Frans de Waal
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Franciscus Bernardus Maria "Frans" de Waal (born October 29, 1948) is a Dutch primatologist and
ethologist Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objectiv ...
. He is the Charles Howard Candler Professor of
Primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
Behavior Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as we ...
in the Department of
Psychology Psychology is the science, scientific study of mind and behavior. Psychology includes the study of consciousness, conscious and Unconscious mind, unconscious phenomena, including feelings and thoughts. It is an academic discipline of immens ...
at
Emory University Emory University is a private research university in Atlanta, Georgia. Founded in 1836 as "Emory College" by the Methodist Episcopal Church and named in honor of Methodist bishop John Emory, Emory is the second-oldest private institution of ...
in
Atlanta, Georgia Atlanta ( ) is the capital city, capital and List of municipalities in Georgia (U.S. state), most populous city of the U.S. state of Georgia (U.S. state), Georgia. It is the county seat, seat of Fulton County, Georgia, Fulton County, the mos ...
, director of the Living Links Center at the
Yerkes National Primate Research Center The Emory National Primate Research Center (formerly known as Yerkes National Primate Research Center) located in Atlanta, Georgia, owned by Emory University, is a center of biomedical and behavioral research, is dedicated to improving human and a ...
at Emory, and author of numerous books including ''Chimpanzee Politics'' (1982) and ''Our Inner Ape'' (2005). His research centers on primate social behavior, including
conflict resolution Conflict resolution is conceptualized as the methods and processes involved in facilitating the peaceful ending of conflict and retribution. Committed group members attempt to resolve group conflicts by actively communicating information abo ...
,
cooperation Cooperation (written as co-operation in British English) is the process of groups of organisms working or acting together for common, mutual, or some underlying benefit, as opposed to working in competition for selfish benefit. Many animal a ...
,
inequity aversion Inequity aversion (IA) is the preference for fairness and resistance to incidental inequalities. The social sciences that study inequity aversion include sociology, economics, psychology, anthropology, and ethology. Human studies Inequity aversion ...
, and food-sharing. He is a member of the
United States National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the N ...
and the
Royal Netherlands Academy of Arts and Sciences The Royal Netherlands Academy of Arts and Sciences ( nl, Koninklijke Nederlandse Akademie van Wetenschappen, abbreviated: KNAW) is an organization dedicated to the advancement of science and literature in the Netherlands. The academy is housed ...
.


Early life and education

De Waal was born in
's-Hertogenbosch s-Hertogenbosch (), colloquially known as Den Bosch (), is a city and municipality in the Netherlands with a population of 157,486. It is the capital of the province of North Brabant and its fourth largest by population. The city is south of ...
on October 29, 1948. He studied at the Dutch universities of
Radboud University Nijmegen Radboud University (abbreviated as RU, nl, Radboud Universiteit , formerly ''Katholieke Universiteit Nijmegen'') is a public research university located in Nijmegen, the Netherlands. The university bears the name of Saint Radboud, a 9th century ...
,
University of Groningen The University of Groningen (abbreviated as UG; nl, Rijksuniversiteit Groningen, abbreviated as RUG) is a public research university of more than 30,000 students in the city of Groningen in the Netherlands. Founded in 1614, the university is th ...
, and
Utrecht Utrecht ( , , ) is the fourth-largest city and a municipality of the Netherlands, capital and most populous city of the province of Utrecht. It is located in the eastern corner of the Randstad conurbation, in the very centre of mainland Net ...
. In 1977, De Waal received his doctorate in
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary ...
from
Utrecht University Utrecht University (UU; nl, Universiteit Utrecht, formerly ''Rijksuniversiteit Utrecht'') is a public research university in Utrecht, Netherlands. Established , it is one of the oldest universities in the Netherlands. In 2018, it had an enrollme ...
after training as a
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...
and
ethologist Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objectiv ...
with professor Jan van Hooff, a well-known expert of emotional facial expressions in primates. His dissertation titled: "Agonistic interactions and relations among Java-monkeys" concerned aggressive behavior and
alliance An alliance is a relationship among people, groups, or states that have joined together for mutual benefit or to achieve some common purpose, whether or not explicit agreement has been worked out among them. Members of an alliance are called ...
formation in
macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principall ...
s. Fellow Dutch ethologist Niko Tinbergen was an inspiration to de Waal.


