The Columbian mammoth (''Mammuthus columbi'') is an extinct species of

mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...

that inhabited the Americas as far north as the

Northern United States The Northern United States, commonly referred to as the American North, the Northern States, or simply the North, is a geographical or historical region of the United States. History Early history Before the 19th century westward expansion, the "N ...

and as far south as Costa Rica during the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus subplanifrons'' in the early

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58DNA studies show that the Columbian mammoth was a

hybrid species Hybrid speciation is a form of speciation where hybridization between two different species leads to a new species, reproductively isolated from the parent species. Previously, reproductive isolation between two species and their parents was tho ...

between woolly mammoths and another lineage descended from steppe mammoths; the hybridization happened more than 420,000 years ago. The pygmy mammoths of the

Channel Islands of California The Channel Islands () are an eight-island archipelago located within the Southern California Bight in the Pacific Ocean, off the coast of California. The four Northern Channel Islands are part of the Transverse Ranges geologic province, ...

evolved from Columbian mammoths. The closest

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

relative of the Columbian and other mammoths is the

Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus '' Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in t ...

. Reaching at the shoulders and in weight, the Columbian mammoth was one of the largest species of mammoth. It had long, curved tusks and four

molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...

, which were replaced six times during the lifetime of an individual. It most likely used its tusks and

trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...

like modern elephants—for manipulating objects, fighting, and foraging. Bones, hair, dung, and stomach contents have been discovered, but no preserved carcasses are known. The Columbian mammoth preferred open areas, such as parkland landscapes, and fed on sedges, grasses, and other plants. It did not live in the Arctic regions of Canada, which were instead inhabited by

woolly mammoth The woolly mammoth (''Mammuthus primigenius'') is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with '' Mammuthus s ...

s. The ranges of the two species may have overlapped, and genetic evidence suggests that they interbred. Several sites contain the skeletons of multiple Columbian mammoths, either because they died in incidents such as a

drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D. Jiang, A. Khan, W. Pokam Mba, D. Rosenfeld, J. Tierney, an ...

, or because these locations were natural traps in which individuals accumulated over time. For a few thousand years prior to their extinction, Columbian mammoths coexisted in North America with Paleoamericans – the first humans to inhabit the Americas – who hunted them for food, used their bones for making tools, and possibly depicted them in ancient art. Columbian mammoth remains have been found in association with

Clovis culture The Clovis culture is a prehistoric Paleoamerican culture, named for distinct stone and bone tools found in close association with Pleistocene fauna, particularly two mammoths, at Blackwater Locality No. 1 near Clovis, New Mexico, in 1936 a ...

artifacts; these remains may have stemmed either from hunting or scavenging. The Columbian mammoth disappeared at the end of the Pleistocene around 11,500 years ago, most likely as a result of habitat loss caused by

climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...

, hunting by humans, or a combination of both.

Taxonomy

The Columbian mammoth was first

scientifically described A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...

in 1857 by naturalist

Hugh Falconer Hugh Falconer MD FRS (29 February 1808 – 31 January 1865) was a Scottish geologist, botanist, palaeontologist, and paleoanthropologist. He studied the flora, fauna, and geology of India, Assam,Burma,and most of the Mediterranean islands a ...

, who named the species '' Elephas columbi'' after the explorer

Christopher Columbus Christopher Columbus * lij, Cristoffa C(or)ombo * es, link=no, Cristóbal Colón * pt, Cristóvão Colombo * ca, Cristòfor (or ) * la, Christophorus Columbus. (; born between 25 August and 31 October 1451, died 20 May 1506) was a ...

. The animal was brought to Falconer's attention in 1846 by

Charles Lyell Sir Charles Lyell, 1st Baronet, (14 November 1797 – 22 February 1875) was a Scottish geologist who demonstrated the power of known natural causes in explaining the earth's history. He is best known as the author of ''Principles of Geolo ...

, who sent him molar fragments found during the 1838 excavation of the Brunswick–Altamaha Canal in

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

, in the southeastern United States. At the time, similar fossils from across North America were attributed to woolly mammoths (then ''Elephas primigenius''). Falconer found that his specimens were distinct, confirming his conclusion by examining their internal structure and studying additional molars from Mexico. Although scientists

William Phipps Blake William Phipps Blake (June 1, 1826 – May 22, 1910) was an American geologist, mining consultant, and educator. Among his best known contributions include being the first college trained chemist to work full-time for a United States chemical m ...

and

Richard Owen Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomist and paleontologist. Owen is generally considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Ow ...

believed that ''E. texianus'' was more appropriate for the species, Falconer rejected the name; he also suggested that ''E. imperator'' and ''E. jacksoni'', two other American elephants described from molars, were based on remains too fragmentary to classify properly. More complete material that may be from the same quarry as Falconer's fragmentary

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

molar (which is cataloged as specimen BMNH 40769 at the

British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum ...

) was reported in 2012, and could help shed more light on that specimen, since doubts about its adequacy as a holotype have been raised. In the early 20th century, the taxonomy of extinct elephants became increasingly complicated. In 1942, paleontologist

Henry Fairfield Osborn Henry Fairfield Osborn, Sr. (August 8, 1857 – November 6, 1935) was an American paleontologist, geologist and eugenics advocate. He was the president of the American Museum of Natural History for 25 years and a cofounder of the American Euge ...

