Hymenaea
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''Hymenaea'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of plants in the legume family
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
. Of the fourteen living species in the genus, all but one are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to the
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also refer ...
of the Americas, with one additional species (''
Hymenaea verrucosa ''Hymenaea verrucosa'' (Zanzibar copal, East African copal, or Amber tree) is a species of flowering plant in the legume family, Fabaceae. It belongs to the paraphyletic subfamily Caesalpinioideae. It is a large tree native to the tropical regio ...
'') on the east coast of Africa. Some authors place the African species in a separate
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
genus, ''
Trachylobium ''Hymenaea'' is a genus of plants in the legume family Fabaceae. Of the fourteen living species in the genus, all but one are native to the tropics of the Americas, with one additional species (''Hymenaea verrucosa'') on the east coast of Afr ...
''.Gwilym Lewis, Brian Schrire, Barbara MacKinder, and Mike Lock. 2005. ''Legumes of the World''. Royal Botanic Gardens, Kew: Richmond, England. In the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
, ''Hymenaea'' is distributed through the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...
islands, and from southern
Mexico Mexico (Spanish language, Spanish: México), officially the United Mexican States, is a List of sovereign states, country in the southern portion of North America. It is borders of Mexico, bordered to the north by the United States; to the so ...
to
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
named the genus in 1753 in ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' for
Hymenaios Hymen ( grc, Ὑμήν), Hymenaios or Hymenaeus, in Hellenistic religion, is a god of marriage ceremonies, inspiring feasts and song. Related to the god's name, a ''hymenaios'' is a genre of Greek lyric poetry sung during the procession of the ...
, the Greek god of marriage ceremonies. The name is a reference to the paired leaflets. Most species of ''Hymenaea'' are large
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s and they are primarily
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
. They may grow to a height of and emerge above the
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
. Some species will grow both as tall forest trees and as smaller shrubby trees depending on their surrounding
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are pinnately bifoliolate, meaning that they have two leaflets attached to the sides of the petiole. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s grow in a
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
or
corymb Corymb is a botanical term for an inflorescence with the flowers growing in such a fashion that the outermost are borne on longer pedicels than the inner, bringing all flowers up to a common level. A corymb has a flattish top with a superficial ...
type of
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
.


