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The Exocoetidae are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of marine fish in the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
Beloniformes
class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differently ...
Actinopterygii Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
, known colloquially as flying fish or flying cod. About 64
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
are grouped in seven to nine
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
. While they cannot fly in the same way a bird does, flying fish can make powerful, self-propelled leaps out of the water where their long wing-like fins enable
gliding Gliding is a recreational activity and competitive air sport in which pilots fly unpowered aircraft known as gliders or sailplanes using naturally occurring currents of rising air in the atmosphere to remain airborne. The word ''soaring'' is a ...
for considerable distances above the water's surface. The main reason for this behavior is thought to be to escape from underwater predators, which include
swordfish Swordfish (''Xiphias gladius''), also known as broadbills in some countries, are large, highly migratory predatory fish characterized by a long, flat, pointed bill. They are a popular sport fish of the billfish category, though elusive. Swordf ...
,
mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...
,
tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...
, and
marlin Marlins are fish from the family Istiophoridae, which includes about 10 species. A marlin has an elongated body, a spear-like snout or bill, and a long, rigid dorsal fin which extends forward to form a crest. Its common name is thought to deri ...
, among others, though their periods of flight expose them to attack by avian predators such as frigate birds.
Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estima ...
is known as "the land of the flying fish", and the fish is one of the national symbols of the country. The Exocet missile is named after them, as variants are launched from underwater, and take a low trajectory, skimming the surface, before striking their targets.


Etymology

The term Exocoetidae is both the scientific name and the general name in
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
for a flying fish. The suffix ''-idae'', common for indicating a family, follows the root of the Latin word ', a transliteration of the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic pe ...
name . This means literally "sleeping outside", from "outside" and "bed", "resting place", verb root κει- "to lie down" (not "untruth"), so named as flying fish were believed to leave the water to sleep ashore, or due to flying fish flying and thus stranding themselves in boats.


Taxonomy

The Exocoetidae is divided into four subfamilies and seven genera: * Subfamily Exocoetinae Risso, 1827 ** Genus '' Exocoetus''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologi ...
* Subfamily Fodiatorinae Fowler, 1925 ** Genus '' Fodiator'' D.S. Jordan &
Meek Meekness is an attribute of human nature and behavior that has been defined as an amalgam of righteousness, inner humility, and patience. Meekness has been contrasted with humility alone insomuch as humility simply refers to an attitude towards o ...
, 1885
* Subfamily Parexocoetinae
Bruun Bruun is a surname of North Germanic origin. The meaning is ''brown'' (''brun'' in Danish, Swedish and Norwegian). In Denmark, the name is known to have been in use since the 13th century in the form Bruun. Other spelling variants are ''Bruhn'' an ...
, 1935
** Genus '' Parexocoetus'' Bleeker, 1865 * Subfamily Cypsellurinae Hubbs, 1933 ** Genus '' Cheilopogon'' Lowe, 1841 ** Genus '' Cypselurus'' Swainson, 1838 ** Genus '' Hirundichthys'' Breder, 1928 ** Genus '' Prognichthys'' Breder, 1928


Distribution and description

Flying fish live in all of the oceans, particularly in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and warm
subtropical The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north a ...
waters. They are commonly found in the epipelagic zone, the top layer of the ocean to a depth of about 200 m (656 ft). Numerous morphological features give flying fish the ability to leap above the surface of the ocean. One such feature is fully broadened
neural arches The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...
, which act as insertion sites for
connective tissue Connective tissue is one of the four primary types of animal tissue, along with epithelial tissue, muscle tissue, and nervous tissue. It develops from the mesenchyme derived from the mesoderm the middle embryonic germ layer. Connective tiss ...
s and ligaments in a fish's skeleton. Fully broadened neural arches act as more stable and sturdier sites for these connections, creating a strong link between the
vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...
and
cranium The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
. This ultimately allows a rigid and sturdy vertebral column (body) that is beneficial in flight. Having a rigid body during glided flight gives the flying fish
aerodynamic Aerodynamics, from grc, ἀήρ ''aero'' (air) + grc, δυναμική (dynamics), is the study of the motion of air, particularly when affected by a solid object, such as an airplane wing. It involves topics covered in the field of fluid dyn ...
advantages, increasing its speed and improving its aim. Furthermore, flying fish have developed vertebral columns and ossified
caudal Caudal may refer to: Anatomy * Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism * Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into the ...
complexes. These features provide the majority of strength to the flying fish, allowing them to physically lift their bodies out of water and glide remarkable distances. These additions also reduce the flexibility of the flying fish, allowing them to perform powerful leaps without weakening midair. At the end of a glide, they fold their
pectoral fins Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
to re-enter the sea, or drop their tails into the water to push against the water to lift for another glide, possibly changing direction. The curved profile of the "wing" is comparable to the aerodynamic shape of a bird wing. The fish is able to increase its time in the air by flying straight into or at an angle to the direction of
updraft In meteorology, an updraft is a small-scale current of rising air, often within a cloud. Overview Localized regions of warm or cool air will exhibit vertical movement. A mass of warm air will typically be less dense than the surrounding regi ...
s created by a combination of air and
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s. Species of genus '' Exocoetus'' have one pair of fins and streamlined bodies to optimize for speed, while '' Cypselurus'' spp. have flattened bodies and two pairs of fins, which maximize their time in the air. From 1900 to the 1930s, flying fish were studied as possible models used to develop airplanes. The Exocoetidae feed mainly on
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
. Predators include
dolphin A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the b ...
s,
tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae (mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max length: ...
,
marlin Marlins are fish from the family Istiophoridae, which includes about 10 species. A marlin has an elongated body, a spear-like snout or bill, and a long, rigid dorsal fin which extends forward to form a crest. Its common name is thought to deri ...
,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s,
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fittin ...
, and
porpoise Porpoises are a group of fully aquatic marine mammals, all of which are classified under the family Phocoenidae, parvorder Odontoceti (toothed whales). Although similar in appearance to dolphins, they are more closely related to narwhals a ...
s.


