Lemon (fruit)
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The lemon (''Citrus limon'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of small
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
trees in the
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
family
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Der ...
, native to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, primarily
Northeast India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...
(
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
), Northern Myanmar or China. The tree's ellipsoidal yellow
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is used for culinary and non-culinary purposes throughout the world, primarily for its
juice Juice is a drink made from the extraction or pressing of the natural liquid contained in fruit and vegetables. It can also refer to liquids that are flavored with concentrate or other biological food sources, such as meat or seafood, such as ...
, which has both culinary and cleaning uses. The pulp and rind are also used in cooking and baking. The juice of the lemon is about 5% to 6%
citric acid Citric acid is an organic compound with the chemical formula HOC(CO2H)(CH2CO2H)2. It is a colorless weak organic acid. It occurs naturally in citrus fruits. In biochemistry, it is an intermediate in the citric acid cycle, which occurs in ...
, with a pH of around 2.2, giving it a sour taste. The distinctive sour taste of lemon juice makes it a key ingredient in drinks and foods such as
lemonade Lemonade is a sweetened lemon-flavored beverage. There are varieties of lemonade found throughout the world. In North America and South Asia, cloudy still lemonade is the most common variety. There it is traditionally a homemade drink using ...
and
lemon meringue pie Lemon meringue pie is a dessert pie consisting of a shortened pastry base filled with lemon curd and topped with meringue. History Fruit desserts covered with baked meringue were found beginning in the 18th century in France. Menon's ''pom ...
.


History

The origin of the lemon is unknown, though lemons are thought to have first grown in
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
(a region in
northeast India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...
), northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
or China. A genomic study of the lemon indicated it was a hybrid between
bitter orange Bitter orange, Seville orange, bigarade orange, or marmalade orange is the citrus tree ''Citrus'' × ''aurantium'' and its fruit. It is native to Southeast Asia and has been spread by humans to many parts of the world. It is probably a cross be ...
(sour orange) and
citron The citron (''Citrus medica''), historically cedrate, is a large fragrant citrus fruit with a thick rind. It is said to resemble a 'huge, rough lemon'. It is one of the original citrus fruits from which all other citrus types developed throu ...
. Lemons are supposed to have entered
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
near southern
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
no later than the second century AD, during the time of
Ancient Rome In modern historiography, ancient Rome refers to Roman civilisation from the founding of the city of Rome in the 8th century BC to the collapse of the Western Roman Empire in the 5th century AD. It encompasses the Roman Kingdom (753–509 BC ...
. They were later introduced to
Persia Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
and then to
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to the north, Iran to the east, the Persian Gulf and K ...
and
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Medit ...
around 700 AD. The lemon was first recorded in literature in a 10th-century
Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walter ...
treatise on farming, and was also used as an ornamental plant in early Islamic gardens. It was distributed widely throughout the Arab world and the
Mediterranean region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and wa ...
between 1000 and 1150. An article on Lemon and lime tree cultivation in
Andalusia Andalusia (, ; es, Andalucía ) is the southernmost autonomous community in Peninsular Spain. It is the most populous and the second-largest autonomous community in the country. It is officially recognised as a "historical nationality". The t ...
of Spain is brought down in
Ibn al-'Awwam Ibn al-'Awwam ( ar, ابن العوام), also called Abu Zakariya Ibn al-Awwam ( ar, أبو زكريا بن العوام), was a Muslim Arab agriculturist who flourished at Seville (modern-day southern Spain) in the later 12th century. He wrote a ...
's 12th-century agricultural work, ''Book on Agriculture''. The first substantial cultivation of lemons in Europe began in
Genoa Genoa ( ; it, Genova ; lij, Zêna ). is the capital of the Italian region of Liguria and the sixth-largest city in Italy. In 2015, 594,733 people lived within the city's administrative limits. As of the 2011 Italian census, the Province of ...
in the middle of the 15th century. The lemon was later introduced to the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
in 1493 when
Christopher Columbus Christopher Columbus * lij, Cristoffa C(or)ombo * es, link=no, Cristóbal Colón * pt, Cristóvão Colombo * ca, Cristòfor (or ) * la, Christophorus Columbus. (; born between 25 August and 31 October 1451, died 20 May 1506) was a ...
brought lemon seeds to Hispaniola on his voyages. Spanish conquest throughout the New World helped spread lemon seeds. It was mainly used as an ornamental plant and for medicine. In the 19th century, lemons were increasingly planted in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
and
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. In 1747,
James Lind James Lind (4 October 1716 – 13 July 1794) was a Scottish doctor. He was a pioneer of naval hygiene in the Royal Navy. By conducting one of the first ever clinical trials, he developed the theory that citrus fruits cured scurvy. Lind ...
's experiments on seamen suffering from
scurvy Scurvy is a disease resulting from a lack of vitamin C (ascorbic acid). Early symptoms of deficiency include weakness, feeling tired and sore arms and legs. Without treatment, decreased red blood cells, gum disease, changes to hair, and bleeding ...
involved adding lemon juice to their diets, though
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) ...
was not yet known as an important dietary ingredient. The origin of the word ''lemon'' may be Middle Eastern. The word draws from the Old French ''limon'', then Italian ''limone'', from the Arabic ''laymūn'' or ''līmūn'', and from the
Persian Persian may refer to: * People and things from Iran, historically called ''Persia'' in the English language ** Persians, the majority ethnic group in Iran, not to be conflated with the Iranic peoples ** Persian language, an Iranian language of the ...
''līmūn'', a generic term for
citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...
fruit, which is a cognate of Sanskrit (''nimbū'', '
lime Lime commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a color between yellow and green Lime may also refer to: Botany ...
').


