The humerus (; ) is a

long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the load during daily activities ...

in the arm that runs from the shoulder to the elbow. It connects the

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eith ...

and the two bones of the lower arm, the

radius In classical geometry, a radius (plural, : radii) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', ...

and

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

, and consists of three sections. The humeral upper extremity consists of a rounded head, a narrow neck, and two short processes (

tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...

s, sometimes called tuberosities). The body is cylindrical in its upper portion, and more prismatic below. The

lower extremity The human leg, in the general word sense, is the entire lower limb of the human body, including the foot, thigh or sometimes even the hip or gluteal region. However, the definition in human anatomy refers only to the section of the lower limb ...

consists of 2

epicondyle An epicondyle () is a rounded eminence on a bone that lies upon a condyle ('' epi-'', "upon" + ''condyle'', from a root meaning "knuckle" or "rounded articular area"). There are various epicondyles in the human skeleton, each named by its anatomic ...

s, 2 processes ( trochlea & capitulum), and 3 fossae ( radial fossa,

coronoid fossa Superior to the anterior portion of the trochlea is a small depression, the coronoid fossa, which receives the coronoid process of the ulna during flexion of the forearm. It is directly adjacent to the radial fossa of the humerus The humerus ...

, and olecranon fossa). As well as its true anatomical neck, the constriction below the greater and lesser tubercles of the humerus is referred to as its surgical neck due to its tendency to fracture, thus often becoming the focus of surgeons.

Etymology

The word "humerus" is derived from la, humerus, umerus meaning upper arm, shoulder, and is linguistically related to

Gothic Gothic or Gothics may refer to: People and languages *Goths or Gothic people, the ethnonym of a group of East Germanic tribes **Gothic language, an extinct East Germanic language spoken by the Goths **Crimean Gothic, the Gothic language spoken b ...

''ams'' shoulder and Greek ''ōmos''.

Structure

Upper extremity

The upper or proximal extremity of the humerus consists of the bone's large rounded head joined to the body by a constricted portion called the neck, and two eminences, the greater and lesser tubercles.

Head

The head (''caput humeri''), is nearly hemispherical in form. It is directed upward, medialward, and a little backward, and articulates with the glenoid cavity of the scapula to form the glenohumeral joint (shoulder joint). The circumference of its articular surface is slightly constricted and is termed the anatomical neck, in contradistinction to a constriction below the tubercles called the surgical neck which is frequently the seat of fracture. Fracture of the anatomical neck rarely occurs.''

Gray's Anatomy ''Gray's Anatomy'' is a reference book of human anatomy written by Henry Gray, illustrated by Henry Vandyke Carter, and first published in London in 1858. It has gone through multiple revised editions and the current edition, the 42nd (Octo ...

'', see infobox The diameter of the humeral head is generally larger in men than in women. Humerus-Head

Anatomical neck

The anatomical neck (''collum anatomicum'') is obliquely directed, forming an obtuse angle with the body. It is best marked in the lower half of its circumference; in the upper half it is represented by a narrow groove separating the head from the tubercles. The line separating the head from the rest of the upper end is called the ''anatomical neck.'' It affords attachment to the articular capsule of the shoulder-joint, and is perforated by numerous vascular foramina. Fracture of the anatomical neck rarely occurs. The anatomical neck of the humerus is an indentation distal to the head of the humerus on which the articular capsule attaches. Neck-of-Humerus

Surgical neck

The surgical neck is a narrow area distal to the tubercles that is a common site of fracture. It makes contact with the axillary nerve and the posterior humeral circumflex artery.

