The comet grouper (''Epinephelus morrhua''), also known as the comet cod or dot-dash grouper, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...

, a

grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

from the subfamily

Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...

which is part of the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some ca ...

, which also includes the

anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...

and sea basses. It is a species of deep coral reefs in the Indo-Pacific region.

Description

The comet grouper has a body with a standard length which is 2.8 to 3.1 times as long as it is deep. The dorsal profile of the head is moderately convex. The preopercle has enlarged serrations at its angle and it is notched just above the angle. The upper edge of the gill cover is straight. The

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

contains 11 spines and 14-15 soft rays, while the anal fin has 3 spines and 7-8 soft rays. The

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

varies from convex to slightly rounded. The head and body tan and they are marked with dark brown bands. There is a forked band starting at the posterior margin of the eye, its upper branch reaching a dark brown saddle-like blotch on the nape just in before of origin of the dorsal fin while the lower branch runs to lowest of the spines on the gill cover before continuing along the flanks and then forking over the

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

. The upper part of this band runs to a dark blotch at the base of the soft rayed part of the dorsal fin. Another dark band runs from the upper margin of the gill cover to the base of spiny part of the dorsal fin. There is also a narrow band which runs from the lower margin of the orbit to the base of the pectoral fin and then continues as a broken line along the lower part of body curving upwards on to the upper part of caudal peduncle. There is another wide band from maxillary groove to rear end of interopercle. The areas between these bands are frequently marked with small brown spots. This species attains a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of , although they a more commonly found at lengths around , and a maximum published weight of .

Distribution

The comet grouper has a wide Indo-Pacific distribution and is found from the

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...

southwards along the African coast to South Africa and east through the Indian Ocean, including the Comoros, Madagascar, Aldabra, Mauritius,

Chagos The Chagos Archipelago () or Chagos Islands (formerly the Bassas de Chagas, and later the Oil Islands) is a group of seven atolls comprising more than 60 islands in the Indian Ocean about 500 kilometres (310 mi) south of the Maldives arc ...

and Réunion, and on into the Pacific. In the

Western Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

it extends as far north as southern Japan and the

Ogasawara Islands The Bonin Islands, also known as the , are an archipelago of over 30 subtropical and tropical islands, some directly south of Tokyo, Japan and northwest of Guam. The name "Bonin Islands" comes from the Japanese word ''bunin'' (an archaic readi ...

and as far south as

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

in Australia. It reaches New Caledonia, Fiji,

Rotuma Rotuma is a Fijian dependency, consisting of Rotuma Island and nearby islets. The island group is home to a large and unique Polynesian indigenous ethnic group which constitutes a recognisable minority within the population of Fiji, known as " ...

and the

Cook Islands ) , image_map = Cook Islands on the globe (small islands magnified) (Polynesia centered).svg , capital = Avarua , coordinates = , largest_city = Avarua , official_languages = , lan ...

Habitat and biology

The comet grouper is found near

coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...

s on slopes off islands, of sea mounts or the continental shelf at depths from where it feeds on benthic fish and large

invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordat ...

. Its biology is poorly understood. This species harbours several parasite species, including the diplectanid

monogenea Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female repro ...

ns ''

Pseudorhabdosynochus morrhua ''Pseudorhabdosynochus'' is a genus of monopisthocotylean monogeneans, included in the family Diplectanidae. The type-species of the genus is ''Pseudorhabdosynochus epinepheli'' ( Yamaguti, 1938). The genus includes more than 80 species, which ...

'' and '' P. variabilis''.

Taxonomy

The comet grouper was first formally described as ''Serranus morhua'' in 1828 by the French

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...

Achille Valenciennes (1794-1865) with the type locality given as Mauritius. It is sometimes confused with its relatives '' E. poecilonotus'', '' E. radiatus'', and '' E. tuamotoensis''.

Utilisation

The comet grouper is not a common fish at market, despite being regarded as an excellent food fish, probably due to the depths at which it is found.

Ciguatoxin 300px, Chemical structure of the ciguatoxin CTX1B Ciguatoxins are a class of toxic polycyclic polyethers found in fish that cause ciguatera. There are several different chemicals in this class. "CTX" is often used as an abbreviation. * - ...

s have been found in fish caught off Mauritius.

References

External links

Fishes of Australia : ''Epinephelus morrhua''
{{Taxonbar, from=Q3236968 comet grouper Marine fish of Northern Australia comet grouper Taxa named by Achille Valenciennes