Career

In 1975, De Waal began a six-year project on the world's largest captive colony of
chimpanzee The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative t ...
s at the Arnhem Zoo. The study resulted in many scientific papers, and resulted in publication of his first book, ''Chimpanzee Politics'', in 1982. This book offered the first description of primate behavior explicitly in terms of planned social strategies. De Waal was first to introduce the thinking of Machiavelli to primatology, leading to the label "Machiavellian Intelligence" that later became associated with it. In his writings, De Waal has never shied away from attributing emotions and
intention Intentions are mental states in which the agent commits themselves to a course of action. Having the plan to visit the zoo tomorrow is an example of an intention. The action plan is the ''content'' of the intention while the commitment is the ''a ...
s to his primates, and as such his work inspired the field of
primate cognition Primate cognition is the study of the intellectual and behavioral skills of non-human primates, particularly in the fields of psychology, behavioral biology, primatology, and anthropology. Primates are capable of high levels of cognition; some m ...
that, three decades later, flourishes around themes of cooperation,
altruism Altruism is the principle and moral practice of concern for the welfare and/or happiness of other human beings or animals, resulting in a quality of life both material and spiritual. It is a traditional virtue in many cultures and a cor ...
, and fairness. His early work also drew attention to
deception Deception or falsehood is an act or statement that misleads, hides the truth, or promotes a belief, concept, or idea that is not true. It is often done for personal gain or advantage. Deception can involve dissimulation, propaganda and sleight o ...
and conflict resolution, nowadays two major areas of research. Initially, all of this was highly controversial. Thus, the label of "reconciliation", which De Waal introduced for reunions after fights, was questioned at first, but is now fully accepted with respect to animal behavior. Recently, De Waal's work has emphasized non-human animal empathy and even the origins of morality. His most widely cited paper, written with his former student Stephanie Preston, concerns the evolutionary origin and
neuroscience Neuroscience is the science, scientific study of the nervous system (the brain, spinal cord, and peripheral nervous system), its functions and disorders. It is a Multidisciplinary approach, multidisciplinary science that combines physiology, an ...
of empathy, not just in primates, but in mammals in general. De Waal's name is also associated with
bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee and less often the dwarf chimpanzee or gracile chimpanzee, is an endangered great ape and one of the two species making up the genus '' Pan,'' the other being the comm ...
s, the "make love – not war" primates that he has made popular. But even his bonobo studies are secondary to the larger goal of understanding what binds primate societies together rather than how competition structures them. Competition is not ignored in his work: the original focus of de Waal's research, before he was well known, was aggressive behavior and social dominance. Whereas his science focuses on the behavior of nonhuman primates (mostly chimpanzees, bonobos,
macaques The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and (in one instance) Gibraltar. Macaques are principal ...
, and
capuchin monkeys The capuchin monkeys () are New World monkeys of the subfamily Cebinae. They are readily identified as the " organ grinder" monkey, and have been used in many movies and television shows. The range of capuchin monkeys includes some tropical fore ...
), his popular books have given de Waal worldwide visibility by relating the insights he has gained from monkey and ape behavior to human society. With his students, he has also worked on elephants, which are increasingly featured in his writings. His research into the innate capacity for
empathy Empathy is the capacity to understand or feel what another person is experiencing from within their frame of reference, that is, the capacity to place oneself in another's position. Definitions of empathy encompass a broad range of social, co ...
among primates has led De Waal to the conclusion that non-human
great ape The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ...
s and humans are simply different types of apes, and that empathic and cooperative tendencies are continuous between these species. His belief is illustrated in the following quote from ''The Age of Empathy'': "We start out postulating sharp boundaries, such as between humans and apes, or between apes and monkeys, but are in fact dealing with sand castles that lose much of their structure when the sea of knowledge washes over them. They turn into hills, leveled ever more, until we are back to where evolutionary theory always leads us: a gently sloping beach." This is quite opposite to the view of some economists and anthropologists, who postulate the differences between humans and other animals. However, recent work on prosocial tendencies in apes and monkeys supports de Waal's position. See, for example, the research of Felix Warneken, a psychologist at the
