Gammarus
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''Gammarus'' is an
amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far descri ...
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Gammaridae. It contains more than 200 described
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
, making it one of the most species-rich genera of crustaceans. Different species have different
optimal Mathematical optimization (alternatively spelled ''optimisation'') or mathematical programming is the selection of a best element, with regard to some criterion, from some set of available alternatives. It is generally divided into two subfi ...
conditions, particularly in terms of
salinity Salinity () is the saltiness or amount of salt (chemistry), salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensio ...
, and different tolerances; '' Gammarus pulex'', for instance, is a purely
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...
species, while '' Gammarus locusta'' is estuarine, only living where the salinity is greater than 25
Per mille (from Latin , "in each thousand") is an expression that means parts per thousand. Other recognised spellings include per mil, per mill, permil, permill, or permille. The associated sign is written , which looks like a percent ...
. Species of ''Gammarus'' are the typical " scuds" of North America and range widely throughout the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
. A considerable number are also found southwards into the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also refer ...
, particularly in
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
.


Species

The following species are included: Four new species were found in 2018 on the
Tibetan Plateau The Tibetan Plateau (, also known as the Qinghai–Tibet Plateau or the Qing–Zang Plateau () or as the Himalayan Plateau in India, is a vast elevated plateau located at the intersection of Central, South and East Asia covering most of the Ti ...
. Four more new species were described from the
Chihuahuan Desert The Chihuahuan Desert ( es, Desierto de Chihuahua, ) is a desert ecoregion designation covering parts of northern Mexico and the southwestern United States. It occupies much of far West Texas, the middle to lower Rio Grande Valley and the lo ...
in 2021. *''
Gammarus abscisus ''Gammarus'' is an amphipod crustacean genus in the family Gammaridae. It contains more than 200 described species, making it one of the most species-rich genera of crustaceans. Different species have different optimal conditions, particularly ...
'' G. Karaman, 1973 *''
Gammarus abstrusus ''Gammarus'' is an amphipod crustacean genus in the family Gammaridae. It contains more than 200 described species, making it one of the most species-rich genera of crustaceans. Different species have different optimal conditions, particularly ...
'' Hou, Platvoet & Li, 2006 *''
Gammarus acalceolatus ''Gammarus'' is an amphipod crustacean genus in the family Gammaridae. It contains more than 200 described species, making it one of the most species-rich genera of crustaceans. Different species have different optimal conditions, particularly ...
'' Pinkster, 1970 *'' Gammarus accolae'' G. Karaman, 1973 *'' Gammarus accretus'' Hou & Li, 2002 *''
Gammarus acherondytes ''Gammarus acherondytes'', the Illinois cave amphipod, is a species of crustacean in the family Gammaridae. The crustacean is Endemism, endemic to the Illinois Sinkhole Plain of Monroe County, Illinois, Monroe County and St. Clair County, Illinoi ...
'' Hubricht & Mackins, 1940 *'' Gammarus aequicauda'' (Martynov, 1931) *'' Gammarus agrarius'' G. Karaman, 1973 *'' Gammarus albimanus'' G. Karaman, 1968 *'' Gammarus alsaticus'' Van Straelen, 1924† *'' Gammarus altus'' Hou & Li, 2018 *'' Gammarus anatoliensis'' Schellenberg, 1937 *'' Gammarus angulatus'' (Martynov, 1930) *'' Gammarus angusticoxalis'' Martynov, 1935 *'' Gammarus annandalei'' (Monod, 1924) *'' Gammarus annulatus'' Smith, 1873 *'' Gammarus anodon'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus aoculus'' Hou & Li, 2003 *'' Gammarus araxenius'' Derzhavin, 1938 *'' Gammarus arduus'' G. Karaman, 1975 *'' Gammarus argaeus'' Vávra, 1905 *'' Gammarus bakhteyaricus'' Khalayi-Pirbalanty & Sari, 2004 *'' Gammarus balcanicus'' Schaferna, 1922 *'' Gammarus