Floral diagram
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Floral diagram is a graphic representation of
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
structure. It shows the number of floral organs, their arrangement and fusion. Different parts of the flower are represented by their respective symbols. Floral diagrams are useful for flower identification or can help in understanding angiosperm
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
. They were introduced in the late 19th century and are generally attributed to A. W. Eichler. They are typically used with the
floral formula A Floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
of that flower to study its
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
.


History

In the 19th century, two contrasting methods of describing the flower were introduced: the textual
floral formula A Floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e and pictorial floral diagrams. Floral diagrams are credited to A. W. Eichler, his extensive work ''Blüthendiagramme'' (1875, 1878) remains a valuable source of information on floral morphology. Eichler inspired later generation of scientists, including John Henry Schaffner. Diagrams were included e.g. in ''Types of Floral Mechanism'' by
Church Church may refer to: Religion * Church (building), a building for Christian religious activities * Church (congregation), a local congregation of a Christian denomination * Church service, a formalized period of Christian communal worship * C ...
(1908). They were used in different textbooks, e.g. ''Organogenesis of Flowers'''' ''by Sattler (1973), ''Botanische Bestimmungsübungen'' by Stützel (2006) or ''Plant Systematics'' by Simpson (2010). ''Floral Diagrams'' (2010) by Ronse De Craene followed Eichler’s approach using the contemporary APG II was contemporary at the time of book’s writing.
APG II system The APG II system (Angiosperm Phylogeny Group II system) of plant classification is the second, now obsolete, version of a modern, mostly molecular-based, system of plant taxonomy that was published in April 2003 by the Angiosperm Phylogeny Gr ...
.


Basic characteristics and significance

A floral diagram is a schematic cross-section through a young flower. It may be also defined as “projection of the flower perpendicular to its axis”. It usually shows the number of floral parts,It also shows organs that are not part of the flower, but may be closely associated with it, such as bracts and bracteoles. their sizes, relative positions and fusion. Different organs are represented by distinguishable symbols, which may be uniform for one organ type, or may reflect concrete
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
. The diagram may also include symbols that don’t represent physical structures, but carry additional information (e.g. symmetry plane orientation). There is no agreement on how floral diagrams should be drawn, it depends on the author whether it is just a rough representation, or whether structural details of the flower are included. Diagrams can describe the
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the s ...
of flowers, or can show evolutionary relationships. They can be generalized to show the typical floral structure of a
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
. It is also possible to represent (partial)
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
s by diagrams. Substantial amount of information may be included in a good diagram. It can be useful for flower identification or comparison between angiosperm taxa. Paleontologists can take advantage of diagrams for reconstruction of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
flowers. Floral diagrams are also of didactic value.


Orientation

Diagrams are usually depicted with the subtending bract ''below'' and the axis ''above'' the flower itself, both in the median line. The axis corresponds to the position of the main stem relative to a lateral flower. When a terminal flower is depicted, the axis is not present and therefore cannot be shown.
Bracteole In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
s, if they are present, are usually drawn on the sides of the diagram.


Symbols used in diagrams

Not only the information contained within diagrams, but also their appearance commonly varies between authors. Just some publications incorporate an overview of used symbols.


Bracts and bracteoles, axes

Bracts and bracteoles are commonly shown as arcs. In ''Floral Diagrams'' by Ronse De Craene they consistently have a black fill and a little triangle on the outer side to distinguish them from the perianth. In Eichler’s ''Blüthendiagramme'' their representation alters between diagrams. The axis relative to the flower is shown as black circle in ''Floral Diagrams''. When inflorescence is depicted, the position of its main stem is illustrated by a crossed circle. Eichler’s depiction of axes alternates between diagrams.


Perianth

Perianth parts are also shown as arcs. They may be colored according to their type. In ''Blüthendiagramme'' the
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s are usually white with black stroke,
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s are hatched and petals are black. Ronse De Craene implies that it may be sometimes impossible to classify the organs, he shows green perianth parts as black and pigmented as white.
Estivation Aestivation ( la, aestas (summer); also spelled estivation in American English) is a state of animal dormancy, similar to hibernation, although taking place in the summer rather than the winter. Aestivation is characterized by inactivity and a ...
can be accurately shown in the diagram.


Androecium

Stamens are represented by a cross-section through anthers. In case there are many stamens in the flower, they can be simplified and drawn as circles. Staminodes have a small black circle inside or are painted black in ''Floral Diagrams'', Eichler also fills them black.


Gynoecium

The pistil is shown as a sectional view of the ovary. Ovary position is highlighted by small triangles in ''Floral Diagrams''. Ronse De Craene also incorporates ovule morphology or shows the position of stigmatic lobes by white shapes.


Nectaries

In ''Floral Diagrams'', nectaries are filled by grey color, Eichler fills them by hatching.


Other

Fusion can be shown in diagrams by full connecting lines between organs. Lost organs can be represented by a star (✶), lost perianth parts or bracts/bracteoles can be shown with dashed stroke. It is possible to show the direction of monosymmetry by a large arrow.
Resupination Resupination is derived from the Latin word ''resupinus'', meaning "bent back with the face upward" or "on the back". " Resupination" is the noun form of the adjective "resupine" which means "being upside-down, supine or facing upward". The word ...
may be illustrated by a curved arrow. Floral parts can be accompanied by numbers to show their sequence of initialization.


Floral diagrams and floral formulae

Each of these two concepts is better in expressing some information. Floral diagrams can show the size and relative position of the organs. On the other hand,
floral formula A Floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e are capable of broader generalization. Prenner ''et al.'' view them as complementary methods and state they make an “identikit” flower when utilized together. Ronse De Craene also approves of their combined use.


Examples


See also

*
Floral formula A Floral formula is a notation for representing the structure of particular types of flowers. Such notations use numbers, letters and various symbols to convey significant information in a compact form. They may represent the floral form of a part ...
e *
Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
* Floral organs *
Dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...
flower *
Monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. Th ...
flower *
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...


Notes


References


Bibliography

*


External links


Floral diagram generator
{{botany Flowers Plant morphology