Greater tubercle

The greater tubercle (''tuberculum majus''; greater tuberosity) is a large, posteriorly placed projection that is placed laterally. The greater tubercle is where ''supraspinatus'', ''infraspinatus'' and ''teres minor'' muscles are attached. The crest of the greater tubercle forms the lateral lip of the bicipital groove and is the site for insertion of ''pectoralis major''. The greater tubercle is just lateral to the anatomical neck. Its upper surface is rounded and marked by three flat impressions: the highest of these gives insertion to the supraspinatus muscle; the middle to the infraspinatus muscle; the lowest one, and the body of the bone for about 2.5 cm. below it, to the teres minor muscle. The lateral surface of the greater tubercle is convex, rough, and continuous with the lateral surface of the body. Greater-Tubercle-of-Right-Humerus

Lesser tubercle

The lesser tubercle (''tuberculum minus''; lesser tuberosity) is smaller, anterolaterally placed to the head of the humerus. The lesser tubercle provides insertion to ''subscapularis'' muscle. Both these tubercles are found in the proximal part of the shaft. The crest of the lesser tubercle forms the medial lip of the bicipital groove and is the site for insertion of ''teres major'' and ''latissimus dorsi'' muscles. The lesser tuberosity, is more prominent than the greater: it is situated in front, and is directed medialward and forward. Above and in front it presents an impression for the insertion of the tendon of the subscapularis muscle. Lesser-Tubercle-of-Right-Humerus

Bicipital groove

The tubercles are separated from each other by a deep groove, the

bicipital groove The bicipital groove (intertubercular groove, sulcus intertubercularis) is a deep groove on the humerus that separates the greater tubercle from the lesser tubercle. It allows for the long tendon of the biceps brachii muscle to pass. Structure ...

(intertubercular groove; bicipital sulcus), which lodges the long tendon of the biceps brachii muscle and transmits a branch of the

anterior humeral circumflex artery The anterior humeral circumflex artery (anterior circumflex artery, anterior circumflex humeral artery) is an artery in the arm. It is one of two circumflexing arteries that branch from the axillary artery, the other being the posterior humeral cir ...

to the shoulder-joint. It runs obliquely downward, and ends near the junction of the upper with the middle third of the bone. In the fresh state its upper part is covered with a thin layer of cartilage, lined by a prolongation of the

synovial membrane The synovial membrane (also known as the synovial stratum, synovium or stratum synoviale) is a specialized connective tissue that lines the inner surface of capsules of synovial joints and tendon sheath. It makes direct contact with the fibrous ...

of the shoulder-joint; its lower portion gives insertion to the tendon of the latissimus dorsi muscle. It is deep and narrow above, and becomes shallow and a little broader as it descends. Its lips are called, respectively, the crests of the greater and lesser tubercles (''bicipital ridges''), and form the upper parts of the anterior and medial borders of the body of the bone. Bicipital-groove

Shaft

The body or shaft of the humerus is triangular to cylindrical in cut section and is compressed anteroposteriorly. It has 3 surfaces, namely: *Anterolateral surface: the area between the lateral border of the humerus to the line drawn as a continuation of the crest of the greater tubercle. The antero-lateral surface is directed lateralward above, where it is smooth, rounded, and covered by the deltoid muscle; forward and lateralward below, where it is slightly concave from above downward, and gives origin to part of the

Brachialis The brachialis (brachialis anticus), also known as the Teichmann muscle, is a muscle in the upper arm that flexes the elbow. It lies deeper than the biceps brachii, and makes up part of the floor of the region known as the cubital fossa (elbow ...

. About the middle of this surface is a rough, rectangular elevation, the

deltoid tuberosity In human anatomy, the deltoid tuberosity is a rough, triangular area on the anterolateral (front-side) surface of the middle of the humerus. It is a site of attachment of deltoid muscle. Structure Variation The deltoid tuberosity has been re ...

for the insertion of the deltoid muscle; below this is the

radial sulcus The radial groove (also known as the musculospiral groove, radial sulcus, or spiral groove) is a broad but shallow oblique depression for the radial nerve and deep brachial artery. It is located on the center of the lateral border of the humerus b ...