barnaulensis'' Schellenberg, 1937 *'' Gammarus baysali'' Özbek et al., 2013 *'' Gammarus belli'' G. Karaman, 1984 *'' Gammarus bergi'' Martynov, 1930 *'' Gammarus birsteini'' Karaman & Pinkster, 1977 *'' Gammarus bosniacus'' Schäferna, 1922 *'' Gammarus bousfieldi'' Cole & Minckley, 1961 *'' Gammarus brevicornis'' (Martynov, 1935) *'' Gammarus brevipodus'' Hou, Li & Platvoet, 2004 *'' Gammarus breviramus'' Bousfield & Elwood, 1971 *'' Gammarus bucharensis'' Martynov, 1935 *'' Gammarus caparti'' Petre-Stroobants, 1981 *'' Gammarus caucasicus'' Martynov, 1932 *'' Gammarus caudisetus'' Viviani, 1805 *'' Gammarus chaohuensis'' Hou & Li, 2002 *'' Gammarus chevreuxi'' Sexton, 1913 *'' Gammarus chimkenti'' G. Karaman, 1984 *'' Gammarus chostensis'' Martynov, 1932 *'' Gammarus cohabitus'' Hosinger, Shafer, Fong & Culver, 2008 *'' Gammarus colei'' sp. nov. *'' Gammarus comosus'' Hou, Li & Gao, 2005 *'' Gammarus craspedotrichus'' Hou & Li, 2002 *'' Gammarus crenulatus'' G. S. Karaman & Pinkster, 1977 *'' Gammarus crinicaudatus'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus crinicornis'' Stock, 1966 *'' Gammarus crispus'' Martynov, 1932 *'' Gammarus curvativus'' Hou & Li, 2003 *'' Gammarus dabanus'' Tachteew & Mekhanikova, 2000 *'' Gammarus daiberi'' Bousfield, 1969 *'' Gammarus decorosus'' Meng, Hou & Li, 2003 *'' Gammarus denticulatus'' Hou, Li & Morino, 2002 *'' Gammarus desperatus'' Cole, 1981 *'' Gammarus dorsosetosus'' Mateus & Mateus, 1990 *'' Gammarus duebeni'' Liljeborg, 1852 *'' Gammarus dulensis'' S. Karaman, 1929 *'' Gammarus edwardsi'' Bate, 1862 *'' Gammarus effultus'' G. Karaman, 1975 *'' Gammarus electrus'' Hou & Li, 2003 *'' Gammarus elevatus'' Hou, Li & Morino, 2002 *'' Gammarus elvirae'' Iannilli & Ruffo, 2002 *'' Gammarus emeiensis'' Hou, Li & Koenemann, 2002 *'' Gammarus fasciatus'' Say, 1818 *'' Gammarus finmarchicus'' Dahl, 1938 *'' Gammarus flabellifera'' Stimpson, 1856 *'' Gammarus fluviatilis'' Milne-Edwards, 1840† *'' Gammarus fontinalis'' Costa, 1883 *'' Gammarus fossarum'' Koch, 1836 *'' Gammarus frater'' Karaman & Pinkster, 1977 *'' Gammarus frigidus'' Hou & Li, 2004 *'' Gammarus galgosensis'' Lee & Kim, 2004 *'' Gammarus galgosensis'' Lee & Kim, 1980 *'' Gammarus gauthieri'' (S. Karaman, 1935) *'' Gammarus glabratus'' Hou & Li, 2003 *'' Gammarus goedmakersae'' G. S. Karaman & Pinkster, 1977 *'' Gammarus gonggaensis'' Hou & Li, 2018 *'' Gammarus gracilis'' Martynov, 1935 *'' Gammarus gregoryi'' Tattersall, 1924 *'' Gammarus halilicae'' G. Karaman, 1969 *'' Gammarus hegmatanensis'' Hekmatara, Sari & Baladehi, 2011 *'' Gammarus hirsutus'' Martynov, 1935 *'' Gammarus hongyuanensis'' Barnard & Dai, 1988 *'' Gammarus hoonsooi'' Lee, 1986 *'' Gammarus hyalelloides'' Cole, 1976 *'' Gammarus ibericus'' Margalef, 1951 *'' Gammarus inaequicauda'' Stock, 1966 *'' Gammarus inberbus'' Karaman & Pinkster, 1977 *'' Gammarus inopinatus'' Mateus & Mateus, 1990 *'' Gammarus insensibilis'' Stock, 1966 *'' Gammarus italicus'' Goedmakers & Pinkster, 1977 *'' Gammarus izmirensis'' Özbek, 2007 *'' Gammarus jacksoni'' Morino & Whitman, 1995 *'' Gammarus jaspidus'' Hou & Li, 2004 *'' Gammarus jenneri'' Bynum & Fox, 1977 *'' Gammarus kamtschaticus'' Tzvetkova, 1972 *'' Gammarus kangdingensis'' Hou & Li, 2018 *'' Gammarus kasymovi'' Aliev, 1997 *'' Gammarus katagani'' Özbek, 2012 *'' Gammarus kesanensis'' Özbek & Camur-Elipek, 2010 *'' Gammarus kesslerianus'' Martynov, 1931 *'' Gammarus kischineffensis'' Schellenberg, 1937 *'' Gammarus komareki'' (Schaferna, 1922) *'' Gammarus korbuensis'' Martynov, 1930 *'' Gammarus koshovi'' Bazikalova, 1946 *'' Gammarus kyonggiensis'' Lee & Seo, 1990 *'' Gammarus laborifer'' Karaman & Pinkster, 1977 *'' Gammarus lacustris'' G. O. Sars, 1864 *'' Gammarus langi'' sp. nov. *'' Gammarus lasaensis'' Barnard & Dai, 1988 *'' Gammarus laticoxalis'' Karaman & Pinkster, 1977 *'' Gammarus lawrencianus'' Bousfield, 1956 *'' Gammarus lecroyae'' Thoma & Heard, 2009 *'' Gammarus ledoyeri'' G. Karaman, 1987 *'' Gammarus lepoliensis'' Jazdzewski & Konopacka, 1989 *'' Gammarus lichuanensis'' Hou & Li, 2002 *'' Gammarus limnaeus'' S. I. smith, 1874 *'' Gammarus limosus'' Hou & Li, 2018 *'' Gammarus lobifer'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus locusta'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologi ...
