Jackal
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Jackals are medium-sized canids native to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
and
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelag ...
. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related
black-backed jackal The black-backed jackal (''Lupulella mesomelas),'' also called the silver-backed jackal, is a medium-sized canine native to eastern and southern Africa. These regions are separated by roughly 900 kilometers. One region includes the southe ...
(''Lupulella mesomelas'') and side-striped jackal (''Lupulella adusta'') of sub-Saharan-Africa, and the golden jackal (''Canis aureus'') of south-central Europe and Asia. The African golden wolf (''Canis lupaster'') was also formerly considered as a jackal. While they do not form a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
, all jackals are opportunistic omnivores, predators of small to medium-sized animals and proficient
scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feedin ...
s. Their long legs and curved canine teeth are adapted for hunting small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s, birds, and
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...
s, and their large feet and fused leg bones give them a physique well-suited for long-distance running, capable of maintaining speeds of for extended periods of time. Jackals are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of dayli ...
, most active at dawn and dusk. Their most common social unit is a
monogamous Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time ( serial monogamy) — as compared to the various forms of non-monogamy (e.g., pol ...
pair, which defends its territory from other pairs by vigorously chasing intruding rivals and marking landmarks around the
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
with their
urine Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excreted from the body through the urethra. Cellul ...
and
feces Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a rela ...
. The territory may be large enough to hold some young adults, which stay with their parents until they establish their own territories. Jackals may occasionally assemble in small packs, for example, to scavenge a carcass, but they normally hunt either alone or in pairs.


Etymology

The English word "jackal" dates back to 1600 and derives from the French ''chacal'', from Turkish çakal, derived from the Persian ', which is in turn derived from the
Sanskrit Sanskrit (; attributively , ; nominally , , ) is a classical language belonging to the Indo-Aryan languages, Indo-Aryan branch of the Indo-European languages. It arose in South Asia after its predecessor languages had Trans-cultural diffusion ...
शृगाल ''śṛgāla'' meaning "the howler".


Taxonomy and relationships

Similarities between jackals and coyotes led
Lorenz Oken Lorenz Oken (1 August 1779 – 11 August 1851) was a German naturalist, botanist, biologist, and ornithologist. Oken was born Lorenz Okenfuss (german: Okenfuß) in Bohlsbach (now part of Offenburg), Ortenau, Baden, and studied natural history and ...
, in the third volume of his ''Lehrbuch der Naturgeschichte'' (1815), to place these species into a new separate genus, ''Thos'', named after the classical Greek word "jackal", but his theory had little immediate impact on taxonomy at the time.
Angel Cabrera In various theistic religious traditions an angel is a supernatural spiritual being who serves God. Abrahamic religions often depict angels as benevolent celestial intermediaries between God (or Heaven) and humanity. Other roles incl ...
, in his 1932 monograph on the mammals of
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
, questioned whether or not the presence of a cingulum on the upper molars of the jackals and its corresponding absence in the rest of ''Canis'' could justify a subdivision of that genus. In practice, Cabrera chose the undivided-genus alternative and referred to the jackals as ''Canis'' instead of ''Thos''. Oken's ''Thos'' theory was revived in 1914 by
Edmund Heller Edmund Heller (May 21, 1875 – July 18, 1939) was an American zoologist. He was President of the Association of Zoos & Aquariums for two terms, from 1935-1936 and 1937-1938. Early life While at Stanford University, he collected specimens in th ...
, who embraced the separate genus theory. Heller's names and the designations he gave to various jackal species and subspecies live on in current taxonomy, although the genus has been changed from ''Thos'' to ''Canis''.Thos vs Canis
The wolf-like canids are a group of large carnivores that are genetically closely related because they all have 78
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
s. The group includes genus ''Canis'', ''Cuon'', and ''Lycaon''. The members are the dog ''(C. lupus familiaris)'',
gray wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
(''C. lupus''),
coyote The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological nich ...
(''C. latrans''), golden jackal (''C. aureus''), Ethiopian wolf (''C. simensis''),
black-backed jackal The black-backed jackal (''Lupulella mesomelas),'' also called the silver-backed jackal, is a medium-sized canine native to eastern and southern Africa. These regions are separated by roughly 900 kilometers. One region includes the southe ...
(''C. mesomelas''), side-striped jackal (''C. adustus''), dhole (''Cuon alpinus''), and African wild dog (''Lycaon pictus''). The latest recognized member is the
African wolf The African wolf (''Canis lupaster'') or golden wolf, formerly known as the African golden jackal, is a canine native to North Africa, West Africa, the Sahel, northern East Africa, and the Horn of Africa. It is listed as least concern on the IU ...
(''C. lupaster''), which was once thought to be an African branch of the golden jackal. As they possess 78 chromosomes, all members of the genus '' Canis'' are karyologically indistinguishable from each other, and from the dhole and the African hunting dog. The two African jackals are shown to be the most basal members of this clade, indicating the clade's origin from Africa. ''
Canis arnensis ''Canis arnensis'', the Arno River dog, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. The Arno River dog has been described as a small jackal-like dog. Its anatomy and morphology relate it ...
'' arrived in Mediterranean Europe 1.9 million years ago and is probably the ancestor of modern jackals. The paraphyletic nature of ''Canis'' with respect to '' Lycaon'' and ''
Cuon The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. ...
'' has led to suggestions that the two African jackals should be assigned to different genera, ''Schaeffia'' for the side-striped jackal and ''Lupulella'' for the black-backed jackal or ''Lupulella'' for both. The intermediate size and shape of the Ethiopian wolf has at times led it to be regarded as a jackal, thus it has also been called the "red jackal" or the "Simien jackal".


