Companion plant
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Companion planting in gardening and agriculture is the planting of different crops in proximity for any of a number of different reasons, including
pest control Pest control is the regulation or management of a species defined as a pest; any animal, plant or fungus that impacts adversely on human activities or environment. The human response depends on the importance of the damage done and will range ...
,
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
, providing
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
for
beneficial insect Beneficial insects (sometimes called beneficial bugs) are any of a number of species of insects that perform valued services like pollination and pest control. The concept of ''beneficial'' is subjective and only arises in light of desired outcome ...
s, maximizing use of space, and to otherwise increase crop productivity. Companion planting is a form of
polyculture In agriculture, polyculture is the practice of growing more than one crop species in the same space, at the same time. In doing this, polyculture attempts to mimic the diversity of natural ecosystems. Polyculture is the opposite of monoculture, i ...
. Companion planting is used by farmers and gardeners in both industrialized and developing countries for many reasons. Many of the modern principles of companion planting were present many centuries ago in
forest garden Forest gardening is a low-maintenance, sustainable, plant-based food production and agroforestry system based on woodland ecosystems, incorporating fruit and nut trees, shrubs, herbs, vines and perennial vegetables which have yields directl ...
s in Asia, and thousands of years ago in
Mesoamerica Mesoamerica is a historical region and cultural area in southern North America and most of Central America. It extends from approximately central Mexico through Belize, Guatemala, El Salvador, Honduras, Nicaragua, and northern Costa Rica ...
.


History

In China, mosquito ferns (''Azolla'' spp.) have been used for at least a thousand years as companion plants for
rice Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and '' Porteresia'', both wild and domesticat ...
crops. They host a
cyanobacterium Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, blu ...
(''Anabaena azollae'') that fixes nitrogen from the atmosphere, and they block light from plants that would compete with the rice. Companion planting was practiced in various forms by the
indigenous peoples of the Americas The Indigenous peoples of the Americas are the inhabitants of the Americas before the arrival of the European settlers in the 15th century, and the ethnic groups who now identify themselves with those peoples. Many Indigenous peoples of the A ...
prior to the arrival of Europeans. These peoples domesticated
squash Squash may refer to: Sports * Squash (sport), the high-speed racquet sport also known as squash racquets * Squash (professional wrestling), an extremely one-sided match in professional wrestling * Squash tennis, a game similar to squash but pla ...
8,000 to 10,000 years ago, then
maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The ...
, then
common bean ''Phaseolus vulgaris'', the common bean, is a herbaceous annual plant grown worldwide for its edible dry seeds or green, unripe pods. Its leaf is also occasionally used as a vegetable and the straw as fodder. Its botanical classification, alo ...
s, forming the Three Sisters agricultural technique. The cornstalk served as a trellis for the beans to climb, the beans fixed nitrogen, benefitting the maize, and the wide leaves of the squash plant provide ample shade for the soil keeping it moist and fertile.


Practice

More recently, starting in the 1920s,
organic farming Organic farming, also known as ecological farming or biological farming,Labelling, article 30 o''Regulation (EU) 2018/848 of the European Parliament and of the Council of 30 May 2018 on organic production and labelling of organic products and re ...
and
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
have made frequent use of companion planting, since many other means of fertilizing, weed reduction and pest control are forbidden. The
list of companion plants This is a list of companion plants. Many more are in the list of beneficial weeds. Companion plants assist in the growth of others by attracting beneficial insects, repelling pests, or providing nutrients, shade, or support. They can be part o ...
used in such systems is large, and includes vegetables, fruit trees, kitchen herbs, garden flowers, and fodder crops. The number of interactions both positive (the pair of species assist each other) and negative (the plants are best not grown together) is also large, though the evidence for such interactions ranges from controlled experiments to hearsay. For example, plants in the cabbage family (
Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some are shrubs. The leav ...
) grow well with celery, onion family plants (''
Allium ''Allium'' is a genus of monocotyledonous flowering plants that includes hundreds of species, including the cultivated onion, garlic, scallion, shallot, leek, and chives. The generic name ''Allium'' is the Latin word for garlic,Gledhill, D ...
''), and aromatic herbs, but are best not grown with strawberry or tomato.


