Echinacea purpurea
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''Echinacea purpurea'', the eastern purple coneflower, purple coneflower, hedgehog coneflower, or echinacea, is a
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n
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
in the family
Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...
. It is
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to parts of eastern North America and present to some extent in the wild in much of the eastern, southeastern and midwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
as well as in the
Canadian Province Within the geographical areas of Canada, the ten provinces and three territories are sub-national administrative divisions under the jurisdiction of the Canadian Constitution. In the 1867 Canadian Confederation, three provinces of British Nor ...
of
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
. It is most common in the
Ozarks The Ozarks, also known as the Ozark Mountains, Ozark Highlands or Ozark Plateau, is a physiographic region in the U.S. states of Missouri, Arkansas, Oklahoma and the extreme southeastern corner of Kansas. The Ozarks cover a significant port ...
and in the
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/
Ohio Valley The Ohio River is a long river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing southwesterly from western Pennsylvania to its mouth on the Mississippi River at the southern tip of Illin ...
. Its
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s include dry open woods, prairies and barrens.


Taxonomy

''Echinacea'' is derived from
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
, meaning 'spiny one', in reference to the spiny sea urchins 'εχίνοι' which the ripe flower heads of species of this genus resemble. The epithet ''purpurea'' means 'reddish-purple'. Originally named ''Rudbeckia purpurea'' by Linnaeus in 1753 in Species plantarum 6, it was reclassified in 1794 by Conrad Moench, in a new genus named ''Echinacea purpurea'' (L.) Moench. In 1818,
Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an English botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle in the West Riding of Yorkshire and ...
describes a new variety that he named ''Rudbeckia purpurea'' var. serotina. Just two decades later, De Candolle raised him to the rank of species of the other genus ''Echinacea serotina'' (Nutt.) DC. (1836). In 2002, Binns et al. discovered a misapplication of the name ''Echinacea purpurea'' for the taxon correctly named ''Echinacea serotina'' in 1836. The authors proposed to retain the names not to cause confusion among gardeners and herbalists. Other names include: Broad-leaved purple coneflower, Eastern Purple Coneflower, Hedgehog Coneflower, Echinacea.


Description

''Echinacea purpurea'' is an
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
up to tall by wide at maturity. Depending on the climate, it blooms throughout summer into autumn. Its cone-shaped
flowering heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
are usually, but not always, purple in the wild. Its individual flowers (florets) within the
flower head A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
are
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have s ...
, having both male and female organs in each flower. It is
pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
by butterflies and bees. The alternate leaves, borne by a petiole from 0 to 17 cm, are oval to
lanceolate The following is a list of terms which are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflet (bo ...
, 5-30 x 5-12 cm; the margin is tightened to toothed. The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
is a capitulum, 7 to 15 cm in diameter, formed by a prominent domed central protuberance consisting of multiple small yellow florets. These are surrounded by a ring of pink or purple
ligulate A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf and leafstalk of many grasses (Poaceae) and sedges. A ligule is also a strap-shaped extension of the corolla, such as that of a r ...
florets. The tubular florets are
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...
while the ligular florets are
sterile Sterile or sterility may refer to: *Asepsis Asepsis is the state of being free from disease-causing micro-organisms (such as pathogenic bacteria, viruses, pathogenic fungi, and parasites). There are two categories of asepsis: medical and surgi ...
. The involucral bracts are linear to lanceolate. The plant prefers well-drained soils in full sun. The fruit is an
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not o ...
, sought after by birds.


Cultivation

''Echinacea purpurea'' is grown as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
in temperate regions. It is ideal for curbs, walkways or beds. The flowers can also go into the composition of fresh bouquets. Numerous
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have been developed for flower quality and plant form. The plant grows in sun or light shade. It thrives in either dry or moist soil and can tolerate drought once established. The cultivars 'Ruby Giant' and ='Elbrook' have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


Propagation

''Echinacea purpurea'' is propagated either
vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or speci ...
or from
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s. Useful vegetative techniques include division, root cuttings, and basal cuttings. Clumps can be
divided Division is one of the four basic operations of arithmetic, the ways that numbers are combined to make new numbers. The other operations are addition, subtraction, and multiplication. At an elementary level the division of two natural numbe ...
, or broken into smaller bunches, which is normally done in the spring or autumn. Cuttings made from roots that are "pencil-sized" will develop into plants when started in late autumn or early winter. Cuttings of basal shoots in the spring may be rooted when treated with rooting hormones, such as IBA at 1000 ppm. Seed
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fe ...
occurs best with daily temperature fluctuations or after stratification, which help to end
dormancy Dormancy is a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps an organism to conserve energy. Dormancy tends to be c ...
. Seeds may be started indoors in advance of the growing season or outdoors after the growing season has started.


Ecology

Many pollinators are attracted to ''E. purpea''. Bees that are attracted to the flowers include
bumblebees A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related gene ...
, sweat bees,
honey bees A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosm ...
, the sunflower leafcutter bee, and the mining bee ''Andrena helianthiformis''. Butterflies that visit include
monarchs A monarch is a head of stateWebster's II New College DictionarMonarch Houghton Mifflin. Boston. 2001. p. 707. for life or until abdication, and therefore the head of state of a monarchy. A monarch may exercise the highest authority and power in ...
, swallowtail butterflies, and sulphur butterflies. Birds, particularly finches, eat the seeds and disperse them through their droppings.
Slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a ...
s and
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit sp ...
s will also eat the foliage when young, or shortly after emerging in the spring. Additionally, roots can be damaged and eaten by gophers.


Chemistry

''Echinacea purpurea'' contains alkamides,
caffeic acid Caffeic acid is an organic compound that is classified as a hydroxycinnamic acid. This yellow solid consists of both phenolic and acrylic functional groups. It is found in all plants because it is an intermediate in the biosynthesis of lignin, one ...
derivatives,
polysaccharide Polysaccharides (), or polycarbohydrates, are the most abundant carbohydrates found in food. They are long chain polymeric carbohydrates composed of monosaccharide units bound together by glycosidic linkages. This carbohydrate can react with w ...
s, and
glycoprotein Glycoproteins are proteins which contain oligosaccharide chains covalently attached to amino acid side-chains. The carbohydrate is attached to the protein in a cotranslational or posttranslational modification. This process is known as glyco ...
s. Nicotiflorin is the dominant
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
in ''E. purpurea'', followed by the flavonoid
rutin Rutin, also called rutoside, quercetin-3-O-rutinoside and sophorin, is the glycoside combining the flavonol quercetin and the disaccharide rutinose (α-L-rhamnopyranosyl-(1→6)-β-D-glucopyranose). It is a flavonoid found in a wide variety of pl ...
.


Traditional medicine

Native Americans used the plant as
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
to treat many ailments.


References

{{Taxonbar, from=Q272661 purpurea Flora of the United States Butterfly food plants Garden plants of North America Medicinal plants of North America Plants described in 1753 Taxa named by Carl Linnaeus