Varieties

The 'Bonnie Brae' is oblong, smooth, thin-skinned and seedless. These are mostly grown in
San Diego County San Diego County (), officially the County of San Diego, is a county in the southwestern corner of the U.S. state of California. As of the 2020 census, the population was 3,298,634, making it California's second-most populous county and the f ...
, USA. The 'Eureka' grows year-round and abundantly. This is the common supermarket lemon, also known as 'Four Seasons' (''Quatre Saisons'') because of its ability to produce fruit and flowers together throughout the year. This variety is also available as a plant to domestic customers. There is also a pink-fleshed Eureka lemon, with a green and yellow
variegated Variegation is the appearance of differently coloured zones in the leaves and sometimes the stems and fruit of plants. Species with variegated individuals are sometimes found in the understory of tropical rainforests, and this habitat is the s ...
outer skin. The Lisbon lemon is very similar to the Eureka and is the other common supermarket lemon. It is smoother than the Eureka, has thinner skin, and has fewer or no seeds. It generally produces more juice than the Eureka. The 'Femminello St. Teresa', or 'Sorrento' originates in Italy. This fruit's zest is high in lemon oils. It is the variety traditionally used in the making of '' limoncello''. The 'Yen Ben' is an Australasian cultivar.


Nutrition and phytochemicals

Lemon is a rich source of
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) ...
, providing 64% of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of healthy ...
in a 100 g reference amount (table). Other essential nutrients are low in content. Lemons contain numerous phytochemicals, including
polyphenol Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some o ...
s,
terpene Terpenes () are a class of natural products consisting of compounds with the formula (C5H8)n for n > 1. Comprising more than 30,000 compounds, these unsaturated hydrocarbons are produced predominantly by plants, particularly conifers. Terpenes ...
s, and
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
s. Lemon juice contains slightly more
citric acid Citric acid is an organic compound with the chemical formula HOC(CO2H)(CH2CO2H)2. It is a colorless weak organic acid. It occurs naturally in citrus fruits. In biochemistry, it is an intermediate in the citric acid cycle, which occurs in ...
than
lime Lime commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a color between yellow and green Lime may also refer to: Botany ...
juice (about 47 g/L), nearly twice the citric acid of grapefruit juice, and about five times the amount of citric acid found in
orange Orange most often refers to: *Orange (fruit), the fruit of the tree species '' Citrus'' × ''sinensis'' ** Orange blossom, its fragrant flower *Orange (colour), from the color of an orange, occurs between red and yellow in the visible spectrum * ...
juice.


Culinary uses

Lemon juice, rind, and peel are used in a wide variety of foods and drinks. The whole lemon is used to make
marmalade Marmalade is a fruit preserve made from the juice and peel of citrus fruits boiled with sugar and water. The well-known version is made from bitter orange. It is also made from lemons, limes, grapefruits, mandarins, sweet oranges, bergamot ...
,
lemon curd Fruit curd is a dessert spread and topping usually made with citrus fruit, such as lemon, lime, orange, grapefruit or tangerine. Other flavor variations include passion fruit, mango, and berries such as raspberries, cranberries or blackberries ...
and lemon liqueur. Lemon slices and lemon rind are used as a garnish for food and drinks. Lemon
zest Zest may refer to: Common usage * Zest (ingredient), the outer peel of a citrus fruit ** Zester, a tool for preparing zest ** Twist (cocktail garnish), a piece of zest * Zest (positive psychology), a component of character Brands * Zest (brand ...
, the grated outer rind of the fruit, is used to add flavor to baked goods, puddings, rice, and other dishes.


