Endomychidae
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Endomychidae, or handsome fungus beetles, is a family of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s with representatives found in all
biogeographic realm A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. De ...
s. There are around 120 genera and 1300 species. The family was established based on the type genus '' Endomychus'', a genus erected in 1795 by
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which was applied to a species that Linnaeus called ''Chrysomela coccinea''. As the common name suggests, Endomychidae feed on
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
. Crowson, in his influential treatment of the beetles, placed the family within the
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark be ...
. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and
Eupsilobiidae Eupsilobiidae is a family of beetles in the superfamily Coccinelloidea, formerly included within the family Endomychidae. Most genera are restricted to the Neotropics, while the genus '' Eidoreus'' is found worldwide. They are fungivore Fungivo ...
. The subfamilies that are included: * Danascelinae * Endomychinae (including Stenotarsinae) * Epipocinae * Leiestinae * Lycoperdininae * Merophysiinae * Pleganophorinae * Xenomycetinae


Genera

These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate): * ''
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'' Gerstaecker, 1858 g * '' Aclemmysa'' Reitter, 1904 g * '' Agaricophilus'' Motschulsky, 1838 g * '' Amphisternus'' Germar, 1843 g * '' Amphistethus'' Strohecker, 1964 g * '' Ancylopus'' Costa, 1854 g * '' Anidrytus'' Gerstaecker, 1858 i c g * '' Aphorista'' Gorham, 1873 i c g b * '' Archipines'' Strohecker, 1953 i c g * '' Asymbius'' Gorham, 1896 g * '' Atopomychus'' Tomaszewska & Szawaryn, 2013 g * '' Atrichonota'' Arrow, 1925 g * '' Avencymon'' Strohecker, 1971 g * '' Beccariola'' Arrow, 1943 g * '' Bolbomorphus'' Gorham, 1888 g * '' Brachytrycherus'' Arrow, 1920 g * '' Bystodes'' Strohecker, 1953 g * '' Cholovocera'' Motschulsky, 1838 g * '' Chondria'' Gorham, 1888 g * '' Corynomalus'' Chevrolat in Dejean, 1836 i c g * '' Cyclotoma'' Mulsant, 1851 i g * '' Danae'' Reiche, 1847 i c g b * '' Danascelis'' Tomaszewska, 1999 i c g * '' Dapsa'' Latreille, 1829 g * '' Dexialia'' Sasaji, 1970 g * ''
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'' Gorham, 1891 g * '' Discolomopsis'' Shockley, 2006 g * '' Displotera'' Reitter, 1877 g * '' Ectomychus'' Gorham, 1888 g * '' Eidorus'' g * '' Endomychus'' Panzer, 1795 i c g b * '' Ephebus'' Chevrolat in Dejean, 1836 i c g * '' Epipocus'' Germar, 1843 i c g b * '' Epopterus'' Chevrolat in Dejean, 1836 i c g * '' Eucteanus'' Gerstaecker, 1857 g * '' Eucymon'' g * '' Eumorphus'' Weber, 1801 g * '' Geoendomychus'' Lea, 1922 g * '' Glesirhanis'' Shockley & Alekseev, 2014 g * '' Hadromychus'' Bousquet & Leschen, 2002 i c g b * '' Holoparamecus'' Curtis, 1833 i c g b * '' Hylaia'' Guerin, 1857 g * '' Hylaperdina'' Tomaszewska, 2012 g * '' Idiophyes'' Blackburn, 1896 g * '' Indalmus'' Gerstaecker, 1858 g * '' Leiestes'' Chevrolat, 1836 g * '' Lycoperdina'' Latreille, 1807 i c g b * '' Meilichius'' Gerstaecker, 1857 g * '' Merophysia'' Lucas, 1852 g * '' Microxenus'' Wallaston, 1861 g * '' Mycetina'' Mulsant, 1846 i c g b * '' Mychothenus'' Strohecker, 1953 g * '' Natalinus'' Tomaszewska, 2011 g * '' Ohtaius'' Chûjô, 1938 g * '' Palaeoestes'' Kirejtshuk & Nel, 2009 g * '' Paniegena'' Heller, 1916 g * '' Parasymbius'' Arrow, 1920 g * '' Phymaphora'' Newman, 1838 i c g b * '' Phymaphoroides'' Motschoulsky, 1857 g * '' Pleganophorus'' Hampe, 1855 g * '' Polymus'' Mulsant, 1846 g * '' Pseudindalmus'' Arrow, 1920 g * '' Rhanidea'' Strohecker, 1953 i c g b * '' Saula'' Gerstaecker, 1858 i c g * '' Sinocymbachus'' Strohecker & Chujo, 1970 g * '' Stenotarsus'' Perty, 1832 i c g b * '' Stethorhanis'' Blaisdell, 1931 i c g * '' Trochoideus'' Westwood, 1833 i c g b * '' Trycherus'' Gerstaecker, 1857 g * '' Xenomycetes'' Horn, 1880 i c g b Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


Fossil genera

* '' Burmalestes'' Tomaszewska and Ślipiński, 2018
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, Late Cretaceous ( Cenomanian) * '' Cretolestes'' Tomaszewska, Ślipiński and Ren, 2018 Burmese amber, Myanmar, Cenomanian * '' Cretaparamecus'' Tomaszewska, Ślipiński, Bai and Zhang, 2018 Burmese amber, Myanmar, Cenomanian * '' Palaeomycetes'' Tomaszewska, Ślipiński and Ren, 2018 Burmese amber, Myanmar, Cenomanian * '' Zemyna'' Tomaszewska, 2018 (replaced name ''Laima'' Alekseev and Tomaszewska, 2018'')'':
Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than ...
, Eocene


References


External links

* * {{Taxonbar, from=Q1076939 Coccinelloidea Polyphaga families