's posthumous monograph on the

Proboscidea The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family ( Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Fr ...

was published, wherein he used various generic and subgeneric names that had previously been proposed for extinct elephant species, such as ''Archidiskodon'', ''Metarchidiskodon'', ''Parelephas'', and ''Mammonteus''. Osborn also retained names for many regional and intermediate subspecies or "varieties", and created recombinations such as ''Parelephas columbi felicis'' and ''Archidiskodon imperator maibeni''. The taxonomic situation was simplified by various researchers from the 1970s onwards; all species of

were retained in the genus ''Mammuthus'', and many proposed differences between species were instead interpreted as

intraspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organis ...

variation. In 2003, paleontologist Larry Agenbroad reviewed opinions about North American mammoth taxonomy, and concluded that several species had been declared junior synonyms, and that ''M. columbi'' (the Columbian mammoth) and ''

M. exilis The pygmy mammoth or Channel Islands mammoth (''Mammuthus exilis'') is an extinct species of dwarf elephant descended from the Columbian mammoth (''M. columbi'') of mainland North America. This species became extinct during the Quaternary extinct ...

'' (the pygmy mammoth) were the only species of mammoth

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

to the Americas (as other species lived both there and in Eurasia). The idea that species such as ''M. imperator'' (the imperial mammoth) and ''M. jeffersoni'' ( Jefferson's mammoth) were either more primitive or advanced stages in Columbian mammoth evolution was largely dismissed, and they were regarded as synonyms. In spite of these conclusions, Agenbroad cautioned that American mammoth taxonomy is not yet fully resolved.

Evolution

The earliest known members of Proboscidea, the

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...

that contains the elephants, existed about 55 million years ago around the

Tethys Sea The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

area. The closest living relatives of the Proboscidea are the

sirenia The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct ...

ns (

dugongs The dugong (; ''Dugong dugon'') is a marine mammal. It is one of four living species of the order Sirenia, which also includes three species of manatees. It is the only living representative of the once-diverse family Dugongidae; its closest ...

and manatees) and the

hyrax Hyraxes (), also called dassies, are small, thickset, herbivorous mammals in the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Typically, they measure between long and weigh between . They are superficially simila ...

es (an order of small, herbivorous mammals). The family

Elephantidae Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are terrestrial large mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the ...

existed six million years ago in

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, and includes the living elephants and the mammoths. Among many now extinct clades, the

mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of the ...

(''Mammut'') is only a distant relative, and part of the distinct family Mammutidae, which diverged 25 million years before the mammoths evolved. The

(''Elephas maximus'') is the closest extant relative of the mammoths. The following

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

shows the placement of the Columbian mammoth among other proboscideans, based on characteristics of the

hyoid bone The hyoid bone (lingual bone or tongue-bone) () is a horseshoe-shaped bone situated in the anterior midline of the neck between the chin and the thyroid cartilage. At rest, it lies between the base of the mandible and the third cervical verteb ...

in the neck: Since many remains of each species of mammoth are known from several localities, reconstructing the evolutionary history of the genus is possible through morphological studies. Mammoth species can be identified from the number of enamel ridges (or lamellar plates) on their molars; primitive species had few ridges, and the number increased gradually as new species evolved to feed on more abrasive food items. The crowns of the teeth became taller in height and the skulls became taller to accommodate this. At the same time, the skulls became shorter from front to back to reduce the weight of the head. The short, tall skulls of woolly and Columbian mammoths are the culmination of this process. The first known members of the genus ''Mammuthus'' are the African species '' M. subplanifrons'' from the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58M. africanavus'' from the

. The former is thought to be the ancestor of later forms. Mammoths entered Europe around 3 million years ago. The earliest European mammoth has been named '' M. rumanus''; it spread across Europe and China. Only its molars are known, which show that it had 8–10 enamel ridges. A population evolved 12–14 ridges, splitting off from and replacing the earlier type, becoming '' M. meridionalis'' about 2.0–1.7 million years ago. In turn, this species was replaced by the

steppe mammoth The steppe mammoth (''Mammuthus trogontherii'', sometimes ''Mammuthus armeniacus'') is an extinct species of Elephantidae that ranged over most of northern Eurasia during the late Early and Middle Pleistocene, approximately 1.8 million-200,000 y ...

(''M. trogontherii'') with 18–20 ridges, which evolved in eastern Asia around 2.0–1.5 million years ago. The Columbian mammoth evolved from a population of ''M. trogontherii'' that had crossed the Bering Strait and entered North America about 1.5 million years ago; it retained a similar number of molar ridges. Mammoths derived from ''M. trogontherii'' evolved molars with 26 ridges 400,000 years ago in Siberia and became the woolly mammoth (''M. primigenius''). Woolly mammoths entered North America about 100,000 years ago. A population of Columbian mammoths that lived between 80,000 and 13,000 years ago on the

, away from the mainland, evolved to be less than half the size of the mainland Columbian mammoths. They are, therefore, considered to be the distinct species ''M. exilis'', the pygmy mammoth (or a subspecies, ''M. c. exilis''). These mammoths presumably reached the islands by swimming there when

sea levels Mean sea level (MSL, often shortened to sea level) is an average surface level of one or more among Earth's coastal bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical datuma standardised ...

were lower, and decreased in size due to the limited food provided by the islands' small areas. Bones of larger specimens have also been found on the islands, but whether these were stages in the dwarfing process, or later arrivals of Columbian mammoths is unknown.

