Liriodendron
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''Liriodendron'' () is a genus of two species of characteristically large trees,
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
over most of their populations, in the magnolia
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
(
Magnoliaceae The Magnoliaceae () are a flowering plant family, the magnolia family, in the order Magnoliales. It consists of two genera: ''Magnolia'' and '' Liriodendron'' (tulip trees). Unlike most angiosperms, whose flower parts are in whorls (rings), t ...
). These trees are widely known by the common name tulip tree or tuliptree for their large flowers superficially resembling
tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
s. It is sometimes referred to as tulip poplar or yellow poplar, and the wood simply as "poplar", although not closely related to the true
poplars ''Populus'' is a genus of 25–30 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar (), aspen, and cottonwood. The w ...
. Other common names include canoewood, saddle-leaf tree, and white wood. The two
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species are ''
Liriodendron tulipifera ''Liriodendron tulipifera''—known as the tulip tree, American tulip tree, tulipwood, tuliptree, tulip poplar, whitewood, fiddletree, and yellow-poplar—is the North American representative of the two-species genus '' Liriodendron'' (the othe ...
'',
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
and '' Liriodendron chinense'', native to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
and
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
. Both species often grow to great size, the North American species may reach as much as in height. The North American species is commonly used horticulturally, the Chinese species is increasing in cultivation, and hybrids have been produced between these two allopatrically distributed species. Various extinct species of ''Liriodendron'' have been described from the fossil record.


Description

''Liriodendron'' trees are easily recognized by their
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
, which are distinctive, having four lobes in most cases and a cross-cut notched or straight apex. Leaf size varies from 8–22 cm long and 6–25 cm wide. They are deciduous in the vast majority of cases for both species; however, each species has a semi-evergreen variety at the southern limit of its range in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
and
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
respectively. The tulip tree is often a large tree, 18–60 m high and 60–120 cm in diameter. The tree is known to reach the height of 191.8 feet (58.49 meters), in groves where they compete for sunlight, somewhat less if growing in an open field. Its trunk is usually columnar, with a long, branch-free bole forming a compact, rather than open, conical crown of slender branches. It has deep roots that spread widely.Michigan Trees Leaves are slightly larger in ''L. chinense,'' compared to ''L. tulipifera'', but with considerable overlap between the species; the petiole is 4–18 cm long. Leaves on young trees tend to be more deeply lobed and larger in size than those on mature trees. In autumn, the leaves turn yellow, or brown and yellow. Both species grow rapidly in rich, moist soils of temperate climates. They hybridize easily.
Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are 3–10 cm in diameter and have nine
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s — three green outer
sepals A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined ...
and six inner
petals Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
which are yellow-green with an orange flare at the base. They start forming after around 15 years and are superficially similar to a
tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
in shape, hence the tree's name. Flowers of ''L. tulipifera'' have a faint
cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated creeping vine plant in the Cucurbitaceae family that bears usually cylindrical fruits, which are used as culinary vegetables.stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s and
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils ...
s are arranged spirally around a central spike or gynaecium; the stamens fall off, and the pistils become the
samara Samara ( rus, Сама́ра, p=sɐˈmarə), known from 1935 to 1991 as Kuybyshev (; ), is the largest city and administrative centre of Samara Oblast. The city is located at the confluence of the Volga and the Samara rivers, with a population ...
s. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a cone-like aggregate of samaras 4–9 cm long, each of which has a roughly tetrahedral seed with one edge attached to the central conical spike and the other edge attached to the wing.


Distribution

''Liriodendron'' trees are also easily recognized by their general shape, with the higher branches sweeping together in one direction, and they are also recognizable by their height, as the taller ones usually protrude above the canopy of oaks, maples, and other trees—more markedly with the American species. Appalachian cove forests often contain several tulip trees of height and girth not seen in other species of eastern hardwoods. In the Appalachian cove forests, trees 150 to 165 ft in height are common, and trees from 166 to nearly 180 ft are also found. More ''Liriodendron'' over 170 ft in height have been measured by the Eastern Native Tree Society than for any other eastern species. The current tallest tulip tree on record has reached 191.9 ft, the tallest native
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants ...
tree known in North America. The tulip tree is rivaled in eastern forests only by white pine, loblolly pine, and
eastern hemlock ''Tsuga canadensis'', also known as eastern hemlock, eastern hemlock-spruce, or Canadian hemlock, and in the French-speaking regions of Canada as ''pruche du Canada'', is a coniferous tree native to eastern North America. It is the state tree of ...
. Reports of tulip trees over 200 ft have been made, but none of the measurements has been confirmed by the Eastern Native Tree Society. Most reflect measurement errors attributable to not accurately locating the highest crown point relative to the base of the tree—a common error made by the users employing only clinometers/ hypsometers when measuring height. Maximum circumferences for the species are between 24 and 30 ft at breast height, although a few historical specimens may have been slightly larger. The
Great Smoky Mountains National Park Great Smoky Mountains National Park is an American national park in the southeastern United States, with parts in North Carolina and Tennessee. The park straddles the ridgeline of the Great Smoky Mountains, part of the Blue Ridge Mountains, w ...
has the greatest population of tulip trees 20 ft and over in circumference. The largest-volume tulip tree known anywhere is the Sag Branch Giant, which has a trunk and limb volume approaching .


