999. * ''Coenagrion mercuriale'' –
southern damselfly ''Coenagrion mercuriale'', the southern damselfly, is a species of damselfly in the family Coenagrionidae. It is found in Algeria, Austria, Belgium, France, Germany, Italy, Liechtenstein, Luxembourg, Morocco, the Netherlands, Portugal, Romania, S ...
Was last seen in 1953, but was rediscovered in
Limburg Limburg or Limbourg may refer to: Regions * Limburg (Belgium), a province since 1839 in the Flanders region of Belgium * Limburg (Netherlands), a province since 1839 in the south of the Netherlands * Diocese of Limburg, Roman Catholic Diocese in ...
in 2011.


Grasshoppers

* ''Tetrix bipunctata'' Tetrix bipunctata is a species of pygmy locust that was not recorded between 1975 and 2011. Therefore, it was listed as being 'extinct' on the Dutch Red List (Rode Lijst) in 1999.


See also

*
List of extinct animals of Europe This list of European animals extinct in the Holocene features animals known to have become extinct in the last 12,000 years on the European continent and its surrounding islands. Dependent territories of European countries in other continents, l ...
* List of molluscs recorded in the Netherlands


References


External links


Dutch Ministry of Agriculture, Nature and Food Quality
{{Extinct Animals by Regions
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
Animals, extinct Animals, extinct