Hybridization

A 2011

ancient DNA Ancient DNA (aDNA) is DNA isolated from ancient specimens. Due to degradation processes (including cross-linking, deamination and fragmentation) ancient DNA is more degraded in comparison with contemporary genetic material. Even under the bes ...

study of the complete mitochondrial genome (inherited through the female line) showed that two examined Columbian mammoths, including the morphologically typical "Huntington mammoth", were grouped within a subclade of woolly mammoths. This suggests that the two populations interbred and produced fertile offspring. One possible explanation is

introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...

of a

haplogroup A haplotype is a group of alleles in an organism that are inherited together from a single parent, and a haplogroup (haploid from the el, ἁπλοῦς, ''haploûs'', "onefold, simple" and en, group) is a group of similar haplotypes that share ...

from woolly to Columbian mammoths, or vice versa. A similar situation has been documented in modern species of African elephant (''Loxodonta''), the

African bush elephant The African bush elephant (''Loxodonta africana'') is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to and a body ...

(''L. africana'') and the African forest elephant (''L. cyclotis''). The authors of the study also suggest that the North American type formerly referred to as ''M. jeffersonii'' may have been a hybrid between the two species, as it is apparently morphologically intermediate. These findings were not expected by scientists;

nuclear DNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. I ...

and more specimens will have to be examined to clarify the situation. A 2015 study of mammoth molars confirmed that ''M. columbi'' evolved from Eurasian ''M. trogontherii'', not ''M. meridionalis'' as had been suggested earlier, and noted that ''M. columbi'' and ''M. trogontherii'' were so similar in morphology that their classification as separate species may be questionable. The study also suggested that the animals in the range where ''M. columbi'' and ''M. primigenius'' overlapped formed a

metapopulation A metapopulation consists of a group of spatially separated populations of the same species which interact at some level. The term metapopulation was coined by Richard Levins in 1969 to describe a model of population dynamics of insect pests in ...

of hybrids with varying morphology. In 2016, a genetic study of North American mammoth specimens confirmed that ''M. columbi'' and ''M. primigenius'' interbred extensively, were both descended from ''M. trogontherii'', and concluded that morphological differences between fossils may, therefore, not be reliable for determining taxonomy. The authors also questioned whether ''M. columbi'' and ''M. primigenius'' should be considered "good species", considering that they were able to interbreed after supposedly being separated for a million years, but cautioned that more specimens need to be sampled. In 2021, DNA older than a million years was sequenced for the first time, from two mammoth teeth of

Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...

age found in eastern Siberia. One tooth from Adyocha (1-1.3 million years old) belonged to a lineage that was ancestral to later woolly mammoths, whereas the other from Krestovka (1.1–1.65 million years old) belonged to new lineage, possibly a distinct species, perhaps descended from steppe mammoths that had become isolated. The study found that half of the ancestry of Columbian mammoths came from the Krestovka lineage, and the other half from woolly mammoths, with the hybridization happening more than 420,000 years ago, during the

Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. Th ...

. Later woolly and Columbian mammoths also interbred occasionally, and mammoth species perhaps hybridized routinely when brought together by glacial expansion. These findings were the first evidence of

hybrid speciation Hybrid speciation is a form of speciation where hybridization between two different species leads to a new species, reproductively isolated from the parent species. Previously, reproductive isolation between two species and their parents was thou ...

from ancient DNA. The study also found that genetic adaptations to cold environments, such as hair growth and fat deposits, were already present in the steppe mammoth lineage, and was not unique to woolly mammoths.

Description

The Columbian mammoth was about tall at the shoulder and weighed about . It was about the same size as the earlier mammoth species ''M. meridionalis'' and ''M. trogontherii'', and was larger than the modern African elephant and the woolly mammoth, both of which reached about . Males were generally larger and more robust. The best indication of sex is the size of the pelvic girdle, since the opening that functions as the birth canal is always wider in females than in males. Like other mammoths, the Columbian mammoth had a high, single-domed head and a sloping back with a high shoulder hump; this shape resulted from the

spinous process The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

es (protrusions) of the back vertebrae decreasing in length from front to rear. Juveniles, though, had convex backs like Asian elephants. Other skeletal features include a short, deep rostrum (front part of the jaws), a rounded

mandibular symphysis In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halve ...

(where the two halves of the lower jaw connected) and the

coronoid process of the mandible In human anatomy, the mandible's coronoid process (from Greek ''korōnē'', denoting something hooked) is a thin, triangular eminence, which is flattened from side to side and varies in shape and size. Its anterior border is convex and is continuou ...