Mechanisms

Companion planting can operate through a variety of mechanisms, which may sometimes be combined. These include
pest control Pest control is the regulation or management of a species defined as a pest; any animal, plant or fungus that impacts adversely on human activities or environment. The human response depends on the importance of the damage done and will range ...
,
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...
, providing
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
for
beneficial insect Beneficial insects (sometimes called beneficial bugs) are any of a number of species of insects that perform valued services like pollination and pest control. The concept of ''beneficial'' is subjective and only arises in light of desired outcome ...
s, and maximizing use of space; all of these may help to increase crop productivity. excerpted from


Nutrient provision

Legumes such as
clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus ...
provide nitrogen compounds to neighbouring plants such as grasses by
fixing nitrogen Nitrogen fixation is a chemical process by which molecular nitrogen (), with a strong triple covalent bond, in the air is converted into ammonia () or related nitrogenous compounds, typically in soil or aquatic systems but also in industry. Atmos ...
from the air with symbiotic bacteria in their root nodules. These enable the grasses or other neighbours to produce more protein and hence to grow more.


Trap cropping

Trap crop A trap crop is a plant that attracts agricultural pests, usually insects, away from nearby crops. This form of companion planting can save the main crop from decimation by pests without the use of pesticides. /sup> A trap crop is used for attracti ...
ping uses alternative plants to attract pests away from a main crop. For example, nasturtium (''Tropaeolum majus'') is a food plant of some caterpillars which feed primarily on members of the cabbage family (brassicas); some gardeners claim that planting them around brassicas protects the food crops from damage, as eggs of the pests are preferentially laid on the nasturtium. However, while many trap crops divert pests from focal crops in small scale greenhouse, garden and field experiments, only a small portion of these plants reduce pest damage at larger commercial scales.


Host-finding disruption

Flying pests are far less successful if their host-plants are surrounded by other plants or even "decoy-plants" coloured green. Pests find hosts in stages, first detecting plant odours which induce it to try to land on the host plant, avoiding bare soil. If the plant is isolated, then the insect simply lands on the patch of green near the odour, making an "appropriate landing". If it finds itself on the wrong plant, an "inappropriate landing", it takes off and flies to another plant; it eventually leaves the area if there are too many "inappropriate" landings. Companion planting of
clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus ...
as ground cover was equally disruptive to eight pest species from four different insect orders. In a test, 36% of cabbage root flies laid eggs beside cabbages growing in bare soil (destroying the crop), compared to only 7% beside cabbages growing in clover (which allowed a good crop). Simple decoys of green cardboard worked just as well as the live ground cover.


Pest suppression

Some companion plants help prevent pest insects or pathogenic
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
from damaging the crop, through chemical means. For example, the smell of the foliage of marigolds is claimed to deter
aphid Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
s from feeding on neighbouring plants. A 2005 study found that oil volatiles extracted from Mexican marigold by
vacuum distillation Vacuum distillation is distillation performed under reduced pressure, which allows the purification of compounds not readily distilled at ambient pressures or simply to save time or energy. This technique separates compounds based on differences i ...
reduced the reproduction of three aphid species ( pea aphid, green peach aphid and glasshouse and potato aphid) by up to 100% after 5 days from exposure. Another example familiar to gardeners is the interaction of onions and carrots with each other's pests: the onion smell puts off carrot root fly, while the smell of carrots puts off onion fly.


Predator recruitment

Companion plants that produce copious nectar or pollen in a vegetable garden ( insectary plants) may help encourage higher populations of
beneficial insect Beneficial insects (sometimes called beneficial bugs) are any of a number of species of insects that perform valued services like pollination and pest control. The concept of ''beneficial'' is subjective and only arises in light of desired outcome ...
s that control pests.