Max Planck Institute for Evolutionary Anthropology The Max Planck Institute for Evolutionary Anthropology (german: Max-Planck-Institut für evolutionäre Anthropologie, shortened to MPI EVA) is a research institute based in Leipzig, Germany, that was founded in 1997. It is part of the Max Plan ...
in
Leipzig Leipzig ( , ; Upper Saxon: ) is the most populous city in the German state of Saxony. Leipzig's population of 605,407 inhabitants (1.1 million in the larger urban zone) as of 2021 places the city as Germany's eighth most populous, as ...
, Germany. In 2011, de Waal and his co-workers were the first to report that chimpanzees given a free choice between helping only themselves or helping themselves plus a partner, prefer the latter. In fact, de Waal does not believe these tendencies to be restricted to humans and apes, but views empathy and sympathy as universal mammalian characteristics, a view that over the past decade has gained support from studies on rodents and other mammals, such as dogs. He and his students have extensively worked on such
cooperation Cooperation (written as co-operation in British English) is the process of groups of organisms working or acting together for common, mutual, or some underlying benefit, as opposed to working in competition for selfish benefit. Many animal a ...
and fairness in animals. In 2011 de Waal gave a
TED Talk TED Conferences, LLC (Technology, Entertainment, Design) is an American-Canadian non-profit media organization that posts international talks online for free distribution under the slogan "ideas worth spreading". TED was founded by Richard Sau ...
entitled "Moral behavior in animals". Part of the talk dealt with inequity aversion among capuchin monkeys, and a video extract of this went viral. It showed the furious reaction of one monkey given a less desirable treat than another. The most recent work in this area was the first demonstration that given a chance to play the
Ultimatum game The ultimatum game is a game that has become a popular instrument of economic experiments. An early description is by Nobel laureate John Harsanyi in 1961. One player, the proposer, is endowed with a sum of money. The proposer is tasked with s ...
, chimpanzees respond in the same way as children and human adults by preferring the equitable outcome. In 1981, de Waal moved to the United States for a position at the Wisconsin National Primate Research Center, and in 1991 took a position at Emory University, in Atlanta, Georgia. He is currently C.H. Candler Professor in the Psychology Department at
Emory University Emory University is a private research university in Atlanta, Georgia. Founded in 1836 as "Emory College" by the Methodist Episcopal Church and named in honor of Methodist bishop John Emory, Emory is the second-oldest private institution of ...
and director of the Living Links Center at the
Yerkes National Primate Research Center The Emory National Primate Research Center (formerly known as Yerkes National Primate Research Center) located in Atlanta, Georgia, owned by Emory University, is a center of biomedical and behavioral research, is dedicated to improving human and a ...
at Emory. He became an American citizen in 2008. His 2013 book ''The Bonobo and the Atheist'' examines human behavior through the eyes of a primatologist, and explores to what extent God and religion are needed for human morality. The main conclusion is that morality comes from within, and is part of human nature. The role of religion is secondary. De Waal also writes a column for ''Psychologie Magazine'', a popular Dutch monthly. Since September 1, 2013, de Waal has been a distinguished professor (''universiteitshoogleraar'') at Utrecht University. This is a part-time appointment—he remains in his position at Emory University, in Atlanta. In October 2016, de Waal was the guest on the BBC Radio Four program ''
The Life Scientific ''The Life Scientific'' is a BBC Radio 4 science programme, presented by Professor Jim Al-Khalili , in which each episode is dedicated to the biography and work of one living scientist. The programme consists of an interview between Al-Khalili ...
''. In June 2018, de Waal was awarded the ''NAT Award'', recently established by the Museum of Natural Sciences of Barcelona. The award, which goes to people or institutions "that are referents for their way of viewing and explaining nature, whether because they have encouraged professional engagement in natural history disciplines or because they have contributed significantly to nature conservation", was awarded to de Waal "for his vision regarding the evolution of animal behaviour in establishing a parallel between primate and human behaviour in aspects such as politics, empathy, morality and justice." Alongside de Waal, broadcaster and naturalist
David Attenborough Sir David Frederick Attenborough (; born 8 May 1926) is an English broadcaster, biologist, natural historian and author. He is best known for writing and presenting, in conjunction with the BBC Natural History Unit, the nine natural histo ...
was awarded an ''Extraordinary Award for a Professional Career'', and biologist and former director of the Barcelona Zoology Museum Roser Nos Ronchera was awarded a ''Honorable Mention''.