(upper protrusion of the jaw bone) extending above the molar surfaces. Archidiskodon imperator121

Apart from its larger size and more primitive molars, the Columbian mammoth also differed from the woolly mammoth by its more downturned mandibular symphysis; the dental alveoli (tooth sockets) of the tusks were directed more laterally away from the midline. Its tail was intermediate in length between that of modern elephants and the woolly mammoth. Since no Columbian mammoth soft tissue has been found, much less is known about its appearance than that of the woolly mammoth. It lived in warmer habitats than the woolly mammoth, and probably lacked many of the adaptations seen in that species. Hair thought to be that of the Columbian mammoth has been discovered in Bechan Cave in Utah, where mammoth dung has also been found. Some of this hair is coarse, and identical to that known to belong to woolly mammoths; however, since this location is so far south, it is unlikely to be woolly mammoth hair. The distribution and density of fur on the living animal is unknown, but it was probably less dense than that of the woolly mammoth due to the warmer habitat. An additional tuft of Columbian mammoth hair is known from near Castroville in California, the hair was noted to be red-orange and was described as being similar in colour to a

Golden Retriever The Golden Retriever is a Scottish breed of retriever dog of medium size. It is characterised by a gentle and affectionate nature and a striking golden coat. It is commonly kept as a pet and is among the most frequently registered breeds ...

Dentition

Columbian mammoths had very long tusks (modified

incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...

teeth), which were more curved than those of modern elephants. The largest known mammoth tusk, long, belonged to a Columbian mammoth, and others range from long. Columbian mammoth tusks were usually not much larger than those of woolly mammoths, which reached . The tusks of females were much smaller and thinner. About a quarter of the tusks' length was inside the sockets; they grew spirally in opposite directions from the base, curving until the tips pointed towards each other, and sometimes crossed. Most of their weight would have been close to the skull, with less

torque In physics and mechanics, torque is the rotational equivalent of linear force. It is also referred to as the moment of force (also abbreviated to moment). It represents the capability of a force to produce change in the rotational motion of th ...

than straight tusks would have generated. The tusks were usually asymmetrical, with considerable variation; some tusks curved down, instead of outwards, or were shorter due to breakage. Columbian mammoth tusks were generally less twisted than those of woolly mammoths. At six months of age, calves developed milk tusks a few centimeters long, which were replaced by permanent tusks a year later. Annual tusk growth of continued throughout life, slowing as the animal reached adulthood. Columbian mammoths had four functional molar teeth at a time, two in the upper jaw and two in the lower. About of the crown was within the jaw, and was above. The crowns of the lower jaw were pushed forward and up as they wore down, comparable to a

conveyor belt A conveyor belt is the carrying medium of a belt conveyor system (often shortened to belt conveyor). A belt conveyor system is one of many types of conveyor systems. A belt conveyor system consists of two or more pulleys (sometimes referred t ...

. The teeth had separated ridges of enamel, which were covered in "prisms" directed towards the chewing surface. Wear-resistant, they were held together with cementum and

dentin Dentin () (American English) or dentine ( or ) (British English) ( la, substantia eburnea) is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by e ...

. A mammoth's molars were replaced five times over the animal's lifetime. The first molars were about the size of those of a human, ; the third ones were long, and the sixth ones were about long and weighed . With each replacement, the molars grew larger and gained more ridges; the number of plates varied between individuals. Growing of ridge took about 10.6 years.

Paleobiology

Like that of modern elephants, the mammoth's sensitive, muscular trunk was a limb-like organ with many functions. It was used for manipulating objects and social interaction. Although healthy adult mammoths could defend themselves from predators with their tusks, trunks, and size, juveniles and weakened adults were vulnerable to pack hunters such as wolves and

big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. Despite enormous differences in size, various cat species are quite similar ...

s. Bones of juvenile Columbian mammoths, accumulated by ''

Homotherium ''Homotherium'', also known as the scimitar-toothed cat or scimitar cat, is an extinct genus of machairodontine saber-toothed predator, often termed scimitar-toothed cats, that inhabited North America, South America, Eurasia, and Africa during ...

'' (the scimitar-toothed cat), have been found in Friesenhahn Cave in

Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...

. Tusks may have been used in intraspecies fighting for territory or mates and for display, to attract females and intimidate rivals. Two Columbian mammoths that died in Nebraska with tusks interlocked provide evidence of fighting behavior. The mammoths could use their tusks as weapons by thrusting, swiping, or crashing them down, and used them in pushing contests by interlocking them, which sometimes resulted in breakage. The tusks' curvature made them unsuitable for stabbing. Although to what extent Columbian mammoths migrated is unclear, an

isotope analysis Isotope analysis is the identification of isotopic signature, abundance of certain stable isotopes of chemical elements within organic and inorganic compounds. Isotopic analysis can be used to understand the flow of energy through a food w ...

