Euphotic depth
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The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a
body of water A body of water or waterbody (often spelled water body) is any significant accumulation of water on the surface of Earth or another planet. The term most often refers to oceans, seas, and lakes, but it includes smaller pools of water such a ...
that receives sunlight, allowing phytoplankton to perform
photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored i ...
. It undergoes a series of physical, chemical, and biological processes that supply nutrients into the upper
water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
. The photic zone is home to the majority of
aquatic life An aquatic ecosystem is an ecosystem formed by surrounding a body of water, in contrast to land-based terrestrial ecosystems. Aquatic ecosystems contain communities of organisms that are dependent on each other and on their environment. The tw ...
due to the activity (
primary production In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...
) of the phytoplankton.


Photosynthesis in photic zone

In the photic zone, the photosynthesis rate exceeds the respiration rate. This is due to the abundant solar energy which is used as an energy source for photosynthesis by
primary producers An autotroph or primary producer is an organism that produces complex organic compounds (such as carbohydrates, fats, and proteins) using carbon from simple substances such as carbon dioxide,Morris, J. et al. (2019). "Biology: How Life Work ...
such as phytoplankton. These phytoplankton grow extremely quickly because of sunlight's heavy influence, enabling it to be produced at a fast rate. In fact, ninety five percent of photosynthesis in the ocean occurs in the photic zone. Therefore, if we go deeper, beyond the photic zone, such as into the compensation point, there is little to no phytoplankton, because of insufficient sunlight. The zone which extends from the base of the euphotic zone to the aphotic zone is sometimes called the dysphotic zone.


Life in the photic zone

Ninety percent of marine life lives in the photic zone, which is approximately two hundred meters deep. This includes phytoplankton (plants), including dinoflagellates, diatoms, cyanobacteria,
coccolithophore Coccolithophores, or coccolithophorids, are single celled organisms which are part of the phytoplankton, the autotrophic (self-feeding) component of the plankton community. They form a group of about 200 species, and belong either to the king ...
s, and
cryptomonad The cryptomonads (or cryptophytes) are a group of algae, most of which have plastids. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anteri ...
s. It also includes
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, the consumers in the photic zone. There are carnivorous meat eaters and
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
plant eaters. Next, copepods are the small
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can ...
s distributed everywhere in the photic zone. Finally, there are
nekton Nekton or necton (from the ) refers to the actively swimming aquatic organisms in a body of water. The term was proposed by German biologist Ernst Haeckel to differentiate between the active swimmers in a body of water, and the passive organisms t ...
(animals that can propel themselves, like fish, squids, and crabs), which are the largest and the most obvious animals in the photic zone, but their quantity is the smallest among all the groups. The depth of the photic zone depends on the transparency of the water. If the water is very clear, the photic zone can become very deep. If it is very murky, it can be only fifty feet (fifteen meters) deep.


Nutrients uptake in the photic zone

Due to biological uptake, the photic zone has relatively low levels of nutrient concentrations. As a result, phytoplankton doesn't receive enough nutrients when there is high water-column stability. The
spatial distribution A spatial distribution in statistics is the arrangement of a phenomenon across the Earth's surface and a graphical display of such an arrangement is an important tool in geographical and environmental statistics. A graphical display of a spatial d ...
of organisms can be controlled by a number of factors. Physical factors include: temperature, hydrostatic pressure, turbulent mixing such as the upward turbulent flux of inorganic nitrogen across the nutricline. Chemical factors include oxygen and trace elements. Biological factors include grazing and migrations. Upwelling carries nutrients from the deep waters into the photic zone, strengthening phytoplankton growth. The remixing and upwelling eventually bring nutrient-rich wastes back into the photic zone. The Ekman transport additionally brings more nutrients to the photic zone. Nutrient pulse frequency affects the phytoplankton competition. Photosynthesis produces more of it. Being the first link in the food chain, what happens to phytoplankton creates a rippling effect for other species. Besides phytoplankton, many other animals also live in this zone and utilize these nutrients. The majority of ocean life occurs in the photic zone, the smallest ocean zone by water volume. The photic zone, although small, has a large impact on those who reside in it.