Flight measurements

In May 2008, a Japanese television crew (
NHK , also known as NHK, is a Japanese public broadcaster. NHK, which has always been known by this romanized initialism in Japanese, is a statutory corporation funded by viewers' payments of a television license fee. NHK operates two terrestr ...
) filmed a flying fish (dubbed "Icarfish") off the coast of Yakushima Island, Japan. The fish spent 45 seconds in flight. The previous record was 42 seconds. The flights of flying fish are typically around 50 m (160 ft),
Ross Piper Ross Piper is a British zoologist, entomologist, and explorer. Biography Piper's fascination by animals began at a young age through early encounters with a violet ground beetle and the caterpillar of an elephant hawk moth. This early interes ...
(2007), ''Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals'',
Greenwood Press Greenwood Publishing Group, Inc. (GPG), also known as ABC-Clio/Greenwood (stylized ABC-CLIO/Greenwood), is an educational and academic publisher (middle school through university level) which is today part of ABC-Clio. Established in 1967 as Gr ...
.
though they can use updrafts at the leading edge of waves to cover distances up to . They can travel at speeds of more than . Maximum altitude is above the surface of the sea. Flying fish often accidentally land on the decks of smaller vessels.


Fishery and cuisine

Flying fish are commercially fished in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
, and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
by
gillnetting Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...
, and in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Gui ...
and
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
by dipnetting. Often in
Japanese cuisine Japanese cuisine encompasses the regional and traditional foods of Japan, which have developed through centuries of political, economic, and social changes. The traditional cuisine of Japan ( Japanese: ) is based on rice with miso soup and oth ...
, the fish is preserved by drying to be used as fish stock for
dashi is a family of stocks used in Japanese cuisine. ''Dashi'' forms the base for miso soup, clear broth soup, noodle broth soup, and many simmering liquids to accentuate the savory flavor known as umami. ''Dashi'' is also mixed into the flour b ...
broth. The roe of '' Cheilopogon agoo'', or Japanese flying fish, is used to make some types of
sushi is a Japanese dish of prepared , usually with some sugar and salt, accompanied by a variety of , such as seafood, often raw, and vegetables. Styles of sushi and its presentation vary widely, but the one key ingredient is "sushi rice," also ...
, and is known as ''
tobiko is flying fish roe in Japanese cuisine, known for its use in ''sushi''. The eggs are small, ranging from 0.5 to 0.8 mm. For comparison, ''tobiko'' is larger than '' masago'' (capelin roe), but smaller than ''ikura'' (salmon roe). Natural ...
''. It is also a staple in the diet of the
Tao people The Tao people are an Austronesian ethnic group native to the tiny outlying Orchid Island of Taiwan. They have a maritime culture, with great ritual and spiritual significance placed on boat-building and fishing. Their ways of life have been th ...
of
Orchid Island Orchid Island, also known by other names, is a volcanic island off the southeastern coast of Taiwan Island. The island is part of Taiwan. It is separated from the Batanes of the Philippines by the Bashi Channel of the Luzon Strait. It is g ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...
. Flying fish is part of the national dish of Barbados, ''
cou-cou Cou-cou, coo-coo (as it is known in the Windward Islands), or fungie (as it is known in the Leeward Islands and Dominica) makes up part of the national dishes of Antigua and Barbuda, Barbados, British Virgin Islands and the U.S. Virgin Islands. ...
'' and flying fish. The taste is close to that of a sardine. Flying fish roe is known as cau-cau in southern Peru, and is used to make several local dishes. In the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
, the fish are caught while they are flying, using nets held from
outrigger canoe Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull. They can range from small dugout canoes to large plank-built vessels. Outrigger ...
s. They are attracted to the light of torches. Fishing is done only when no moonlight is available.


