Conodont biostratigraphy
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Conodont Conodonts ( Greek ''kōnos'', " cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, whi ...
s are an extinct class of animals whose feeding apparatuses called ''teeth'' or ''elements'' are common microfossils found in strata dating from the Stage 10 of the Furongian, the fourth and final series of the Cambrian, to the Rhaetian stage of the Late Triassic. These elements can be used alternatively to or in correlation with other types of fossils (
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian ( Miaolingian, Wuliuan) through t ...
s,
trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the ...
s,
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefis ...
s, ...) in the subfield of the
stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers ( strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...
named
biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of ...
.


Paleozoic conodonts


Cambrian conodonts

It is suggested that '' Eoconodontus notchpeakensis'' can be a marker of the Stage 10 of the
Furongian The Furongian is the fourth and final epoch and series of the Cambrian. It lasted from to million years ago. It succeeds the Miaolingian series of the Cambrian and precedes the Lower Ordovician Tremadocian Stage. It is subdivided into three sta ...
, the fourth and final series of the Cambrian. In 2006, a working group proposed the first appearance of '' Cordylodus andresi''. Currently the first appearance of ''E. notchpeakensis'' is favored by many authors because it is globally widespread and is independent of facies (known from continental rise to peritidal environments). The ''Eoconodontus notchpeakensis'' proposal would also incorporate a non-biostratigraphic marker to correlate the beginning of Stage 10 globally. A carbon isotope excursion (the HERB-event) occurs in the lower part of the E. notchpeakensis range.


Ordovician conodonts


Early Ordovician

The base of the Tremadocian, the lowest stage of
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...
, is defined as the first appearance of ''
Iapetognathus fluctivagus ''Iapetognathus fluctivagus'' is a species of denticulate cordylodan conodonts belonging to the genus ''Iapetognathus''. It existed during the Tremadocian Age ( million years ago) of the Ordovician. It is an important index fossil in biostratigr ...
'' at the GSSP section in Newfoundland. No conodont species is associated with the Floian, the second stage of the Ordovician.


Middle Ordovician

The base of the Dapingian, the third stage of the Ordovician, is defined as the
first appearance In American comic books and other stories with a long history, first appearance refers to the first issue to feature a fictional character. These issues are often highly valued by collectors due to their rarity and iconic status. Reader interes ...
of ''
Baltoniodus ''Baltoniodus'' is an extinct genus of conodonts. Use in stratigraphy The base of the Dapingian, the third stage of the Middle Ordovician, is defined as the first appearance of ''Baltoniodus triangularis''. The Whiterock Stage refers mainly ...
triangularis''. The base of the
Darriwilian The Darriwilian is the upper stage of the Middle Ordovician. It is preceded by the Dapingian and succeeded by the Upper Ordovician Sandbian Stage. The lower boundary of the Darriwilian is defined as the first appearance of the graptolite specie ...
, the fourth stage of the Ordovician, lies just above the North Atlantic '' Microzarkodina parva'' conodont zone. The base also lies in the upper part of the North American '' Histiodella altifrons'' conodont zone. The Whiterock Stage refers mainly to the early Middle Ordovician in North America. It is often used in the older literature in a global sense. The Whiterock Stage is given a range from 471.8 (ca. 472) to 462 m.y.a., spanning close to 10 million years. Officially its start is defined by the potentially lowest occurrence of the conodont ''
Protoprioniodus aranda ''Protoprioniodus'' is an extinct genus of conodonts. Use in stratigraphy The Whiterock Stage refers mainly to the early Middle Ordovician in North America. It is often used in the older literature in a global sense. The Whiterock Stage is g ...
'' or ''
Baltoniodus ''Baltoniodus'' is an extinct genus of conodonts. Use in stratigraphy The base of the Dapingian, the third stage of the Middle Ordovician, is defined as the first appearance of ''Baltoniodus triangularis''. The Whiterock Stage refers mainly ...
triangularis''.