Photic zone depth

The depth is, by definition, where radiation is degraded down to 1% of its surface strength. Accordingly, its thickness depends on the extent of light
attenuation In physics, attenuation (in some contexts, extinction) is the gradual loss of flux intensity through a medium. For instance, dark glasses attenuate sunlight, lead attenuates X-rays, and water and air attenuate both light and sound at variabl ...
in the water column. As incoming light at the surface can vary widely, this says little about the net growth of phytoplankton. Typical euphotic depths vary from only a few centimetres in highly
turbid Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality. Fluids can ...
eutrophic Eutrophication is the process by which an entire body of water, or parts of it, becomes progressively enriched with minerals and nutrients, particularly nitrogen and phosphorus. It has also been defined as "nutrient-induced increase in phytoplan ...
lakes, to around 200 meters in the open
ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...
. It also varies with seasonal changes in turbidity, which can be strongly driven by phytoplankton concentrations, such that the depth of the photic zone often decreases as
primary production In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...
increases. Moreover, the
respiration rate The respiration rate is a parameter which is used in ecological and agronomical modeling. In theoretical production ecology and aquaculture, it typically refers to respiration per unit of time (usually loss of biomass by respiration per unit of ...
is actually greater than the photosynthesis rate. The reason why phytoplankton production is so important is because it plays a prominent role when interwoven with other
food web A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one o ...
s.


Light attenuation

Most of the solar energy reaching the Earth is in the range of visible light, with wavelengths between about 400-700 nm. Each colour of visible light has a unique wavelength, and together they make up white light. The shortest wavelengths are on the violet and ultraviolet end of the spectrum, while the longest wavelengths are at the red and infrared end. In between, the colours of the visible spectrum comprise the familiar “ROYGBIV”; red, orange, yellow, green, blue, indigo, and violet.Webb, Paul (2019
''Introduction to Oceanography''
chapter 6.5 Light, Rebus Community, Roger Williams University, open textbook. Material was copied from this source, which is available under
Creative Commons Attribution 4.0 International License
Water is very effective at absorbing incoming light, so the amount of light penetrating the ocean declines rapidly (is attenuated) with depth. At one metre depth only 45% of the solar energy that falls on the ocean surface remains. At 10 metres depth only 16% of the light is still present, and only 1% of the original light is left at 100 metres. No light penetrates beyond 1000 metres. In addition to overall attenuation, the oceans absorb the different wavelengths of light at different rates. The wavelengths at the extreme ends of the visible spectrum are attenuated faster than those wavelengths in the middle. Longer wavelengths are absorbed first; red is absorbed in the upper 10 metres, orange by about 40 metres, and yellow disappears before 100 metres. Shorter wavelengths penetrate further, with blue and green light reaching the deepest depths. This is why things appear blue underwater. How colours are perceived by the eye depends on the wavelengths of light that are received by the eye. An object appears red to the eye because it reflects red light and absorbs other colours. So the only colour reaching the eye is red. Blue is the only colour of light available at depth underwater, so it is the only colour that can be reflected back to the eye, and everything has a blue tinge under water. A red object at depth will not appear red to us because there is no red light available to reflect off of the object. Objects in water will only appear as their real colours near the surface where all wavelengths of light are still available, or if the other wavelengths of light are provided artificially, such as by illuminating the object with a dive light. Water in the
open ocean The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
appears clear and blue because it contains much less particulate matter, such as phytoplankton or other suspended particles, and the clearer the water, the deeper the light penetration. Blue light penetrates deeply and is scattered by the water molecules, while all other colours are absorbed; thus the water appears blue. On the other hand, coastal water often appears greenish. Coastal water contains much more suspended
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel ...
and algae and
microscopic organisms A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
than the open ocean. Many of these organisms, such as phytoplankton, absorb light in the blue and red range through their photosynthetic pigments, leaving green as the dominant wavelength of reflected light. Therefore the higher the phytoplankton concentration in water, the greener it appears. Small silt particles may also absorb blue light, further shifting the colour of water away from blue when there are high concentrations of suspended particles. The ocean can be divided into depth layers depending on the amount of light penetration, as discussed in pelagic zone. The upper 200 metres is referred to as the photic or euphotic zone. This represents the region where enough light can penetrate to support photosynthesis, and it corresponds to the epipelagic zone. From 200 to 1000 metres lies the dysphotic zone, or the twilight zone (corresponding with the mesopelagic zone). There is still some light at these depths, but not enough to support photosynthesis. Below 1000 metres is the aphotic (or midnight) zone, where no light penetrates. This region includes the majority of the ocean volume, which exists in complete darkness.