, directed obliquely from behind, forward, and downward, and transmitting the

radial nerve The radial nerve is a nerve in the human body that supplies the posterior portion of the upper limb. It innervates the medial and lateral heads of the triceps brachii muscle of the arm, as well as all 12 muscles in the posterior osteofascial comp ...

and profunda artery. *Anteromedial surface: the area between the medial border of the humerus to the line drawn as a continuation of the crest of the greater tubercle. The antero-medial surface, less extensive than the antero-lateral, is directed medialward above, forward and medialward below; its upper part is narrow, and forms the floor of the intertubercular groove which gives insertion to the tendon of the latissimus dorsi muscle; its middle part is slightly rough for the attachment of some of the fibers of the tendon of insertion of the coracobrachialis muscle; its lower part is smooth, concave from above downward, and gives origin to the brachialis muscle. *Posterior surface: the area between the medial and lateral borders. The posterior surface appears somewhat twisted, so that its upper part is directed a little medialward, its lower part backward and a little lateralward. Nearly the whole of this surface is covered by the lateral and medial heads of the Triceps brachii, the former arising above, the latter below the radial sulcus. Its three borders are: *Anterior: the anterior border runs from the front of the greater tubercle above to the

below, separating the antero-medial from the antero-lateral surface. Its upper part is a prominent ridge, the crest of the greater tubercle; it serves for the insertion of the tendon of the

pectoralis major muscle The pectoralis major () is a thick, fan-shaped or triangular convergent muscle, situated at the chest of the human body. It makes up the bulk of the chest muscles and lies under the breast. Beneath the pectoralis major is the pectoralis minor ...

. About its center it forms the anterior boundary of the

, on which the deltoid muscle attaches; below, it is smooth and rounded, affording attachment to the brachialis muscle. *Lateral: the lateral border runs from the back part of the greater tubercle to the lateral

, and separates the anterolateral from the posterior surface. Its upper half is rounded and indistinctly marked, serving for the attachment of the lower part of the insertion of the teres minor muscle, and below this giving origin to the lateral head of the

triceps brachii muscle The triceps, or triceps brachii (Latin for "three-headed muscle of the arm"), is a large muscle on the back of the upper limb of many vertebrates. It consists of 3 parts: the medial, lateral, and long head. It is the muscle principally respon ...

; its center is traversed by a broad but shallow oblique depression, the spiral groove (''musculospiral groove''). The

runs in the spiral groove. Its lower part forms a prominent, rough margin, a little curved from backward, forward the

lateral supracondylar ridge The lateral supracondylar ridge is a prominent, rough margin on the lower part of the lateral border of the humerus. It presents an anterior lip for the origin of forearm extensors, including the brachioradialis muscle above, and the extensor car ...

, which presents an anterior lip for the origin of the

brachioradialis muscle The brachioradialis is a muscle of the forearm that flexes the forearm at the elbow. It is also capable of both pronation and supination, depending on the position of the forearm. It is attached to the distal styloid process of the radius by way o ...

above, and

extensor carpi radialis longus muscle The extensor carpi radialis longus is one of the five main muscles that control movements at the wrist. This muscle is quite long, starting on the lateral side of the humerus, and attaching to the base of the second metacarpal bone (metacarpal of ...

above, a posterior lip for the

, and an intermediate ridge for the attachment of the lateral intermuscular septum. *Medial: the medial border extends from the lesser tubercle to the medial epicondyle. Its upper third consists of a prominent ridge, the crest of the lesser tubercle, which gives insertion to the tendon of the teres major muscle. About its center is a slight impression for the insertion of the coracobrachialis muscle, and just below this is the entrance of the nutrient canal, directed downward; sometimes there is a second nutrient canal at the commencement of the

. The inferior third of this border is raised into a slight ridge, the

medial supracondylar ridge The inferior third of the medial border of the humerus is raised into a slight ridge, the medial supracondylar ridge (or medial supracondylar line), which becomes very prominent below; it presents an anterior lip for the origins of the Brachiali ...

, which became very prominent below; it presents an anterior lip for the origins of the brachialis muscle and the pronator teres muscle, a posterior lip for the medial head of the

, and an intermediate ridge for the attachment of the medial intermuscular septum. Anterior-view-of-the-humerus-showing-borders-and-surfaces

Posterior-view-of-the-humerus-showing-borders-and-surfaces

The Deltoid tuberosity is a roughened surface on the lateral surface of the shaft of the Humerus and acts as the site of insertion of ''deltoideus'' muscle. The posterorsuperior part of the shaft has a crest, beginning just below the surgical neck of the humerus and extends till the superior tip of the deltoid tuberosity. This is where the lateral head of ''triceps brachii'' is attached. Deltoid-Tuberosity