Blackwater Draw Blackwater Draw is an intermittent stream channel about long, with headwaters in Roosevelt County, New Mexico, about southwest of Clovis, New Mexico, and flows southeastward across the Llano Estacado toward the city of Lubbock, Texas, where i ...

in New Mexico indicated that they spent part of the year in the Rocky Mountains, away. The study of tusk rings may aid further study of mammoth migration. On

Goat Rock Beach Goat Rock Beach is a sand beach in northwestern Sonoma County, California, United States. This landform is a sub-unit of Sonoma Coast State Beach, owned and managed by the State of California. At the northern terminus of Goat Rock Beach is the mo ...

Sonoma Coast State Park Sonoma Coast State Park is a State of California property in Sonoma County consisting of public access use on lands adjoining the Pacific Ocean. This extent of beach runs from a coastal point about north of Jenner and continues for approximat ...

blueschist Blueschist (), also called glaucophane schist, is a metavolcanic rock that forms by the metamorphism of basalt and rocks with similar composition at high pressures and low temperatures (), approximately corresponding to a depth of . The blue ...

and

chert Chert () is a hard, fine-grained sedimentary rock composed of microcrystalline or cryptocrystalline quartz, the mineral form of silicon dioxide (SiO2). Chert is characteristically of biological origin, but may also occur inorganically as a ...

outcrops (nicknamed "Mammoth Rocks") show evidence of having been rubbed by Columbian mammoths or mastodons. The rocks have polished areas above the ground, primarily near their edges, and are similar to African rubbing rocks used by elephants and other herbivores to rid themselves of mud and parasites. Similar rocks exist in Hueco Tanks, Texas, and on Cornudas Mountain in New Mexico.

Social behavior

Like modern elephants, Columbian mammoths were probably social and lived in

matriarchal Matriarchy is a social system in which women hold the primary power positions in roles of authority. In a broader sense it can also extend to moral authority, social privilege and control of property. While those definitions apply in general En ...

(female-led) family groups; most of their other social behavior was also similar to that of modern elephants. This is supported by fossil assemblages such as the

Dent site The Dent site is a Clovis culture (about 11,000 years before present) site located in Weld County, Colorado, near Milliken, Colorado. It provided evidence that humans and mammoths co-existed in the Americas. The site is located on an alluvial f ...

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the ...

and the Waco Mammoth National Monument in

Waco, Texas Waco ( ) is the county seat of McLennan County, Texas, United States. It is situated along the Brazos River and I-35, halfway between Dallas and Austin. The city had a 2020 population of 138,486, making it the 22nd-most populous city in the s ...

, where groups consisting entirely of female and juvenile Columbian mammoths have been found (implying female-led family groups). The latter assemblage includes 22 skeletons, with 15 individuals representing a herd of females and juveniles that died in a single event. The herd was originally proposed to have been killed by a

flash flood A flash flood is a rapid flooding of low-lying areas: washes, rivers, dry lakes and depressions. It may be caused by heavy rain associated with a severe thunderstorm, hurricane, or tropical storm, or by meltwater from ice or snow flowing ...

, and the arrangement of some of the skeletons suggests that the females may have formed a defensive ring around the juveniles. In 2016, the herd was suggested to have died by

near a diminishing watering hole;

scavenging Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding ...

traces on the bones contradict rapid burial, and the absence of calves and the large diversity of other animal species found gathered at the site support this scenario. Another group, consisting of a bull and six females, was found at the same site; although both groups died between 64,000 and 73,000 years ago, whether they died in the same event is unknown. At the Murray Springs Clovis Site in Arizona, where several Columbian mammoth skeletons have been excavated, a

trackway Historic roads (historic trails in USA and Canada) are paths or routes that have historical importance due to their use over a period of time. Examples exist from prehistoric times until the early 20th century. They include ancient trackways ...

similar to that left by modern elephants leads to one of the skeletons. The mammoth may have made the trackway before it died, or another individual may have approached the dead or dying animal—similar to the way modern elephants guard dead relatives for several days. Accumulations of modern elephant remains have been called " elephants' graveyards", because these sites were erroneously thought to be where old elephants went to die. Similar accumulations of mammoth bones have been found; these are thought to be the result of individuals dying near or in rivers over thousands of years and their bones being accumulated by the water (such as in the

Aucilla River The Aucilla River rises in Brooks County, Georgia, USA, close to Thomasville, and passes through the Big Bend region of Florida, emptying into the Gulf of Mexico at Apalachee Bay. Some early maps called it the Ocilla River. It is long and h ...

in Florida), or animals dying after becoming mired in mud. Some accumulations are thought to be the remains of herds that died at the same time, perhaps due to flooding. Columbian mammoths are occasionally preserved in

volcanic A volcano is a rupture in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface. On Earth, volcanoes are most often found where tectonic plat ...

deposits such as those in Tocuila, Texcoco, Mexico, where a volcanic

lahar A lahar (, from jv, ꦮ꧀ꦭꦲꦂ) is a violent type of mudflow or debris flow composed of a slurry of pyroclastic material, rocky debris and water. The material flows down from a volcano, typically along a river valley. Lahars are extr ...

mudflow A mudflow or mud flow is a form of mass wasting involving fast-moving flow of debris that has become liquified by the addition of water. Such flows can move at speeds ranging from 3 meters/minute to 5 meters/second. Mudflows contain a significa ...

covered at least seven individuals 12,500 years ago. How many mammoths lived at one location at a time is unknown, but the number likely varied by season and lifecycle. Modern elephants can form large herds, sometimes consisting of multiple family groups, and these herds can include thousands of animals migrating together. Mammoths may have formed large herds more often than modern elephants, since animals living in open areas are more likely to do this than those in forested areas.