Awards

* 2021 Doctor ''Honoris Causa'', Université Jean-Monnet-Saint-Étienne (France) * 2020 PEN / EO Wilson Literary Science Writing Award (US) * 2018 Doctor ''Honoris Causa'',
Yale University Yale University is a private research university in New Haven, Connecticut. Established in 1701 as the Collegiate School, it is the third-oldest institution of higher education in the United States and among the most prestigious in the w ...
(US) * 2018 NAT Award for the Dissemination of Natural Science, Barcelona (Spain) * 2017 Doctor ''Honoris Causa'', Radboud University (Netherlands) * 2015 ASP Distinguished Primatologist (American Society of Primatologists) * 2014 Galileo Prize (Premio Letterario Galileo), Padua (Italy) * 2014 Eugène Dubois Chair, Maastricht University (Netherlands) * 2013 Edward O. Wilson Biodiversity Technology Pioneer Award * 2013 Foreign Member, Royal Holland Society of Sciences and Humanities * 2013 Doctor ''
Honoris Causa An honorary degree is an academic degree for which a university (or other degree-awarding institution) has waived all of the usual requirements. It is also known by the Latin phrases ''honoris causa'' ("for the sake of the honour") or ''ad hono ...
'',
Utrecht University Utrecht University (UU; nl, Universiteit Utrecht, formerly ''Rijksuniversiteit Utrecht'') is a public research university in Utrecht, Netherlands. Established , it is one of the oldest universities in the Netherlands. In 2018, it had an enrollme ...
(Netherlands) * 2012
Ig Nobel Prize The Ig Nobel Prize ( ) is a satiric prize awarded annually since 1991 to celebrate ten unusual or trivial achievements in scientific research. Its aim is to "honor achievements that first make people laugh, and then make them think." The name o ...
winner, in the Anatomy category * 2011 ''
Discover Discover may refer to: Art, entertainment, and media * ''Discover'' (album), a Cactus Jack album * ''Discover'' (magazine), an American science magazine Businesses and brands * DISCover, the ''Digital Interactive Systems Corporation'' * D ...
'' magazine's "47 (all time) Great Minds of Science" * 2011 Doctor ''Honoris Causa'',
Colgate University Colgate University is a private liberal arts college in Hamilton, New York. The college was founded in 1819 as the Baptist Education Society of the State of New York and operated under that name until 1823, when it was renamed Hamilton Theolog ...
(US) * 2010 Knight of the
Order of the Netherlands Lion The Order of the Netherlands Lion, also known as the Order of the Lion of the Netherlands ( nl, De Orde van de Nederlandse Leeuw, french: L'Ordre du Lion Néerlandais) is a Dutch order of chivalry founded by King William I of the Netherlands on ...
* 2009 Medal, Società di Medicina & Scienze Naturali, Parma (Italy) * 2009 Ariëns Kappers Medal,
Royal Netherlands Academy of Arts and Sciences The Royal Netherlands Academy of Arts and Sciences ( nl, Koninklijke Nederlandse Akademie van Wetenschappen, abbreviated: KNAW) is an organization dedicated to the advancement of science and literature in the Netherlands. The academy is housed ...
* 2009 Doctor ''Honoris Causa'', University for Humanistics (Netherlands) * 2008 Member of the
American Academy of Arts and Sciences The American Academy of Arts and Sciences (abbreviation: AAA&S) is one of the oldest learned societies in the United States. It was founded in 1780 during the American Revolution by John Adams, John Hancock, James Bowdoin, Andrew Oliver, a ...
(AAAS) * 2007 ''Time'' 100 world's most influential people * 2005 Member of the
American Philosophical Society The American Philosophical Society (APS), founded in 1743 in Philadelphia, is a scholarly organization that promotes knowledge in the sciences and humanities through research, professional meetings, publications, library resources, and communit ...
(APS) * 2005 Arthur W. Staats Award,
American Psychological Foundation The American Psychological Foundation (abbreviated APF) is an American philanthropic organization dedicating to awarding research grants to psychologists in the early stages of their careers. It is affiliated with the American Psychological Assoc ...
* 2004 Member of the (US)
National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nat ...
(NAS) * 1993 Corresponding member of the
Royal Netherlands Academy of Arts and Sciences The Royal Netherlands Academy of Arts and Sciences ( nl, Koninklijke Nederlandse Akademie van Wetenschappen, abbreviated: KNAW) is an organization dedicated to the advancement of science and literature in the Netherlands. The academy is housed ...
(KNAW) * 1989 ''Los Angeles Times'' Book Award for ''Peacemaking among Primates''