Uses and properties

The pulpy center of the
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
s is edible and contains starch. The fruit is sold in local markets in the Americas. The leaves may be used to make a tea. The trees produce a dense
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
used for
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
in making ships and furniture. The thick bark of some species is used by indigenous peoples of the Amazon to make
canoe A canoe is a lightweight narrow water vessel, typically pointed at both ends and open on top, propelled by one or more seated or kneeling paddlers facing the direction of travel and using a single-bladed paddle. In British English, the ter ...
s. Seeds contain large amounts (40% of dry weight) of a highly viscous polysaccharide (
xyloglucan Xyloglucan is a hemicellulose that occurs in the primary cell wall of all vascular plants; however, all enzymes responsible for xyloglucan metabolism are found in Charophyceae algae.LEV Del Bem and M Vincentz (2010) Evolution of xyloglucan-relate ...
) which can be used in several industrial sectors such as food, paper, cosmetic and pharmaceutical. The trees also make hard
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...
s that are used to manufacture varnish, especially the resin from ''Hymenaea courbaril'' (jatobá) in Brazil. The resin that is produced in Brazil is known as
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
n
copal Copal is tree resin, particularly the aromatic resins from the copal tree ''Protium copal'' (Burseraceae) used by the cultures of pre-Columbian Mesoamerica as ceremonially burned incense and for other purposes. More generally, copal includes re ...
, and ''
Hymenaea verrucosa ''Hymenaea verrucosa'' (Zanzibar copal, East African copal, or Amber tree) is a species of flowering plant in the legume family, Fabaceae. It belongs to the paraphyletic subfamily Caesalpinioideae. It is a large tree native to the tropical regio ...
'' is the source of the valuable
Zanzibar Zanzibar (; ; ) is an insular semi-autonomous province which united with Tanganyika in 1964 to form the United Republic of Tanzania. It is an archipelago in the Indian Ocean, off the coast of the mainland, and consists of many small islan ...
copal. Resin may be collected from living trees, or from the soil near the place where a tree once stood. Throughout its American range, indigenous peoples use the resin for
incense Incense is aromatic biotic material that releases fragrant smoke when burnt. The term is used for either the material or the aroma. Incense is used for aesthetic reasons, religious worship, aromatherapy, meditation, and ceremony. It may also b ...
and as a cement. Resin from the extinct species ''
Hymenaea protera ''Hymenaea protera'' is an extinct prehistoric leguminous tree, the probable ancestor of present-day '' Hymenaea'' species. Most neotropical ambers come from its fossilized resin, including the famous Dominican amber. ''H. protera'' once grew in ...
'' is the source of Dominican amber, while the extinct ''
Hymenaea mexicana ''Hymenaea mexicana'' is an extinct legume species in the family Fabaceae described from a series of isolated fossil petals, leaflets, and amber. The species is known from a group of Late Oligocene to Early Miocene locations in southern Mexico. ...
'' produced the resin which is the source of Mexican amber. ''
Hymenaea courbaril ''Hymenaea courbaril'', the courbaril or West Indian locust, is a tree common in the Caribbean, Central America, and South America. It is a hardwood that is used for furniture, flooring, and decoration. Its hard fruit pods have edible dry pulp ...
'' has been used as a
model organism A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workin ...
to study the effect of increased concentration on the rate of
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
in
neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
regions. When the concentration of was increased from an ambient reference level of 360 ppm to 720 ppm, the photosynthetic assimilation in the seedlings doubled. This suggests the species could play an important role in greenhouse gas sequestration, as atmospheric is expected to reach ca. 700 ppm by the year 2075 if current levels of fossil fuel consumption are maintained. ''Hymenaea courbaril'' is a very important species in programmes of recuperation of degraded
rain forests Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest, ...
in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
. It appears late in the natural regeneration process being classified as a 'late successional' or
climax species Climax species, also called late seral, late-successional, K-selected or equilibrium species, are plant species that can germinate and grow with limited resources; e.g., they need heat exposure or low water availability. They are the species wi ...
.


Species

*'' Hymenaea allendis'' *'' Hymenaea aurea'' *''
Hymenaea courbaril ''Hymenaea courbaril'', the courbaril or West Indian locust, is a tree common in the Caribbean, Central America, and South America. It is a hardwood that is used for furniture, flooring, and decoration. Its hard fruit pods have edible dry pulp ...
'' *'' Hymenaea eriogyne'' *'' Hymenaea intermedia'' *'' Hymenaea maranhensis'' *''
Hymenaea martiana ''Hymenaea martiana'' is a tree species in the genus '' Hymenaea'' found in Brazil (Alagoas, Bahia, Ceara, Goias, Mato Grosso, Minas Gerais, Pernambuco) and Paraguay. Chemicals The three rhamnosides eucryphin, astilbin Astilbin is a flavanonol ...
'' *''
Hymenaea mexicana ''Hymenaea mexicana'' is an extinct legume species in the family Fabaceae described from a series of isolated fossil petals, leaflets, and amber. The species is known from a group of Late Oligocene to Early Miocene locations in southern Mexico. ...
'' *'' Hymenaea oblongifolia'' *'' Hymenaea parvifolia'' *''
Hymenaea protera ''Hymenaea protera'' is an extinct prehistoric leguminous tree, the probable ancestor of present-day '' Hymenaea'' species. Most neotropical ambers come from its fossilized resin, including the famous Dominican amber. ''H. protera'' once grew in ...
'' *'' Hymenaea reticulata'' *'' Hymenaea rubriflora'' *'' Hymenaea stigonocarpa'' *'' Hymenaea torrei'' *'' Hymenaea velutina'' *''
Hymenaea verrucosa ''Hymenaea verrucosa'' (Zanzibar copal, East African copal, or Amber tree) is a species of flowering plant in the legume family, Fabaceae. It belongs to the paraphyletic subfamily Caesalpinioideae. It is a large tree native to the tropical regio ...
'' †Extinct


See also

* Animé, an oleo-resin from ''Hymenaea'' species.


References


Further reading

* * * * * * * * * {{Taxonbar, from=Q290340 Fabaceae genera