Elateroidea
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The Elateroidea are a large
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species.


Description

Elateroidea is a morphologically diverse group, including hard-bodied beetles with 5 abdominal ventrites, soft-bodied beetles with 7-8 ventrites connected with membranes (formerly known as cantharoids), and beetles with intermediate forms. They have a range of sizes and colours, but in terms of shape, they are usually narrow and parallel-sided as adults. Many of the sclerotised elateroids ( Cerophytidae, Eucnemidae, Throscidae, Elateridae) have a clicking mechanism. This is a peg on the prothorax which fits into a cavity in the mesothorax. When a click beetle bends its body, the peg snaps into the cavity, causing the beetle's body to straighten so suddenly that it jumps into the air. Most beetles capable of
bioluminescence Bioluminescence is the production and emission of light by living organisms. It is a form of chemiluminescence. Bioluminescence occurs widely in marine vertebrates and invertebrates, as well as in some fungi, microorganisms including some b ...
are in the Elateroidea, in the families Lampyridae (~2000 species), Phengodidae (~200 species), Rhagophthalmidae (100 species) and Elateridae (>100 species). Females in several lineages, including
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
, Lampyridae, Phengogidae and Rhagophthalmidae, do not pupate and remain in a larval form. This trait is estimated to have evolved independently at least three times within the superfamily. Some Elateroidea, including species of Cantharidae and Lycidae, have bright aposematic colours to signal to predators that they are poisonous and so should not be eaten.


Families

The validity and relationships of some families, such as Podabrocephalidae are not fully resolved. The family Rhinorhipidae has recently been removed to its own superfamily, with evidence that it is a basal taxon within Elateriformia dating to an Upper Triassic/Lower Jurassic split from other extant beetle lineages. * Artematopodidae Lacordaire, 1857 – soft-bodied plant beetles (= Eurypogonidae) * Brachypsectridae Leconte & Horn, 1883 – Texas beetles *
Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the red coats of early British soldiers, hence the ...
Imhoff, 1856 – soldier beetles * Cerophytidae Latreille, 1834 – rare click beetles * Elateridae
Leach Leach may refer to: * Leach (surname) * Leach, Oklahoma, an unincorporated community, United States * Leach, Tennessee, an unincorporated community, United States * Leach Highway, Western Australia * Leach orchid * Leach phenotype, a mutatio ...
, 1815
– click beetles (including Ampedidae, Balgidae, Dicronychidae, Drilidae, Lissomidae, Plastoceridae, Prosternidae, Protelateridae, Pyrophoridae, Synaptidae) * Eucnemidae Eschscholtz, 1829false click beetles (including Anischiidae and Perothopidae) * Iberobaeniidae Bocak ''et al.'', 2016 * Jurasaidae Rosa, Costa, Klamp & Kundrata, 2020 *
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
– net-winged beetles * Omethidae LeConte, 1861false firefly beetles (including Telegeusidae) * Throscidae Laporte, 1840false metallic wood-boring beetles (= Trixagidae) * † Mysteriomorphidae Alekseev and Ellenberger, 2019 (Cretaceous) * Lampyroid clade ** † Cretophengodidae Li, Kundrata, Tihelka & Cai, 2021 (Cretaceous) **
Lampyridae The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
Latreille, 1817 – firefly beetles ** Phengodidae LeConte 1861 – glowworm beetles (including Cydistinae) ** Rhagophthalmidae
Olivier Olivier is the French form of the given name Oliver. It may refer to: * Olivier (given name), a list of people and fictional characters * Olivier (surname), a list of people * Château Olivier, a Bordeaux winery * Olivier, Louisiana, a rural po ...
, 1907
(sometimes in Phengodidae, might belong in Lampyridae) ** Sinopyrophoridae Bi & Li, 2019 *''Incertae sedis'': **†'' Anoeuma'' Li, Kundrata & Cai, 2022 (Cretaceous)


Phylogeny

Some morphological and molecular phylogenetic analyses find that
Byrrhoidea Byrrhoidea is a superfamily of beetles belonging to Elateriformia that includes several families which are either aquatic or associated with a semi-aquatic habitat. Other than the superfamily Hydrophiloidea, most of the remaining Polyphagan beetl ...
is either a monophyletic or paraphyletic group closely related to Elateroidea. Based on Kusy et al. 2018 and 2020


References


External links


Tree of Life - Elateroidea
{{DEFAULTSORT:Elateroidea Beetle superfamilies