Lepidus (dinosaur)
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''Lepidus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of extinct
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
from the
Upper Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. ...
of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
. It lived in the
Otis Chalk Otis may refer to: Arts and entertainment Characters * Otis (Superman), in the films ''Superman'' and ''Superman II'' and related DC Comics media ** Otis Graves, in the TV series ''Supergirl'' * Otis (''The Walking Dead''), in the Image Comics ...
localities of the
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Colora ...
in
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
, around 223 million years ago.


Discovery

It was first described in 2015 by Nesbitt & Ezcurra, who decided it warranted a new taxon, which they named ''Lepidus praecisio''. The generic name is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "fascinating", and the specific name is Latin for "fragment", or "scrap". The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
material includes a
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
,
astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...
, and
fibula The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity i ...
, and other referred material includes a
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
and
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. T ...
. The holotypic material is well preserved and shows signs of muscle scars. The
astragalus ''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to tempe ...
and
calcaneum In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel) or heel bone is a bone of the tarsus of the foot which constitutes the heel. In some other animals, it is the point of the hock. ...
are clearly fused into one bone, with no visible suture. The tibia resembles that of neotheropods, in overall morphology. The shape of the tibia resembles the same in ''
Camposaurus ''Camposaurus'' ( ) is a coelophysid dinosaur genus from the Norian stage of the Late Triassic period of North America. The pertinent fossil remains date back to the early to middle Norian stage, and is widely regarded as the oldest known neot ...
'', ''
Coelophysis ''Coelophysis'' ( traditionally; or , as heard more commonly in recent decades) is an extinct genus of coelophysid theropod dinosaur that lived approximately 228 to 201.3 million years ago during the latter part of the Triassic Period fro ...
'', '' Tawa'', ''
Eodromaeus ''Eodromaeus'' (meaning "dawn runner") is an extinct genus of probable basal theropod dinosaurs from the Late Triassic of Argentina. Like many other of the earliest-known dinosaurs, it hails from the Carnian-age (~230 Ma) Ischigualasto Form ...
'', and ''
Herrerasaurus ''Herrerasaurus'' is a genus of saurischian dinosaur from the Late Triassic period. This genus was one of the earliest dinosaurs from the fossil record. Its name means "Herrera's lizard", after the rancher who discovered the first specimen in ...
''. ''Lepidus'' was one of eighteen dinosaur taxa from 2015 to be described in open access or free-to-read journals.


Classification

A phylogenetic analysis was conducted, including with and without referred material. The results are shown below:


See also

* Timeline of coelophysoid research


References

Late Triassic dinosaurs of North America Coelophysoids Fossil taxa described in 2015 Taxa named by Sterling Nesbitt Paleontology in Texas {{theropod-stub