Natural traps

Many specimens also accumulated in natural traps, such as

sinkhole A sinkhole is a depression or hole in the ground caused by some form of collapse of the surface layer. The term is sometimes used to refer to doline, enclosed depressions that are locally also known as ''vrtače'' and shakeholes, and to openi ...

s and

tar pit Tar pits, sometimes referred to as asphalt pits, are large asphalt deposits. They form in the presence of oil, which is created when decayed organic matter is subjected to pressure underground. If this crude oil seeps upward via fractures, cond ...

The Mammoth Site The Mammoth Site is a museum and paleontological site near Hot Springs, South Dakota, in the Black Hills. It is an active paleontological excavation site at which research and excavations are continuing. The facility encloses a prehistoric sink ...

Hot Springs A hot spring, hydrothermal spring, or geothermal spring is a spring produced by the emergence of geothermally heated groundwater onto the surface of the Earth. The groundwater is heated either by shallow bodies of magma (molten rock) or by circ ...

South Dakota South Dakota (; Sioux: , ) is a U.S. state in the North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota and Dakota Sioux Native American tribes, who comprise a large po ...

, is a 26,000-year-old, roughly -long sinkhole that functioned for 300 to 700 years before filling with

sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand ...

. The site is the opposite scenario of that in Waco; all but one of the at least 55 skeletons—additional skeletons are excavated each year—are male, and accumulated over time rather than in a single event. Like modern male elephants, male mammoths primarily are assumed to have lived alone, to be more adventurous (especially young males), and to be more likely to encounter dangerous situations than the females. The mammoths may have been lured to the hole by warm water or vegetation near the edges, slipping in and drowning or starving. Isotope studies of growth rings have shown that most of the mammoths died during spring and summer, which may have correlated with vegetation near the sinkhole. One individual, nicknamed "Murray", lies on its side, and probably died in this pose while struggling to get free. Deep footprints of mammoths attempting to free themselves from the sinkhole's mud can be seen in vertically excavated sections of the site. Smilodon and Canis dirus

Since the early 20th century, excavations at the

La Brea Tar Pits La Brea Tar Pits is an active paleontological research site in urban Los Angeles. Hancock Park was formed around a group of tar pits where natural asphalt (also called asphaltum, bitumen, or pitch; ''brea'' in Spanish) has seeped up from the gr ...

Los Angeles Los Angeles ( ; es, Los Ángeles, link=no , ), often referred to by its initials L.A., is the largest city in the state of California and the second most populous city in the United States after New York City, as well as one of the world ...

have yielded of fossils from 600 species of flora and fauna, including several Columbian mammoths. Many of the fossils are the remains of animals that became stuck in

asphalt Asphalt, also known as bitumen (, ), is a sticky, black, highly viscous liquid or semi-solid form of petroleum. It may be found in natural deposits or may be a refined product, and is classed as a pitch. Before the 20th century, the term ...

puddles that seeped to the surface of the pits, 40,000 to 11,500 years ago. Dust and leaves likely concealed the liquid asphalt, which then trapped unwary animals. Mired animals died from hunger or exhaustion; their corpses attracted predators, which sometimes became stuck, themselves. The fossil record of the tar pits is dominated by the remains of predators, such as large

canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...

s and

felid Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the dom ...

s. Fossils of different animals are found stuck together when they are excavated from the pits. The tar pits do not preserve soft tissue, and a 2014 study concluded that asphalt may degrade the DNA of animals mired in it after an attempt was made to extract DNA from a Columbian mammoth. A site in at airport construction area in Mexico nicknamed " mammoth central" is believed to have been the boggy shores of an ancient lake bed where animals were trapped 10,0000 to 20,000 years ago. Human tools have been found at the site. It remains unclear whether the 200 Columbian mammoths found there died of natural causes and were then carved by humans. Some have hypothesized that humans drove the Columbian mammoths into the area to kill them. The site is only from artificial pits which were once used by humans to trap and kill large mammals.

Diet

An adult Columbian mammoth would have needed more than of food per day, and may have foraged for 20 hours a day. Mammoths chewed their food using their powerful jaw muscles to move the mandible forward and close the mouth, then backward while opening; the sharp enamel ridges thereby cut across each other, grinding the food. The ridges were wear-resistant, enabling the animal to chew large quantities of food that contained grit. The trunk could be used for pulling up large tufts of grass, picking buds and flowers, or tearing leaves and branches from trees and shrubs, and the tusks were used to dig up plants and strip bark from trees. Digging is indicated on preserved tusks by flat, polished sections of the surface that would have reached the ground. Isotope studies of Columbian mammoths from Mexico and the United States have shown that their diet varied by location, consisting of a mix of C3 (most plants) and C4 plants (such as grasses), and they were not restricted to

grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and ot ...

browsing Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...