)
*'' Gammarus longipedis'' Karaman & Pinkster, 1987 *'' Gammarus longisaeta'' Lee & Seo, 1992 *'' Gammarus lophacanthus'' Hou & Li, 2002 *'' Gammarus lordeganensis'' Khalayi-Pirbalanty & Sari, 2004 *'' Gammarus lychnidensis'' Schellenberg, 1943 *'' Gammarus macedonicus'' G. Karaman, 1976 *'' Gammarus madidus'' Hou & Li, 2005 *'' Gammarus malpaisensis'' sp. nov. *'' Gammarus marmouchensis'' Fadil, 2006 *'' Gammarus maroccanus'' Fadil & Dakki, 2001 *'' Gammarus martensi'' Hou & Li, 2004 *'' Gammarus matienus'' Derzhavin, 1938 *'' Gammarus miae'' Mateus & Mateus, 1990 *'' Gammarus microps'' Pinkster & Goedmakers, 1975 *'' Gammarus minus'' Say, 1818 *'' Gammarus mladeni'' Karaman & Pinkster, 1977 *'' Gammarus monospeliensis'' Pinkster, 1972 *'' Gammarus montanus'' Hou, Li & Platvoet, 2004 *''
Gammarus mucronatus ''Gammarus mucronatus'' is a species of scud in the family Gammaridae. It is found in the coasts of the North American Atlantic seaboard and the Gulf of Mexico. Phylogenetics ''Gammarus mucronatus'' is part of the Arthropod phylum: it has an ...
'' Say, 1818 *'' Gammarus murarius'' Hou & Li, 2004 *'' Gammarus nekkensis'' Uchida, 1935 *'' Gammarus ninglangensis'' Hou & Li, 2003 *'' Gammarus nipponensis'' Uéno, 1940 *'' Gammarus nox'' Stock, 1995 *'' Gammarus nudus'' Martynov, 1931 *'' Gammarus obnixus'' Karaman & Pinkster, 1977 *'' Gammarus obruki'' Özbek, 2012 *'' Gammarus oceanicus'' Segerstråle, 1947 *'' Gammarus ocellatus'' Martynov, 1930 *'' Gammarus ochridensis'' (Schäferna, 1926) *'' Gammarus odaensis'' Lee & Kim, 1980 *'' Gammarus odettae'' Mateus & Mateus, 1990 *'' Gammarus oeningensis'' Heer, 1865† *'' Gammarus orinos'' Pinkster & Schol, 1984 *'' Gammarus oronticus'' Alouf, 1979 *'' Gammarus osellai'' Karaman & Pinkster, 1977 *'' Gammarus pageti'' Mateus & Mateus, 1990 *'' Gammarus palustris'' Bousfield, 1969 *'' Gammarus parechiniformis'' G. Karaman, 1977 *'' Gammarus paricrenatus'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus parthicus'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus paucispinus'' Hou & Li, 2002 *'' Gammarus pavlovici'' *'' Gammarus pavo'' Karaman & Pinkster, 1977 *'' Gammarus paynei'' Delong, 1992 *'' Gammarus pecos'' Cole & Bousfield, 1970 *'' Gammarus pellucidus'' Gurjanova, 1930 *'' Gammarus percalacustris'' sp. nov. *'' Gammarus pexus'' Hou & Li, 2005 *'' Gammarus platvoeti'' Hou & Li, 2003 *'' Gammarus pljakici'' G. Karaman, 1964 *'' Gammarus plumipes'' Mateus & Mateus, 1990 *'' Gammarus praecyrius'' Derzhavin, 1941† *'' Gammarus preciosus'' Wang, Hou & Li, 2009 *'' Gammarus pretzmanni'' Mateus & Mateus, 1990 *''
Gammarus proiectus ''Gammarus'' is an amphipod crustacean genus in the family Gammaridae. It contains more than 200 described species, making it one of the most species-rich genera of crustaceans. Different species have different optimal conditions, particularly i ...