Paleoclimatology

Phytoplankton are
unicellular A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...
microorganisms which form the base of the ocean food chains. They are dominated by diatoms, which grow silicate shells called
frustule A frustule is the hard and porous cell wall or external layer of diatoms. The frustule is composed almost purely of silica, made from silicic acid, and is coated with a layer of organic substance, which was referred to in the early literature on ...
s. When diatoms die their shells can settle on the
seafloor The seabed (also known as the seafloor, sea floor, ocean floor, and ocean bottom) is the bottom of the ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth an ...
and become
microfossil A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, ...
s. Over time, these microfossils become buried as
opal Opal is a hydrated amorphous form of silica (SiO2·''n''H2O); its water content may range from 3 to 21% by weight, but is usually between 6 and 10%. Due to its amorphous property, it is classified as a mineraloid, unlike crystalline form ...
deposits in the marine sediment.
Paleoclimatology Paleoclimatology (American and British English spelling differences, British spelling, palaeoclimatology) is the study of climates for which direct measurements were not taken. As instrumental records only span a tiny part of Earth's history, the ...
is the study of past climates.
Proxy data In the study of past climates (" paleoclimatology"), climate proxies are preserved physical characteristics of the past that stand in for direct meteorological measurements and enable scientists to reconstruct the climatic conditions over a longe ...
is used in order to relate elements collected in modern-day sedimentary samples to climatic and oceanic conditions in the past. Paleoclimate proxies refer to preserved or fossilized physical markers which serve as substitutes for direct meteorological or ocean measurements. An example of proxies is the use of diatom isotope records of
δ13C In geochemistry, paleoclimatology, and paleoceanography ''δ''13C (pronounced "delta c thirteen") is an isotopic signature, a measure of the ratio of stable isotopes 13C : 12C, reported in parts per thousand (per mil, ‰). The measure is also ...
,
δ18O In geochemistry, paleoclimatology and paleoceanography ''δ''18O or delta-O-18 is a measure of the ratio of stable isotopes oxygen-18 (18O) and oxygen-16 (16O). It is commonly used as a measure of the temperature of precipitation, as a measur ...
, δ30Si (δ13Cdiatom, δ18Odiatom, and δ30Sidiatom). In 2015, Swann and Snelling used these isotope records to document historic changes in the photic zone conditions of the north-west
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
, including nutrient supply and the efficiency of the soft-tissue
biological pump The biological pump (or ocean carbon biological pump or marine biological carbon pump) is the ocean's biologically driven sequestration of carbon from the atmosphere and land runoff to the ocean interior and seafloor sediments.Sigman DM & GH ...
, from the modern day back to
marine isotope stage 5e Marine Isotope Stage 5 or MIS 5 is a marine isotope stage in the geologic temperature record, between 130,000 and 80,000 years ago. Sub-stage MIS 5e, called the Eemian or Ipswichian, covers the last major interglacial period before the Holocene, w ...
, which coincides with the last interglacial period. Peaks in opal productivity in the marine isotope stage are associated with the breakdown of the regional halocline stratification and increased nutrient supply to the photic zone. Material was copied from this source, which is available under
Creative Commons Attribution 3.0 International License
File:Ocean changes from the last interglacial to the modern day A.png File:Ocean changes from the last interglacial to the modern day B.png The initial development of the halocline and stratified water column has been attributed to the onset of major Northern Hemisphere glaciation at 2.73 Ma, which increased the flux of freshwater to the region, via increased monsoonal rainfall and/or glacial meltwater, and sea surface temperatures. The decrease of
abyssal The abyssal zone or abyssopelagic zone is a layer of the pelagic zone of the ocean. "Abyss" derives from the Greek word , meaning bottomless. At depths of , this zone remains in perpetual darkness. It covers 83% of the total area of the ocean a ...
water upwelling associated with this may have contributed to the establishment of globally cooler conditions and the expansion of glaciers across the Northern Hemisphere from 2.73 Ma. While the halocline appears to have prevailed through the
late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
and early Quaternary glacial–interglacial cycles, other studies have shown that the stratification boundary may have broken down in the
late Quaternary The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene together ...
at glacial terminations and during the early part of interglacials.


See also

* Mesophotic coral reef *
Electromagnetic absorption by water The absorption of electromagnetic radiation by water depends on the state of the water. The absorption in the gas phase occurs in three regions of the spectrum. Rotational transitions are responsible for absorption in the microwave and far-inf ...
*
Epipelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral ree ...


References

{{physical oceanography, expanded=none Aquatic ecology Oceanographical terminology