The radial sulcus, also known as the spiral groove is found on the posterior surface of the shaft and is a shallow oblique groove through which the radial nerve passes along with deep vessels. This is located posteroinferior to the deltoid tuberosity. The inferior boundary of the spiral groove is continuous distally with the lateral border of the shaft. Radial-groove-continuing-as-lateral-border-of-shaft

Radial-groove-continuing-as-lateral-border-of-shaft

The nutrient foramen of the humerus is located in the anteromedial surface of the humerus. The nutrient arteries enter the humerus through this foramen. Nutrient-Foramen

Distal humerus

The distal or lower extremity of the humerus is flattened from before backward, and curved slightly forward; it ends below in a broad, articular surface, which is divided into two parts by a slight ridge. Projecting on either side are the lateral and medial

Articular surface

The articular surface extends a little lower than the epicondyles, and is curved slightly forward; its medial extremity occupies a lower level than the lateral. The ''lateral'' portion of this surface consists of a smooth, rounded eminence, named the capitulum of the humerus; it articulates with the cup-shaped depression on the head of the radius, and is limited to the front and lower part of the bone.

Fossae

Above the front part of the trochlea is a small depression, the

, which receives the coronoid process of the ulna during flexion of the forearm. Above the back part of the trochlea is a deep triangular depression, the olecranon fossa, in which the summit of the olecranon is received in

extension Extension, extend or extended may refer to: Mathematics Logic or set theory * Axiom of extensionality * Extensible cardinal * Extension (model theory) * Extension (predicate logic), the set of tuples of values that satisfy the predicate * Ext ...

of the forearm. Olecranon-fossa

The coronoid fossa is the medial hollow part on the anterior surface of the distal humerus. The coronoid fossa is smaller than the olecranon fossa and receives the coronoid process of the ulna during maximum flexion of the elbow. Coronoid-Fossa

Above the front part of the capitulum is a slight depression, the radial fossa, which receives the anterior border of the head of the radius, when the forearm is flexed. Radial-Fossa

These fossæ are separated from one another by a thin, transparent lamina of bone, which is sometimes perforated by a

supratrochlear foramen The supratrochlear foramen is a small hole located above the trochlea of the humerus of several mammals, it is found frequently in dogs and sometimes in humans, especially women and in the left humerus. It is covered by a layer of connective tiss ...

; they are lined in the fresh state by the synovial membrane of the elbow-joint, and their margins afford attachment to the anterior and posterior ligaments of this articulation. The Capitulum is a rounded eminence forming the lateral part of the distal humerus. The head of the radius articulates with the capitulum. Capitulum-and-trochlea

The trochlea is spool-shaped medial portion of the distal humerus and articulates with the ulna.

Epicondyles

The epicondyles are continuous above with the supracondylar ridges. * The lateral epicondyle is a small, tuberculated eminence, curved a little forward, and giving attachment to the radial collateral ligament of the elbow-joint, and to a tendon common to the origin of the

Supinator In human anatomy, the supinator is a broad muscle in the posterior compartment of the forearm, curved around the upper third of the radius. Its function is to supinate the forearm. Structure Supinator consists of two planes of fibers, between whic ...

and some of the Extensor muscles. * The medial epicondyle, larger and more prominent than the lateral, is directed a little backward; it gives attachment to the ulnar collateral ligament of the elbow-joint, to the

Pronator teres The pronator teres is a muscle (located mainly in the forearm) that, along with the pronator quadratus, serves to pronate the forearm (turning it so that the palm faces posteriorly when from the anatomical position). It has two attachments, to t ...

, and to a common tendon of origin of some of the Flexor muscles of the forearm; the ulnar nerve runs in a groove on the back of this epicondyle. Medial-epicondyle-and-lateral-epicondyle

Medial-epicondyle-and-lateral-epicondyle

Medial-supracondylar-crest-and-lateral-supracondylar-crest

The Medial supracondylar crest forms the sharp medial border of the distal humerus continuing superiorly from the medial epicondyle. The Lateral Supracondylar crest forms the sharp lateral border of the distal humerus continuing superiorly from the lateral epicondyle.