Fossils

Liriodendrons have been reported as fossils from the late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
and early
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
of North America and central Asia. They are known widely as Tertiary-age
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s in Europe and well outside their present range in Asia and North America, showing a once- circumpolar northern distribution. Like many "Arcto-Tertiary" genera, ''Liriodendron'' apparently became extinct in Europe due to large-scale
glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate bet ...
and aridity of climate during glacial phases. (The name should not be confused with '' Lepidodendron,'' an important group of long-extinct
pteridophyte A pteridophyte is a vascular plant (with xylem and phloem) that disperses spores. Because pteridophytes produce neither flowers nor seeds, they are sometimes referred to as " cryptogams", meaning that their means of reproduction is hidden. Fer ...
s in the phylum Lycopodiophyta common as
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ...
coal-age fossils).


Cultivation and use

''Liriodendron'' trees prefer a
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
climate, sun or part shade, and deep,
fertile Fertility is the capability to produce offspring through reproduction following the onset of sexual maturity. The fertility rate is the average number of children born by a female during her lifetime and is quantified demographically. Fertili ...
, well-drained and slightly acidic
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
. Propagation is by seed or
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
. Plants grown from seed may take more than eight years to flower. Grafted plants flower depending on the age of the scion plant. The wood of the North American species (called poplar or tulipwood) is fine grained and stable. It is easy to work and commonly used for cabinet and furniture framing, i.e. internal structural members and subsurfaces for veneering. Additionally, much inexpensive furniture, described for sales purposes simply as "hardwood", is in fact primarily stained poplar. In the literature of American furniture manufacturers from the first half of the 20th century, it is often referred to as "gum wood". The wood is only moderately rot-resistant and is not commonly used in shipbuilding, but has found some recent use in light-craft construction. The wood is readily available, and when air dried, has a density around . The name canoewood probably refers to the tree's use for construction of
dugout canoe A dugout canoe or simply dugout is a boat made from a hollowed tree. Other names for this type of boat are logboat and monoxylon. ''Monoxylon'' (''μονόξυλον'') (pl: ''monoxyla'') is Greek – ''mono-'' (single) + '' ξύλον xylon'' (t ...
s by eastern Native Americans, for which its fine grain and large trunk size is eminently suited. Tulip tree leaves are eaten by the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sy ...
s of some
Lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described speci ...
, for example the
eastern tiger swallowtail ''Papilio glaucus'', the eastern tiger swallowtail, is a species of butterfly native to eastern North America. It is one of the most familiar butterflies in the eastern United States, ranging north to southern Ontario, Canada, and is common in m ...
(''Papilio glaucus'').


Species and cultivars

*'' Liriodendron chinense'' *''
Liriodendron tulipifera ''Liriodendron tulipifera''—known as the tulip tree, American tulip tree, tulipwood, tuliptree, tulip poplar, whitewood, fiddletree, and yellow-poplar—is the North American representative of the two-species genus '' Liriodendron'' (the othe ...
'' **'Ardis' is a small-leaf, compact cultivar **'Aureomarginatum' is variegated with yellow-margined leaves **'Fastigiatum' grows with an erect or columnar habit ( fastigiate) **'Florida' strain — a fast-growing early bloomer, leaves have round lobes **'Glen Gold' bears yellow-gold colored leaves **'Mediopictum' is a variegated cultivar with gold-centered leaves *'Chapel Hill' and 'Doc Deforce's Delight' are hybrids of the above two species


References


External links

* Hunt, D. (ed). 1998. ''Magnolias and their allies''. International Dendrology Society & Magnolia Society. () * Parks, C.R., Wendel, J.F., Sewell, M.M., & Qiu, Y.-L. (1994). The significance of allozyme variation and introgression in the Liriodendron tulipifera complex (Magnoliaceae). ''Am. J. Bot.'' 81 (7): 878-88
abstract and first page
* Parks, C.R., Miller, N.G., Wendel, J.F. and McDougal, K.M. (1983). Genetic diversity within the genus Liriodendron (Magnoliaceae). ''Annals of the Missouri Botanical Garden'' 70 (4): 658-66
abstract and first page
* Collingwood, G.H., Brush, W.D. (1984) ''Knowing your trees''. American Forestry Association. (L.O.C. card no. 78-52994):286-287







* ttp://www.fna.org/china/mss/volume07/Magnoliaceae-CAS_coauthoring.htm ''Flora of China'' draft account of Magnoliaceae(site currently down; se
google cache


* ''Botanicas Trees & Shrubs'', Random House, Sydney, 2005


USDA Plants Tulip Popular


* ttp://www.psu.edu/dept/nkbiology/naturetrail/speciespages/poplar.htm Penn State Yellow Popular
USDA Forest Service Yellow Popular
{{Taxonbar, from=Q157236 Magnoliaceae Magnoliales genera