Selected bibliography


Books

* ''Different: Gender Through the Eyes of a Primatologist'', 2022. * ''Mama's Last Hug: Animal Emotions and What They Tell Us about Ourselves'', 2019. * ''Are We Smart Enough to Know How Smart Animals Are?'', 2016. * ''The Bonobo and the Atheist'', 2013. * ''The Age of Empathy: Nature's Lessons for a Kinder Society'', 2009. * ''Primates and Philosophers: How Morality Evolved'', 2006. * ''Our Inner Ape''. New York: Riverhead Books, 2005. * ''Animal Social Complexity: Intelligence, Culture, and Individualized Societies'', Edited with Peter L. Tyack. Cambridge:
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retir ...
, 2003. . * ''My Family Album, Thirty Years of Primate Photography'' 2003. * ''Tree of Origin: What Primate Behavior Can Tell Us about Human Social Evolution'',
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retir ...
, 2001. . * '' The Ape and the Sushi Master, Cultural reflections by a primatologist''. New York:
Basic Books Basic Books is a book publisher founded in 1950 and located in New York, now an imprint of Hachette Book Group. It publishes books in the fields of psychology, philosophy, economics, science, politics, sociology, current affairs, and history. H ...
, 2001. * ''Chimpanzee Politics: Power and Sex Among Apes'' (25th Anniversary ed.). Baltimore, MD: JHU Press; 2007. . * ''Natural Conflict Resolution''. 2000 (with Filippo Aureli) * ''Bonobo: The Forgotten Ape''. Berkeley: University of California Press, 1997. (with
Frans Lanting Frans Lanting (born 13 July 1951) is a Dutch National Geographic photographer, author and speaker. Life Lanting was born in Rotterdam in Netherlands. He studied economics at the Erasmus university in Rotterdam and later immigrated to the United St ...
) * ''Good Natured: The Origins of Right and Wrong in Humans and Other Animals''. Cambridge:
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retir ...
, 1996. * ''Chimpanzee Cultures'', Edited with Richard Wrangham, W.C. McGrew, and Paul Heltne. Foreword by
Jane Goodall Dame Jane Morris Goodall (; born Valerie Jane Morris-Goodall on 3 April 1934), formerly Baroness Jane van Lawick-Goodall, is an English primatologist and anthropologist. Seen as the world's foremost expert on chimpanzees, Goodall is best kn ...
. Cambridge:
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retir ...
, 1994. . * ''Peacemaking Among Primates''. Cambridge:
Harvard University Press Harvard University Press (HUP) is a publishing house established on January 13, 1913, as a division of Harvard University, and focused on academic publishing. It is a member of the Association of American University Presses. After the retir ...
, 1989.


Articles

* 201
Opinion piece about the discovery of ''Homo naledi'' in ''The New York Times''
* 201
Opinion piece about animal intelligence in ''The Wall Street Journal''
* 201
Opinion piece about God and morality in ''The New York Times''
* 2010 * 2009, * 2008 * 2007

''
Skeptic Skepticism, also spelled scepticism, is a questioning attitude or doubt toward knowledge claims that are seen as mere belief or dogma. For example, if a person is skeptical about claims made by their government about an ongoing war then the ...
'', (August 8, 2007). * 2006, * 2005
"The empathic ape"
''New Scientist'', October 8, 2005 * 2001, "Do Humans Alone 'Feel Your Pain'?"

October 26, 2001) * 1999, * 1995,


Additional reading

*


See also

*''
The Genius of Charles Darwin ''The Genius of Charles Darwin'' is a three-part television documentary, written and presented by evolutionary biologist Richard Dawkins. It was first shown in August 2008 on Channel 4. It won Best TV Documentary Series 2008 at the British ...
'' (Richard Dawkins interviews De Waal) *'' The Family of Chimps'', a Dutch documentary film based on de Waal's book, ''Chimpanzee Politics and Our Inner Ape''


References


External links


''The surprising science of alpha males''
TED talk TED Conferences, LLC (Technology, Entertainment, Design) is an American-Canadian non-profit media organization that posts international talks online for free distribution under the slogan "ideas worth spreading". TED was founded by Richard Sau ...
by Frans de Waal.
''Do animals have morals?''
TED talk by Frans de Waal.
Interview with Frans de Waal
on the
BBC Radio 4 BBC Radio 4 is a British national radio station owned and operated by the BBC that replaced the BBC Home Service in 1967. It broadcasts a wide variety of spoken-word programmes, including news, drama, comedy, science and history from the BBC's ...
programme
The Life Scientific ''The Life Scientific'' is a BBC Radio 4 science programme, presented by Professor Jim Al-Khalili , in which each episode is dedicated to the biography and work of one living scientist. The programme consists of an interview between Al-Khalili ...
. * .
Frans de Waal
An extended film interview with transcripts for the 'Why Are We Here?' documentary series. {{DEFAULTSORT:Waal, Frans de 1948 births Living people List of atheists (surnames C to D) Animal cognition writers Dutch animal welfare scholars Dutch biologists Dutch expatriates in the United States Emory University faculty Ethologists Fellows of the American Academy of Arts and Sciences Human evolution theorists Knights of the Order of the Netherlands Lion Members of the American Philosophical Society Members of the Royal Netherlands Academy of Arts and Sciences Members of the United States National Academy of Sciences People from 's-Hertogenbosch Primatologists Utrecht University alumni