Juice

Lemon juice is used to make
lemonade Lemonade is a sweetened lemon-flavored beverage. There are varieties of lemonade found throughout the world. In North America and South Asia, cloudy still lemonade is the most common variety. There it is traditionally a homemade drink using ...
, soft drinks, and cocktails. It is used in
marinade Marinating is the process of soaking foods in a seasoned, often acidic, liquid before cooking. The origin of the word alludes to the use of brine (''aqua marina'' or sea water) in the pickling process, which led to the technique of adding flav ...
s for fish, where its acid neutralizes
amine In chemistry, amines (, ) are compounds and functional groups that contain a basic nitrogen atom with a lone pair. Amines are formally derivatives of ammonia (), wherein one or more hydrogen Hydrogen is the chemical element wi ...
s in fish by converting them into nonvolatile ammonium salts. In meat, the acid partially hydrolyzes tough collagen fibers,
tenderizing Tenderness is a quality of meat gauging how easily it is chewed or cut. Tenderness is a desirable quality, as tender meat is softer, easier to chew, and generally more palatable than harder meat. Consequently, tender cuts of meat typically command h ...
it. In the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
, lemon juice is frequently added to
pancakes A pancake (or hotcake, griddlecake, or flapjack) is a flat cake, often thin and round, prepared from a starch-based batter that may contain eggs, milk and butter and cooked on a hot surface such as a griddle or frying pan, often frying w ...
, especially on
Shrove Tuesday Shrove Tuesday is the day before Ash Wednesday (the first day of Lent), observed in many Christian countries through participating in confession and absolution, the ritual burning of the previous year's Holy Week palms, finalizing one's Lenten ...
. Lemon juice is also used as a short-term
preservative A preservative is a substance or a chemical that is added to products such as food products, beverages, pharmaceutical drugs, paints, biological samples, cosmetics, wood, and many other products to prevent decomposition by microbial growth or by ...
on certain foods that tend to oxidize and turn brown after being sliced ( enzymatic browning), such as
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus ''Malus''. The tree originated in Central Asia, where its wild ancestor, ' ...
s, bananas, and
avocado The avocado (''Persea americana'') is a medium-sized, evergreen tree in the laurel family ( Lauraceae). It is native to the Americas and was first domesticated by Mesoamerican tribes more than 5,000 years ago. Then as now it was prized for ...
s, where its acid denatures the enzymes.


Peel

In
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
, lemons are preserved in jars or barrels of salt. The salt penetrates the peel and rind, softening them, and curing them so that they last almost indefinitely. The preserved lemon is used in a wide variety of dishes. Preserved lemons can also be found in Sicilian, Italian, Greek, and French dishes. The peel can be used in the manufacture of pectin, a polysaccharide used as a gelling agent and stabilizer in food and other products.


Oil

Lemon oil is extracted from oil-containing cells in the skin. A machine breaks up the cells, and uses a water spray to flush off the oil. The oil/water mixture is then filtered and separated by centrifugation.


Leaves

The leaves of the lemon tree are used to make a tea and for preparing cooked meats and seafoods.


Other uses


Industrial

Lemons were the primary commercial source of
citric acid Citric acid is an organic compound with the chemical formula HOC(CO2H)(CH2CO2H)2. It is a colorless weak organic acid. It occurs naturally in citrus fruits. In biochemistry, it is an intermediate in the citric acid cycle, which occurs in ...
before the development of fermentation-based processes.


Aroma

Lemon oil may be used in
aromatherapy Aromatherapy is based on the usage of aromatic materials including essential oils and other aroma compounds, with claims for improving psychological and physical well-being. It is offered as a complementary therapy or as a form of alternative ...
. Lemon oil aroma does not influence the human
immune system The immune system is a network of biological processes that protects an organism from diseases. It detects and responds to a wide variety of pathogens, from viruses to parasitic worms, as well as cancer cells and objects such as wood splint ...
, but may contribute to relaxation.