Importance


Barbados

Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estima ...
is known as "the land of the flying fish", and the fish is one of the national symbols of the country. Once abundant, it migrated between the warm,
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
-filled
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
surrounding the island of Barbados and the plankton-rich outflows of the
Orinoco The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wor ...
River in
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. Just after the completion of the Bridgetown Harbor / Deep Water Harbor in Bridgetown, Barbados had an increase of ship visits, linking the island to the world. The overall health of the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s surrounding Barbados suffered due to ship-based
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, th ...
. Additionally, Barbadian
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in t ...
pushed them closer to the Orinoco delta, no longer returning to Barbados in large numbers. Today, the flying fish only migrate as far north as
Tobago Tobago () is an island and ward within the Republic of Trinidad and Tobago. It is located northeast of the larger island of Trinidad and about off the northeastern coast of Venezuela. It also lies to the southeast of Grenada. The offic ...
, around southwest of Barbados. Despite the change, flying fish remain a coveted delicacy. Many aspects of Barbadian culture center around the flying fish; it is depicted on coins, as sculptures in fountains, in artwork, and as part of the official logo of the Barbados Tourism Authority. Additionally, the Barbadian coat of arms features a
pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before ...
and
dolphinfish ''Coryphaena'' is a genus of marine ray-finned fishes known as the dolphinfishes, and is currently the only known genus in the family Coryphaenidae. The generic name is from Greek κορυφή (''koryphē'', "crown, top") and -αινα (-''ain ...
on either side of the shield, but the dolphinfish resembles a flying fish. Furthermore, actual artistic renditions and holograms of the flying fish are also present within the
Barbadian passport A Barbados passport is a travel document issued to citizens of Barbados, in accordance with Citizenship Act (CAP. 186) from 1978, the Immigration Act (CAP. 190) from 1997, and the Barbados Constitution, for the purpose of facilitating internation ...
.


Maritime disputes

Flying fish have also been gaining in popularity in other islands, fueling several maritime disputes. In 2006, the council of the
United Nations Convention on the Law of the Sea The United Nations Convention on the Law of the Sea (UNCLOS), also called the Law of the Sea Convention or the Law of the Sea Treaty, is an international agreement that establishes a legal framework for all marine and maritime activities. , 167 ...
fixed the
maritime boundaries A maritime boundary is a conceptual division of the Earth's water surface areas using physiographic or geopolitical criteria. As such, it usually bounds areas of exclusive national rights over mineral and biological resources,VLIZ Maritime Bound ...
between
Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estima ...
and
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
over the flying fish dispute, which gradually raised tensions between the neighbours. The ruling stated both countries must preserve stocks for the future. Barbadian fishers still follow the flying fish southward.


Prehistoric analogues

The oldest known fossil of a flying or gliding fish are those of the extinct family
Thoracopteridae Thoracopteridae is an extinct family of prehistoric bony fish; classified with the order Peltopleuriformes. This lineage of Triassic flying fish-like Perleidiformes converted their pectoral and pelvic fins into broad wings very similar to those ...
, dating back to the
Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma an ...
, 235–242 million years ago. However, they are not related to modern flying fish, with the wing-like pectoral fins being
convergently evolved Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
in both lineages.


See also

*
Flying and gliding animals A number of animals are capable of aerial locomotion, either by powered flight or by gliding. This trait has appeared by evolution many times, without any single common ancestor. Flight has evolved at least four times in separate animals: i ...
* Neuston *
Tradeoffs for locomotion in air and water Certain species of fish and birds are able to locomote in both air and water, two fluid media with very different properties. A fluid is a particular phase of matter that deforms under shear stresses and includes any type of liquid or gas. Becau ...


References


External links


Flying Fish, National Geographic Society
{{Authority control Exocoetidae Beloniformes Gliding animals Fish of Oceania Marine fish of Africa Marine fish of Asia Fish of Hawaii Fish of Japan Extant Miocene first appearances Miocene fish Quaternary fish Barbadian cuisine