Species


Folklore and literature

Like foxes and coyotes, jackals are often depicted as clever sorcerers in the myths and legends of their regions. They are mentioned roughly 14 times in the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity Christianity is an Abrahamic monotheistic religion based on the life and teachings of Jesus ...
. It is frequently used as a literary device to illustrate desolation, loneliness, and abandonment, with reference to its habit of living in the ruins of former cities and other areas abandoned by humans. It is called "wild dog" in several translations of the Bible. In the
King James Bible The King James Version (KJV), also the King James Bible (KJB) and the Authorized Version, is an English translation of the Christian Bible for the Church of England, which was commissioned in 1604 and published in 1611, by sponsorship of ...
, Isaiah 13:21 refers to 'doleful creatures', which some commentators suggest are either jackals or hyenas. In the Indian ''
Panchatantra The ''Panchatantra'' (IAST: Pañcatantra, ISO: Pañcatantra, sa, पञ्चतन्त्र, "Five Treatises") is an ancient Indian collection of interrelated animal fables in Sanskrit verse and prose, arranged within a frame story ...
'' stories, the jackal is mentioned as wily and wise. In Bengali tantrik tradition, they represent the goddess
Kali Kali (; sa, काली, ), also referred to as Mahakali, Bhadrakali, and Kalika ( sa, कालिका), is a Hindu goddess who is considered to be the goddess of ultimate power, time, destruction and change in Shaktism. In this tra ...
. It is said she appears as jackals when meat is offered to her. The
Serer religion The Serer religion, or ''a ƭat Roog'' ("the way of the Divine"), is the original religious beliefs, practices, and teachings of the Serer people of Senegal in West Africa. The Serer religion believes in a universal supreme deity called Roog ...
and
creation myth A creation myth (or cosmogonic myth) is a symbolic narrative of how the world began and how people first came to inhabit it., "Creation myths are symbolic stories describing how the universe and its inhabitants came to be. Creation myths develo ...
posits jackals were among the first animals created by Roog, the supreme deity of the
Serer people The Serer people are a West African ethnoreligious group.
.


References


Further reading

* ''The New Encyclopedia of Mammals'' edited by David Macdonald, Oxford University Press, 2001; * ''Cry of the Kalahari'', by Mark and Delia Owens, Mariner Books, 1992. * ''The Velvet Claw: A Natural History of the Carnivores'', by David MacDonald, BBC Books, 1992. * ''Foxes, Wolves, and Wild Dogs of the World'', by David Alderton, Facts on File, 2004.


External links


Jackal
at the African Wildlife Foundation
Jackals
at A-Z Animals
''Jackals of the African Crater''
at PBS.org
Jackal is a meaningless term
{{DEFAULTSORT:Jackal Jackals Mammals described in 1758 Taxa named by Carl Linnaeus Mammal common names