Other

One educational science experiment involves attaching
electrode An electrode is an electrical conductor used to make contact with a nonmetallic part of a circuit (e.g. a semiconductor, an electrolyte, a vacuum or air). Electrodes are essential parts of batteries that can consist of a variety of materials d ...
s to a lemon and using it as a battery to produce electricity. Although very low power, several lemon batteries can power a small digital watch. These experiments also work with other fruits and vegetables. Lemon juice may be used as a simple
invisible ink Invisible ink, also known as security ink or sympathetic ink, is a substance used for writing, which is invisible either on application or soon thereafter, and can later be made visible by some means, such as heat or ultraviolet light. Invisible ...
, developed by heat. Lemon juice can be used to increase the
blonde Blond (male) or blonde (female), also referred to as fair hair, is a hair color characterized by low levels of the dark pigment eumelanin. The resultant visible hue depends on various factors, but always has some yellowish color. The color can ...
colour of hair, acting as a natural highlight after the moistened hair is exposed to sunlight. This is due to the citric acid that acts as bleach.


Horticulture

Lemons need a minimum temperature of around , so they are not hardy year-round in temperate climates, but become hardier as they mature. Citrus require minimal pruning by trimming overcrowded branches, with the tallest branch cut back to encourage bushy growth. Throughout summer, pinching back tips of the most vigorous growth assures more abundant canopy development. As mature plants may produce unwanted, fast-growing shoots (called "water shoots"), these are removed from the main branches at the bottom or middle of the plant. The tradition of urinating near a lemon tree may be beneficial as urine is a fertilizer. In cultivation in the UK, the cultivars "Meyer" and "Variegata" have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's Award of Garden Merit (confirmed 2017).


Production

In 2020, world production of lemons (combined with
limes Limes may refer to: * the plural form of lime (disambiguation) Lime commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a ...
for reporting) was 21.4 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton ( United State ...
s. The top producers – India, Mexico, China, Argentina, Brazil, and Turkey – collectively accounted for 65% of global production (table).


Other citrus called "lemons"

* Flat lemon, a mandarin hybrid. *
Meyer lemon ''Citrus'' × ''meyeri'', the Meyer lemon (), is a hybrid citrus fruit native to China. It is a cross between a citron and a mandarin/pomelo hybrid. Mature trees are around tall with dark green shiny leaves. The flowers are white with a purp ...
, a cross between a citron and a mandarin/pomelo hybrid distinct from sour or sweet orange, named after Frank N. Meyer, who first introduced it to the United States in 1908. Thin-skinned and slightly less acidic than the Lisbon and Eureka lemons, Meyer lemons require more care when shipping and are not widely grown on a commercial basis. Meyer lemons often mature to a yellow-orange color. They are slightly more frost-tolerant. *
Ponderosa lemon The ponderosa lemon (''Citrus'' × ''pyriformis'') (also called Skierniewice lemonKonarska, A., and E. Weryszko-Chmielewska. "Micromorphology, anatomy and ultrastructure of nectaries in two types of flowers of Citrus limon cv.‘Ponderosa’." Ac ...
, more cold-sensitive than true lemons, the fruit are thick-skinned and very large. Genetic analysis showed it to be a complex hybrid of citron and pomelo. * Rough lemon, a citron-mandarin cross, cold-hardy and often used as a citrus rootstock *
Sweet lemon Sweet lemon and Sweet lime refer to groups of citrus hybrids that contain low acid pulp and juice. They are hybrids often similar to non-sweet lemons or limes, but with less citron parentage. Sweet limes and lemons are not sharply separated: ...
s or sweet limes, a mixed group including the lumia (pear lemon), limetta, and Palestinian sweet lime. Among them is the Jaffa lemon, a pomelo-citron hybrid. * Volkamer lemon, like the rough lemon, a citron-mandarin cross


Gallery

File:Flowers blossoms.jpg, Flower File:Lemon-citrus limon seedling.jpg, Lemon seedling File:P2101990,lemon.jpg, Mature lemons File:Citrus limon - Lemon tree - Limonero - Limoeiro.JPG, Full-sized tree File:Citrus x limon 'Variegated Pink' - Lemon.jpg,
Variegated Variegation is the appearance of differently coloured zones in the leaves and sometimes the stems and fruit of plants. Species with variegated individuals are sometimes found in the understory of tropical rainforests, and this habitat is the s ...
pink lemon File:Lemon 2 Bangladesh .jpg, Lemon with leaves


See also

* List of lemon dishes and drinks *


References


External links

* * (with illustrations) {{Authority control Citrus hybrids Cocktail garnishes Tropical agriculture Sour fruits Fruit trees