Late Ordovician

No conodont species is associated with the
Sandbian The Sandbian is the first stage of the Upper Ordovician. It follows the Darriwilian and is succeeded by the Katian. Its lower boundary is defined as the first appearance datum of the graptolite species '' Nemagraptus gracilis'' around million yea ...
, the Katian, nor with the Hirnantian, the fifth, sixth and seventh and final stages of the Ordovician. ''
Baltoniodus ''Baltoniodus'' is an extinct genus of conodonts. Use in stratigraphy The base of the Dapingian, the third stage of the Middle Ordovician, is defined as the first appearance of ''Baltoniodus triangularis''. The Whiterock Stage refers mainly ...
gerdae'' has been found in the early Sandbian
Bromide Formation The Bromide Formation is a geological formation in Oklahoma, USA. It is well known for its diverse echinoderm and trilobite fossil fauna. Location The Bromide Formation crops out in the Arbuckle and Wichita Mountains and in the Criner Hills ...
, in Oklahoma, United States.


Silurian conodonts


Llandovery

The Llandovery epoch lasted from 443.8 ± 1.5 to 433.4 ± 2.8 mya, and is subdivided into three stages: the
Rhuddanian In the geologic timescale, the Rhuddanian is the first age of the Silurian Period and of the Llandovery Epoch. The Silurian is in the Paleozoic Era of the Phanerozoic Eon. The Rhuddanian Age began 443.8 ± 1.5 Ma and ended 440.8 ± 1.2 Ma ( ...
, lasting until 440.8 million years ago, the Aeronian, lasting to 438.5 million years ago, and the Telychian. The
Telychian In the geologic timescale, the Telychian is the age of the Llandovery Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon. The Telychian Age was between 438.5 ± 1.2 million years ago (Ma) and 433.4 ± 0.8 Ma. The Telych ...
(Late Llandovery) of Estonia can be defined by five conodont zones (''
Pterospathodus ''Pterospathodus'' is an extinct genus of conodont from the Silurian period. Use in stratigraphy The Telychian (Late Llandovery) of Estonia can be defined by five conodont zones (''Pterospathodus eopennatus ssp. n. 1'', ''P. eopennatus ssp. ...
eopennatus ssp. n. 1'', ''P. eopennatus ssp. n. 2'', ''P. amorphognathoides angulatus'', ''P. a. lennarti'' and ''P. a. lithuanicus'').An updated Telychian (Late Llandovery, Silurian) conodont zonation based on Baltic faunas. Peep Männik, Lethaia, Volume 40, Issue 1, pp. 45–60, March 2007,


Wenlock

The Wenlock, which lasted from 433.4 ± 1.5 to 427.4 ± 2.8 mya, is subdivided into the Sheinwoodian (to 430.5 million years ago) and Homerian ages. The
Sheinwoodian In the geologic timescale, the Sheinwoodian is the age of the Wenlock Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that is comprehended between 433.4 ± 0.8 Ma and 430.5 ± 0.7 Ma (million years ago), approximatel ...
(Wenlock) is defined between the acritarch biozone 5 and the last appearance of ''Pterospathodus amorphognathoides''. The global boundary stratotype point is in Hughley Brook in Apedale, U.K.


Ludlow

The Ludlow, lasting from 427.4 ± 1.5 to 423 ± 2.8 mya, comprises the Gorstian stage, lasting until 425.6 million years ago, and the Ludfordian stage.


Přídolí


Devonian conodonts


Early Devonian

The Early Devonian lasts from 419.2 ± 2.8 to 393.3 ± 2.5 and begins with the Lochkovian stage, which lasts until the Pragian. This spans from 410.8 ± 2.8 to 407.6 ± 2.5, and is followed by the Emsian, which lasts until the Middle Devonian begins, 393.3± 2.7 million years ago.


Middle Devonian

The Middle Devonian comprises two subdivisions, the Eifelian giving way to the Givetian 387.7± 2.7 million years ago.