. Stomach contents from Columbian mammoths are rare, since no carcasses have been found, but plant remains were discovered between the pelvis and ribs of the "Huntington mammoth" when it was excavated in Utah.

Microscopy Microscopy is the technical field of using microscopes to view objects and areas of objects that cannot be seen with the naked eye (objects that are not within the resolution range of the normal eye). There are three well-known branches of micr ...

showed that these chewed remains consisted of

sedge The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus '' Carex'' ...

s, grasses, fir twigs and needles, oak, and

maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since h ...

. A large amount of mammoth dung has been found in two caves in Utah. The dry conditions and stable temperature of Bechan Cave (''bechan'' is

Navajo The Navajo (; British English: Navaho; nv, Diné or ') are a Native Americans in the United States, Native American people of the Southwestern United States. With more than 399,494 enrolled tribal members , the Navajo Nation is the largest fe ...

for "large faeces") has preserved 16,000- to 13,500-year-old elephant dung, most likely from Columbian mammoths. The dung consists of 95% grasses and sedges, and varies from 0 to 25% woody plants between dung boluses, including

saltbush Saltbush is a vernacular plant name that most often refers to ''Atriplex'', a genus of about 250 plants distributed worldwide from subtropical to subarctic regions. ''Atriplex'' species are native to Australia, North and South America, and Eurasia. ...

sagebrush Sagebrush is the common name of several woody and herbaceous species of plants in the genus '' Artemisia''. The best known sagebrush is the shrub '' Artemisia tridentata''. Sagebrushes are native to the North American west. Following is an al ...

, water birch, and blue spruce. This is similar to the diet documented for the woolly mammoth, although browsing seems to have been more important for the Columbian mammoth. The cover of dung is thick, and has a volume of 227 m³ (8,000 cubic ft), with the largest boluses in diameter. The Bechan dung could have been produced by a small group of mammoths over a relatively short time, since adult African elephants drop an average of of dung every two hours and each day. Giant North American fruits of plants such as the Osage-orange,

Kentucky coffeetree The Kentucky coffeetree (''Gymnocladus dioicus''), also known as American coffee berry, Kentucky mahogany, nicker tree, and stump tree, is a tree in the subfamily Caesalpinioideae of the legume family Fabaceae, native to the Midwest, Upper South, ...

, pawpaw and

honey locust The honey locust (''Gleditsia triacanthos''), also known as the thorny locust or thorny honeylocust, is a deciduous tree in the family Fabaceae, native to central North America where it is mostly found in the moist soil of river valleys. Hon ...

have been proposed to have evolved in tandem with now-extinct American megafauna such as mammoths and other proboscideans, since no extant endemic herbivores are able to ingest these fruits and disperse their seeds. Introduced cattle and horses have since taken over this ecological role.

Life history

The lifespan of the Columbian mammoth is thought to have been about 80 years. The lifespan of a mammal is related to its size; Columbian mammoths are larger than modern elephants, which have a lifespan of about 60 years. The age of a mammoth can be roughly determined by counting the

growth rings Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and at ...

of its tusks when viewed in cross section. However, ring-counting does not account for a mammoth's early years; early growth is represented in tusk tips, which are usually worn away. In the remainder of the tusk, each major line represents a year, with weekly and daily lines found in between. Dark bands correspond to summer, making determining the season in which a mammoth died possible. Tusk growth slowed when foraging became more difficult, such as during illness or when a male mammoth was banished from the herd (male elephants live with their herds until about the age of 10). Mammoths continued growing during adulthood, as do other elephants. Males grew until age 40, and females until age 25. Mammoths may have had gestation periods of 21–22 months, like those of modern elephants. Columbian mammoths had six sets of molars in the course of a lifetime. At 6–12 months, the second set of molars would erupt, with the first set worn out at 18 months of age. The third set of molars lasted for 10 years, and this process was repeated until the sixth set emerged at 30 years of age. When the last set of molars wore out, the animal would be unable to chew, and would die of starvation. Almost all vertebrae of the "Huntington mammoth", a very aged specimen, were deformed by arthritic disease, and four of its lumbar vertebrae were fused; some bones also indicate bacterial infection, such as

osteomyelitis Osteomyelitis (OM) is an infection of bone. Symptoms may include pain in a specific bone with overlying redness, fever, and weakness. The long bones of the arms and legs are most commonly involved in children e.g. the femur and humerus, while the ...

. The condition of the bones suggests the specimen died of old age and malnutrition.

Distribution and habitat

Columbian mammoths inhabited the southern half of North America, ranging from the northern United States across Mexico as far south as

Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...

. One Costa Rican specimen, a molar, was reported in 1963, but has since been lost. The environment in these areas may have had more varied habitats than those inhabited by woolly mammoths in the north (the

mammoth steppe During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was the Earth's most extensive biome. It spanned from Spain eastward across Eurasia to Canada and from the arctic islands southward to China. The mammoth step ...

). Some areas were covered by grasses, herbaceous plants, trees, and shrubs; their composition varied from region to region, and included

grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...

savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground ...

