Crotalus cerberus
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''Crotalus cerberus'' is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
found in the southwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
. It is known as the Arizona black rattlesnake, black rattlesnake, and several other common names. Wright AH, Wright AA. 1957. Handbook of Snakes. 2 volumes. Comstock Publishing Associates. (7th printing, 1985). 1105 pp. .


Description

According to Wright and Wright (1957), adults grow to an average length of . Klauber (1997) reports the maximum length to be less at , with the smallest gravid female measuring . Klauber LM. 1997. Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. 2 volumes. Reprint, University of California Press, Berkeley. . Hubbs and O'Connor (2012) list an adult size range of .Hubbs, Brian and Brendan O'Connor. 2012. '' A Guide to the Rattlesnakes and other Venomous Serpents of the United States''. Tricolor Books. Tempe, Arizona, 129 pp. . (''Crotalus cerberus'', pp. 34-35.) The color pattern consists of a dark grayish, brownish black, reddish brown, or blackish ground color, overlaid with a dorsal pattern of blotches that are rectangular anteriorly, becoming subhexagonal posteriorly, eventually becoming crossbands just before the tail. However, specimens also may be a uniform dark color without any clear dorsal pattern, or the dorsal blotches may be even darker and bordered with white, cream, or yellow transverse rows of scales, or the color pattern may be quite pale with a significant amount of yellow mixed in. A postocular stripe is evident in lightly colored specimens, but not so much in darker ones.Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. 2 volumes. Comstock Publishing Associates, Ithaca and London. 870 pp. 1500 plates. .


Color change

Arizona black rattlesnakes undergo morphological color change (gradual change due to changes in number or quality of chromatophores); adults are less patterned than juveniles (also called ontogenetic color change). Some adults can change color relatively quickly (physiological color change: rapid change due to movement of organelles within chromatophores), an ability shared not only with
chameleon Chameleons or chamaeleons (family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 202 species described as of June 2015. The members of this family are best known for their distinct range of colors, bein ...
s and other lizards but also with other snakes such as some
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
'' Tropidophis''. The mechanism for their physiological color change is likely the same as that documented in the closely related prairie rattlesnake (melanin movement within their dermal melanophores), but further research is needed to determine the stimuli for this phenomenon in this rattlesnake.


Common names

Arizona black rattlesnake, black rattlesnake, black diamond rattlesnake, brown rattlesnake, Cerberus rattlesnake, mountain diamond-back. Also often incorrectly referred to as a timber rattlesnake.


Geographic range

Found in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
, in
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
from the
Hualapai Mountains The Hualapai Mountains are a mountain range located in Mohave County, east of Kingman, Arizona. Rising up to 8,417 feet at its highest peak, the higher elevations of the Hualapai Mountains support Madrean Sky Island habitats, and are host to ...
and Cottonwood Cliffs in the northwest of the state, southeast to the Santa Catalina, Rincon, Pinaleno and Blue Mountains. Also found at Steeple Rock, in extreme western
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
. The type locality given is "San Francisco Mountains" (
Coconino County, Arizona Coconino County is a county in the north-central part of the U.S. state of Arizona. Its population was 145,101 at the 2020 census. The county seat is Flagstaff. The county takes its name from ''Cohonino'', a name applied to the Havasupa ...
, USA).


Diet

It preys upon suitably sized
amphibian Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbo ...
s,
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchocephalia ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s and their eggs, and
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s.


Reproduction

Sexually mature females bear live young in broods of 4 to 21
neonate An infant or baby is the very young offspring of human beings. ''Infant'' (from the Latin word ''infans'', meaning 'unable to speak' or 'speechless') is a formal or specialised synonym for the common term ''baby''. The terms may also be used to ...
s. The Arizona black rattlesnake is the first species of snake observed to exhibit complex social behavior, and like all temperate pit vipers, care for their babies. Females remain with their young in nests for 7 to 14 days, and mothers have been observed cooperatively parenting their broods.


References


External links


Arizona Black Rattlesnake, ''Crotalus cerberus''
at th
National Park Service
Accessed 18 June 2008. {{Taxonbar, from1=Q3003829, from2=Q5188959
cerberus In Greek mythology, Cerberus (; grc-gre, Κέρβερος ''Kérberos'' ), often referred to as the hound of Hades, is a multi-headed dog that guards the gates of the Underworld to prevent the dead from leaving. He was the offspring of the ...
Snakes of North America Reptiles of the United States Endemic fauna of the United States Fauna of the Southwestern United States Fauna of the Sonoran Desert Natural history of Arizona Natural history of New Mexico Taxa named by Elliott Coues Reptiles described in 1875