Cucujoidea is a

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

beetles Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describe ...

. This group formerly included all of the families now included in the superfamily

Coccinelloidea Coccinelloidea is a superfamily of beetles in the order Coleoptera, formerly included in the superfamily Cucujoidea. There are more than 10,000 species in Coccinelloidea, including more than 6000 in the lady beetle family Coccinellidae. Morphol ...

. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (

Scolytinae A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although the ...

), which are

weevil Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...

s (superfamily

Curculionoidea Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...

Morphology

The morphology of Cucujoidea is varied and there are no features uniting all members of the superfamily. Adults can be recognised by the procoxal cavities being internally open in most taxa, females having tarsal formula 5-5-5 and males 5-5-5 or 5-5-4 (rarely 4-4-4), females with tergite VIII concealed dorsally by tergite VII, and males with tergite X completely membraneous. Larvae have frontal arms usually lyriform, the mandible mesal surface usually with well-developed mola, a maxillary articulating area usually present, a hypopharyngeal sclerome usually present, and two pretarsal setae.

Taxonomy

According to a 2015 revision, the following 25 families make up superfamily Cucujoidea: * Agapythidae Sen Gupta and Crowson, 1969 *

Boganiidae Boganiidae is a family of beetles, in the superfamily Cucujoidea. Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and flowering plants of the f ...

Sen Gupta & Crowson, 1966 *

Cavognathidae Cavognathidae is a family of beetles, in the superfamily Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles an ...

Sen Gupta & Crowson, 1966 * Cryptophagidae Kirby, 1826 – silken fungus beetles * Cucujidae Latreille, 1802 – flat bark beetles * Cybocephalidae Jacquelin du Val, 1858 *

Cyclaxyridae Cyclaxyridae are a family of beetles in the superfamily Cucujoidea. The only living genus is '' Cyclaxyra'', with two species endemic to New Zealand. Other species have been named from fossils. They are also known as sooty mould beetles due to the ...

Gimmel, Leschen & Ślipiński, 2009 – sooty mould beetles *

Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscription, it contains 6 tribes ( Tritomini, Dacnini, Megalodacnini, Er ...

Latreille, 1802 –

pleasing fungus beetle file:Ischyrus quadripunctatus P1390172a.jpg, ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family (biology), family of beetles belonging to Cucujoidea containing over 100 genus, genera. In the present circumscrip ...

s *

Helotidae Helotidae is a family of beetles, in the suborder Polyphaga. The family includes about five extant genera, ''Helota'' MacLeay, ''Neohelota'' Ohta, ''Afrohelotina'' Kirejtshuk, ''Metahelotella'' Kirejtshuk, and ''Strophohelota'' Kirejtshuk. Helo ...

Reitter, 1876 *

Hobartiidae Hobartiidae is a family of beetles, in the superfamily Cucujoidea. There are only two known genera '' Hobartius'' (Sen Gupta & Crowson, 1966) and '' Hydnobioides'' (Sen Gupta & Crowson, 1966), with six species, five of which are native to Austra ...

Sen Gupta & Crowson, 1966 *

Kateretidae Kateretidae also known as short-winged flower beetles are a family of beetles in the superfamily Cucujoidea. There are 10 extant and 4 extinct genera, and at least 40 described species. They are found worldwide except in New Zealand. Adults are ...

Erichson in Agassiz, 1846 –

short-winged flower beetle Kateretidae also known as short-winged flower beetles are a family of beetles in the superfamily Cucujoidea. There are 10 extant and 4 extinct genera, and at least 40 described species. They are found worldwide except in New Zealand. Adults are a ...

s (= Brachypteridae) * Laemophloeidae Ganglbauer 1899 – lined flat bark beetles *

Lamingtoniidae ''Lamingtonium'' is the only genus in the family Lamingtoniidae, of the beetle superfamily Cucujoidea. It contains three species endemic to Australia.Lawrence, J. F., and Leschen, R. A. B. (2003). Review of Lamingtoniidae (Coleoptera: Cucujoidea ...

Sen Gupta & Crowson, 1966 * Monotomidae Laporte, 1840 – root-eating beetles *

Myraboliidae ''Myrabolia'' is the only genus in the beetle family Myraboliidae in the superfamily Cucujoidea. It has about 13 species, found in Australia. Adults and possibly larvae live under the bark of ''Eucalyptus ''Eucalyptus'' () is a genus of over s ...

Lawrence and Britton, 1991 * Nitidulidae Latreille, 1802 – sap beetles * Passandridae Erichson, 1845 – parasitic flat bark beetles * Phalacridae Leach, 1815 – shining flower beetles * Phloeostichidae Reitter, 1911 * Priasilphidae Crowson, 1973 * Protocucujidae Crowson, 1954 *

Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vo ...

Kirby, 1937 – silvanid flat bark beetles * Smicripidae Horn, 1879 – palmetto beetles * Sphindidae Jacquelin du Val, 1860 – dry-fungus beetles * Tasmosalpingidae Lawrence and Britton, 1991 * Parandrexidae Kirejtshuk, 1994 (Middle Jurassic-Early Cretaceous)

Extinct genera

* '' Alloterocucus'' Li et al., 2022

Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...

, Myanmar, mid-

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

(latest

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...

- earliest

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in ...

)

References

External links

* * Beetle superfamilies Taxa named by Pierre André Latreille {{Cucujoidea-stub