'' Stock, Mirzajani, Vonk, Naderi & Kiabi, 1998 *'' Gammarus pseudoanatoliensis'' Karaman & Pinkster, 1987 *'' Gammarus pseudolimnaeus'' Bousfield, 1958 *'' Gammarus pseudosyriacus'' Karaman & Pinkster, 1977 *'' Gammarus pulex'' Linnaeus, 1758 *'' Gammarus qiani'' Hou & Li, 2002 *'' Gammarus rambouseki'' S. Karaman, 1931 *'' Gammarus retzi'' Maikovsky, 1941† *'' Gammarus rifatlensis'' Fadil, 2006 *'' Gammarus riparius'' Hou & Li, 2002 *'' Gammarus roeselii'' Gervais, 1835 *'' Gammarus rouxi'' Pinkster & Goedmakers, 1975 *'' Gammarus salemaai'' G. Karaman, 1985 *'' Gammarus salinus'' Spooner, 1947 *'' Gammarus sepidannus'' Zamanpoore, Poeckl, Grabowski & Schiemer, 2009 *''
Gammarus setosus ''Gammarus setosus'' is a gammarid amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There ar ...
'' Dementieva, 1931 *'' Gammarus shanxiensis'' Barnard & Dai, 1988 *'' Gammarus shenmuensis'' Hou & Li, 2004 *'' Gammarus sichuanensis'' Hou, Li & Zheng, 2002 *'' Gammarus sinuolatus'' Hou & Li, 2004 *'' Gammarus sirvannus'' Hekmatara, Sari & Baladehi, 2011 *'' Gammarus sketi'' G. Karaman, 1989 *'' Gammarus sobaegensis'' G. Karaman, 1984 *'' Gammarus solidus'' G. Karaman, 1977 *'' Gammarus songirdaki'' G. Karaman, 1984 *'' Gammarus soyoensis'' Lee & Kim, 1980 *'' Gammarus spelaeus'' Martynov, 1931 *'' Gammarus spinipalmus'' (Chen, 1939) *'' Gammarus spooneri'' G. Karaman, 1991 *'' Gammarus stagnarius'' Hou, Li & Morino, 2002 *'' Gammarus stalagmiticus'' Hou & Li, 2005 *'' Gammarus stankokaramani'' G. Karaman, 1976 *'' Gammarus stojicevici'' (S. Karaman, 1929) *'' Gammarus stupendus'' Pinkster, 1983 *'' Gammarus subtypicus'' Stock, 1966 *'' Gammarus suifunensis'' Martynov, 1925 *'' Gammarus syriacus'' Chevreux, 1895 *'' Gammarus takesensis'' Hou, Li & Platvoet, 2004 *'' Gammarus taliensis'' Shen, 1954 *'' Gammarus tauricus'' Martynov, 1931 *'' Gammarus teletzkensis'' Martynov, 1930 *'' Gammarus tigrinus'' Sexton, 1939 *'' Gammarus topkarai'' Özbek & Balik, 2009 *'' Gammarus translucidus'' Hou, Li & Li, 2004 *'' Gammarus troglophilus'' Hubricht & Mackins, 1940 *'' Gammarus truncatus'' Martynov, 1930 *'' Gammarus turanus'' Martynov, 1935 *'' Gammarus uludagi'' G. S. Karaman, 1975 *'' Gammarus unguiserratus'' Costa, 1853 *'' Gammarus ustaoglui'' Özbek & Guloglu, 2005 *'' Gammarus varsoviensis'' Jazdzewski, 1975 *'' Gammarus vignai'' Pinkster & Karaman, 1978 *'' Gammarus wangbangensis'' Lee & Seo, 1990 *'' Gammarus wautieri'' Roux, 1967 *'' Gammarus wilkitzkii'' Birula, 1897 *'' Gammarus xiangfengensis'' Hou & Li, 2002 *'' Gammarus zaddachi'' Sexton, 1912 *'' Gammarus zagrosensis'' Zamanpoore, Poeckl, Grabowski & Schiemer, 2009 *'' Gammarus zeongogensis'' Lee & Kim, 1980


References

{{Taxonbar, from=Q548451 Gammaridea Taxonomy articles created by Polbot Crustacean genera