Late Devonian

Finally, the Late Devonian starts with the Frasnian, 382.7 ± 2.8 to 372.2 ± 2.5, followed by the Famennian subdivision, the beginning and end of which are marked with extinction events. This lasted until the end of the Devonian, 358.9± 2.5 million years ago. During that stage, a biologic event occurred (Upper Kellwasser Extinction of all '' Ancyrodella'' and '' Ozarkodina'' and most '' Palmatolepis'', '' Polygnathus'' and '' Ancyrognathus''). The
Famennian The Famennian is the latter of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used ...
(372.2 ± 1.6 mya) is defined by a
GSSP Golden Spike A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted by ...
located at Coumiac quarry, Montagne Noire, France where there is a biologic abundant occurrence of '' Palmatolepis triangularis''.


Carboniferous conodonts


Mississippian (also known as Lower Carboniferous)

The Tournaisian, the oldest age of the Mississippian contains eight conodont biozones: * the zone of '' Gnathodus pseudosemiglaber'' and '' Scaliognathus anchoralis'' * the zone of ''Gnathodus semiglaber'' and '' Polygnathus communis'' * the zone of '' Dollymae bouckaerti'' * the zone of ''Gnathodus typicus'' and '' Siphonodella isosticha'' * the zone of ''Siphonodella quadruplicata'' and '' Patrognathus andersoni'' (upper zone of ''Patrognathus andersoni'') * the lower zone of ''Patrognathus andersoni'' * the zone of ''Patrognathus variabilis'' * the zone of ''Patrognathus crassus'' The Visean, the second age of the Mississippian, contains four conodont biozones: * the zone of '' Lochriea nodosa'' * the zone of ''Lochriea mononodosa'' * the zone of ''Gnathodus bilineatus'' * the zone of ''Gnathodus texanus'' The Serpukhovian, the third or youngest age of the Mississippian, includes four conodont biozones: * the zone of ''Gnathodus postbilineatus'' * the zone of ''Gnathodus bollandensis'' * the zone of ''Lochriea cruciformis'' * the zone of ''Lochriea ziegleri''


Pennsylvanian (also known as Upper Carboniferous)

The
Bashkirian The Bashkirian is in the ICS geologic timescale the lowest stage or oldest age of the Pennsylvanian. The Bashkirian age lasted from to Ma, is preceded by the Serpukhovian and is followed by the Moscovian. The Bashkirian overlaps with the ...
, the oldest age of the Pennsylvanian, contains six biozones based on conodont
index fossil Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...
s: * the zone of ''
Neognathodus ''Neognathodus'' is an extinct genus of conodonts. Use in stratigraphy The Bashkirian, the oldest age of the Pennsylvanian (also known as Upper Carboniferous), contains six biozones based on conodont index fossils, two of which contain ''Neo ...
atokaensis'' * the zone of ''
Declinognathodus ''Declinognathodus'' is an extinct genus of platform conodonts. Use in stratigraphy The Bashkirian, the oldest age of the Pennsylvanian (also known as Upper Carboniferous), contains six biozones based on conodont index fossils, two of which ...
marginodosus'' * the zone of ''
Idiognathodus ''Idiognathodus'' is an extinct conodont genus in the family Idiognathodontidae. Use in stratigraphy The species ''Idiognathodus simulator'' made its first appearance during the Gzhelian, the youngest age of the Pennsylvanian (late Carbonifer ...
sinuosus'' * the zone of ''Neognathodus askynensis'' * the zone of ''
Idiognathoides ''Idiognathoides'' is an extinct genus of conodonts. Glen K. Merrill stated in 1963 that "conodont workers have considered ''Idiognathoides'' to be a junior synonym of '' Polygnathodella'' but it now proves to be a junior synonym of ''Cavusgna ...
sinuatus'' * the zone of ''Declinognathodus noduliferus'' The base of the
Moscovian Moscovian may refer to: *An inhabitant of Moscow, the capital of Russia *Something of, from, or related to Moscow * Moscovian (Carboniferous), a stage of the Carboniferous in the ICS geologic timescale