Borders

The medial portion of the articular surface is named the trochlea, and presents a deep depression between two well-marked borders; it is convex from before backward, concave from side to side, and occupies the anterior, lower, and posterior parts of the extremity. * The ''lateral border'' separates it from the groove which articulates with the margin of the head of the radius. * The ''medial border'' is thicker, of greater length, and consequently more prominent, than the lateral. The grooved portion of the articular surface fits accurately within the semilunar notch of the ulna; it is broader and deeper on the posterior than on the anterior aspect of the bone, and is inclined obliquely downward and forward toward the medial side.

Articulations

At the shoulder, the head of the humerus articulates with the glenoid fossa of the

. More distally, at the elbow, the capitulum of the humerus articulates with the head of the

, and the trochlea of the humerus articulates with the trochlear notch of the

Nerves

The axillary nerve is located at the proximal end, against the shoulder girdle. Dislocation of the humerus's

glenohumeral joint The shoulder joint (or glenohumeral joint from Greek ''glene'', eyeball, + -''oid'', 'form of', + Latin ''humerus'', shoulder) is structurally classified as a synovial ball-and-socket joint and functionally as a diarthrosis and multiaxial joint ...

has the potential to injure the axillary nerve or the axillary artery. Signs and symptoms of this dislocation include a loss of the normal shoulder contour and a palpable depression under the acromion. The

follows the humerus closely. At the midshaft of the humerus, the radial nerve travels from the posterior to the anterior aspect of the bone in the ''spiral groove''. A fracture of the humerus in this region can result in radial nerve injury. The ulnar nerve lies at the distal end of the humerus near the elbow. When struck, it can cause a distinct tingling sensation, and sometimes a significant amount of pain. It is sometimes popularly referred to as 'the funny bone', possibly due to this sensation (a "funny" feeling), as well as the fact that the bone's name is a homophone of 'humorous'. It lies posterior to the medial epicondyle, and is easily damaged in elbow injuries.

Function

Muscular attachment

The

deltoid Deltoid (delta-shaped) can refer to: * The deltoid muscle, a muscle in the shoulder * Kite (geometry), also known as a deltoid, a type of quadrilateral * A deltoid curve, a three-cusped hypocycloid * A leaf shape * The deltoid tuberosity, a part o ...

originates on the lateral third of the

clavicle The clavicle, or collarbone, is a slender, S-shaped long bone approximately 6 inches (15 cm) long that serves as a strut between the shoulder blade and the sternum (breastbone). There are two clavicles, one on the left and one on the r ...

acromion In human anatomy, the acromion (from Greek: ''akros'', "highest", ''ōmos'', "shoulder", plural: acromia) is a bony process on the scapula (shoulder blade). Together with the coracoid process it extends laterally over the shoulder joint. The ...

and the crest of the

spine Spine or spinal may refer to: Science Biology * Vertebral column, also known as the backbone * Dendritic spine, a small membranous protrusion from a neuron's dendrite * Thorns, spines, and prickles, needle-like structures in plants * Spine (zoolo ...

of the scapula. It is inserted on the

of the humerus and has several actions including abduction, extension, and circumduction of the shoulder. The supraspinatus also originates on the spine of the scapula. It inserts on the greater tubercle of the humerus, and assists in abduction of the shoulder. The

pectoralis major The pectoralis major () is a thick, fan-shaped or triangular convergent muscle, situated at the chest of the human body. It makes up the bulk of the chest muscles and lies under the breast. Beneath the pectoralis major is the pectoralis minor, ...

, teres major, and

latissimus dorsi The latissimus dorsi () is a large, flat muscle on the back that stretches to the sides, behind the arm, and is partly covered by the trapezius on the back near the midline. The word latissimus dorsi (plural: ''latissimi dorsorum'') comes from ...

insert at the ''intertubercular groove'' of the humerus. They work to adduct and medially, or internally, rotate the humerus. The infraspinatus and teres minor insert on the greater tubercle, and work to laterally, or externally, rotate the humerus. In contrast, the subscapularis muscle inserts onto the lesser tubercle and works to medially, or internally, rotate the humerus. The

biceps brachii The biceps or biceps brachii ( la, musculus biceps brachii, "two-headed muscle of the arm") is a large muscle that lies on the front of the upper arm between the shoulder and the elbow. Both heads of the muscle arise on the scapula and join ...

brachialis The brachialis (brachialis anticus), also known as the Teichmann muscle, is a muscle in the upper arm that flexes the elbow. It lies deeper than the biceps brachii, and makes up part of the floor of the region known as the cubital fossa (elbow ...