Protective shelter

Some crops are grown under the protective shelter of different kinds of plant, whether as
wind break A windbreak (shelterbelt) is a planting usually made up of one or more rows of trees or shrubs planted in such a manner as to provide shelter from the wind and to protect soil from erosion. They are commonly planted in hedgerows around the edge ...
s or for shade. For example,
shade-grown coffee Shade-grown coffee is a form of the crop produced from coffee plants grown under a canopy of trees. A canopy of assorted types of shade trees is created to cultivate shade-grown coffee. Because it incorporates principles of natural ecology to pro ...
, especially '' Coffea arabica'', has traditionally been grown in light shade created by scattered trees with a thin canopy, allowing light through to the coffee bushes but protecting them from overheating. Suitable Asian trees include ''
Erythrina subumbrans ''Erythrina subumbrans'' is a flowering plant species in the genus ''Erythrina ''Erythrina'' is a genus of plants in the pea family, Fabaceae. It contains about 130 species, which are distributed in tropical and subtropical regions worldwide. ...
'' (tton tong or dadap), ''
Gliricidia sepium ''Gliricidia sepium'', often simply referred to as its genus name '' Gliricidia'', is a medium size leguminous tree belonging to the family Fabaceae. Common names include quickstick, ''mata ratón''; ''cacao de nance'', ''cachanance''; ''balo'' ...
'' (khae falang), ''
Cassia siamea ''Senna siamea'', also known as Siamese cassia, kassod tree, cassod tree and cassia tree, is a legume in the subfamily Caesalpinioideae. It is native to South and Southeast Asia, although its exact origin is unknown. It is a medium-size, ev ...
'' (khi lek), ''
Melia azedarach ''Melia azedarach'', commonly known as the chinaberry tree, pride of India, bead-tree, Cape lilac, syringa berrytree, Persian lilac, Indian lilac, or white cedar, is a species of deciduous tree in the mahogany family, Meliaceae, that is native ...
'' (khao dao sang), and ''
Paulownia tomentosa ''Paulownia tomentosa'', common names princess tree, empress tree, or foxglove-tree, is a deciduous hardwood tree in the family Paulowniaceae, native to central and western China. It is an extremely fast-growing tree with seeds that disperse rea ...
'', a useful timber tree.


Approaches

Companion planting approaches in use or being trialled include: * Square foot gardening attempts to protect plants from issues such as weed infestation by packing them as closely together as possible. This is facilitated by using companion plants, which can be closer together than normal. *
Forest gardening Forest gardening is a low-maintenance, sustainable, plant-based food production and agroforestry system based on woodland ecosystems, incorporating fruit and nut trees, shrubs, herbs, vines and perennial vegetables which have yields directly us ...
, where companion plants are intermingled to simulate an ecosystem, emulates the interaction of plants of up to seven different heights in a
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (se ...
.. See also


See also

*
Satoyama is a Japanese term applied to the border zone or area between mountain foothills and arable flat land. Literally, ''sato'' () means village, and ''yama'' () means hill or mountain. Satoyama have been developed through centuries of small-scale ...
*
Intercropping Intercropping is a multiple cropping practice that involves growing two or more crops in proximity. In other words, intercropping is the cultivation of two or more crops simultaneously on the same field. The most common goal of intercropping is ...
*
Ecological facilitation Ecological facilitation or probiosis describes species interactions that benefit at least one of the participants and cause harm to neither.Stachowicz, J. J. 2001. Mutualism, facilitation, and the structure of ecological communities. BioScience 51 ...
*
Vegan organic gardening Vegan organic (or veganic) agriculture is the organic production of food and other crops with minimal animal inputs. Vegan organic agriculture is the organic form of animal-free agriculture. Animal-free farming methods use no animal products ...
*
Three Sisters (agriculture) The Three Sisters are the three main agricultural crops of various Indigenous peoples of North America: squash, maize ("corn"), and climbing beans (typically tepary beans or common beans). In a technique known as companion planting, the maize ...
*
List of beneficial weeds This is a list of undomesticated or feral plants, generally considered weeds, yet having some positive effects or uses, often being ideal as companion plants in gardens. Beneficial weeds can accomplish a number of roles in the garden or yard, in ...
* List of pest-repelling plants


References

{{Permaculture Sustainable gardening Permaculture Crops Biological pest control Sustainable technologies Chemical ecology