, and

brachioradialis The brachioradialis is a muscle of the forearm that flexes the forearm at the elbow. It is also capable of both pronation and supination, depending on the position of the forearm. It is attached to the distal styloid process of the radius by way ...

(which attaches distally) act to flex the elbow. (The biceps do not attach to the humerus.) The triceps brachii and anconeus extend the elbow, and attach to the posterior side of the humerus. The four muscles of supraspinatus, infraspinatus, teres minor and subscapularis form a musculo-ligamentous girdle called the rotator cuff. This cuff stabilizes the very mobile but inherently unstable

. The other muscles are used as counterbalances for the actions of lifting/pulling and pressing/pushing.

Other animals

Primitive fossils of

amphibian Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbo ...

s had little, if any, shaft connecting the upper and lower extremities, making their limbs very short. In most living

tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids ( pelycosaurs, extinct t ...

s, however, the humerus has a similar form to that of humans. In many

reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...

s and some

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...

s (where it is the primitive state), the lower extremity includes a large opening called the entepicondylar foramen to allow the passage of nerves and blood vessels.

Additional images

File:Humerus animation.gif, Position of humerus (shown in red). Animation. File:Left humerus - close-up - animation.gif, Left humerus. Animation. File:Human humerus 2.stl, 3D image File:Human arm bones diagram.svg, Human arm bones diagram. File:Inferior epiphysis of humerus.jpg, Humerus - inferior epiphysis. Anterior view. File:Slide10DEN.JPG, Trochlea. Posterior view. File:LeftHumanPosteriorDistalHumerusFlexed.jpg, Humerus - inferior epiphysis. Posterior view. File:AnteroMedialHeadofLeftHumerus.jpg, Humerus - superior epiphysis. Anterior view. File:Superior epiphysis.jpg, Humerus - superior epiphysis. Posterior view. File:Elbow joint - deep dissection (anterior view, human cadaver).jpg, Elbow joint. Deep dissection. Anterior view. File:Slide1bgbg.JPG, Elbow joint. Deep dissection. Posterior view. File:Slide3bgbg.JPG, Elbow joint. Deep dissection. Posterior view. File:Gray326.png, The left shoulder and acromioclavicular joints, and the proper ligaments of the scapula File:Gray207.png, Left humerus. Anterior view. File:Gray208.png, Left humerus. Posterior view. File:AnteriorHeadOfLeftHumerus.jpg, Left humerus. Anteriolateral view. File:MedialHeadOfLeftHumerus.jpg, Left humerus. Medial view. File:ProxHumeralFracture.png, Fracture of the proximal humerus File:Gray329.png, Left elbow-joint, showing anterior and ulnar collateral ligaments. File:Gray332.png, Capsule of elbow-joint (distended). Posterior aspect. File:Humerus Anatomy by Jason Christian.webm, Humerus Anatomy

Ossification

During embryonic development, the humerus is one of the first structures to ossify, beginning with the first ossification center in the shaft of the bone. Ossification of the humerus occurs predictably in the embryo and fetus, and is therefore used as a fetal biometric measurement when determining gestational age of a fetus. At birth, the neonatal humerus is only ossified in the shaft. The epiphyses are cartilaginous at birth. The medial humeral head develops an ossification center around 4 months of age and the greater tuberosity around 10 months of age. These ossification centers begin to fuse at 3 years of age. The process of ossification is complete by 13 years of age, though the epiphyseal plate (growth plate) persists until skeletal maturity, usually around 17 years of age.

References

External links

*
Humerus
- BlueLink Anatomy, University of Michigan Medical School {{Authority control Bones of the upper limb Long bones