, and

aspen parkland Aspen parkland refers to a very large area of transitional biome between prairie and boreal forest in two sections, namely the Peace River Country of northwestern Alberta crossing the border into British Columbia, and a much larger area stretchi ...

habitats. Wooded areas also occurred; although mammoths would not have preferred forests, clearings in them could provide the animals with grasses and herbs. The Columbian mammoth shared its habitat with other now-extinct Pleistocene mammals such as ''

Glyptotherium ''Glyptotherium'' (from Greek for 'grooved or carved beast') is a genus of glyptodont (an extinct group of large, herbivorous armadillos) that lived from the Early Pliocene, about 4.9 million years ago, to the Early Holocene, around 7,000 yea ...

'', ''

Smilodon ''Smilodon'' is a genus of the extinct machairodont subfamily of the felids. It is one of the most famous prehistoric mammals and the best known saber-toothed cat. Although commonly known as the saber-toothed tiger, it was not closely rela ...

'',

ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribb ...

s, ''

Camelops ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...

'', mastodons, horses, and bison. It did not live in Arctic Canada or Alaska, which was inhabited by woolly mammoths. Fossils of woolly and Columbian mammoths have been found in the same place in a few areas of North America where their ranges overlapped, including the Hot Springs Site. Whether the two species were

sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species s ...

and lived there simultaneously, or if the woolly mammoths entered southern areas when Columbian mammoth populations were absent is unknown. The Columbian mammoth coexisted with the other extinct proboscideans '' Stegomastodon mirificus'' and '' Cuvieronius tropicus'' at sites in Texas and New Mexico during the early

Irvingtonian The Irvingtonian North American Land Mammal Age on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), spanning from 1.9 million – 250,000 years BP.

Relationship with humans

Humans entered the Americas through the

Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...

landbridge, and evidence documents their interactions with Columbian mammoths. Tools made from Columbian mammoth remains have been discovered in several North American sites. At Tocuila, Mexico, mammoth bones were quarried 13,000 years ago to produce

lithic flake In archaeology, a lithic flake is a "portion of rock removed from an objective piece by percussion or pressure,"Andrefsky, W. (2005) ''Lithics: Macroscopic Approaches to Analysis''. 2d Ed. Cambridge, Cambridge University Press and may also be ref ...

s and cores. At the Lange-Ferguson Site in South Dakota, the remains of two mammoths were found with two 12,800-year-old cleaver choppers made from a mammoth shoulder blade; the choppers had been used to butcher the mammoths. At the same site, a flake knife made from a long mammoth bone was also found wedged against mammoth vertebrae. At Murray Springs, archeologists discovered a 13,100-year-old object made from a mammoth femur; the object is thought to be a shaft wrench, a tool for straightening wood and bone to make spear-shafts (the

Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...

use similar tools). The earliest suggested evidence of Columbian mammoth butchering in America (from Lovewell, Kansas) dates from 18,000 to 21,000 years ago. Paleoamericans of the

, which arose roughly 7,000 years later, may have been the first humans to hunt mammoths extensively. These people are thought to have hunted Columbian mammoths with Clovis pointed spears which were thrown or thrust. Although Clovis points have been found with Columbian mammoth remains at several sites, archeologists disagree about whether the finds represent hunting, scavenging dead mammoths, or are coincidental. A female mammoth at the

Naco-Mammoth Kill Site The Naco Mammoth Kill Site is an archaeological site in southeast Arizona, 1 mile northwest of Naco in Cochise County. The site was reported to the Arizona State Museum in September 1951 by Marc Navarrete, a local resident, after his father fou ...

in Arizona, found with eight Clovis points near its skull, shoulder blade, ribs, and other bones, is considered the most convincing evidence for hunting. In modern experiments, replica spears have been able to penetrate the rib cages of African elephants with reuse causing little damage to the points. Other sites show more circumstantial evidence of mammoth hunting, such as piled bones bearing butcher marks. Some of these sites are not closely associated with Clovis points. The Dent site (the first evidence of mammoth hunting in North America, discovered in 1932) and the

Lehner Mammoth-Kill Site The Lehner Mammoth-Kill Site is a location in southern Arizona that is significant for its association with evidence that mammoths were killed here by Paleo-Indians 9000 years BCE. In 1952, Ed Lehner discovered extinct mammoth bone fragments o ...

, where multiple juvenile and adult mammoths have been found with butcher marks and in association with Clovis points, were once interpreted as the killing of entire herds by Clovis hunters. However, isotope studies have shown that the accumulations represent individual deaths at different seasons of the year, so are not herds killed in single incidents. Many other such assemblages of bones with butcher marks may also represent accumulations over time, so are ambiguous as evidence for large-scale hunting. A 2021 article by the American paleontologist Metin I. Eren and colleagues suggested mammoths were not very susceptible to Clovis point weapons due to their thick skin, hair, muscles, ribs, and fat, which would have impeded most types of attacks humans could pull off at that time. Experiments wherein most spear points used to calculate their effectiveness against simulated mammoth skin shattered on impact rather than penetrating, suggested to these researchers that ancient humans probably preferred to scavenge mammoth carcasses for their meat and other resources and threw spears to drive other scavengers away from carcasses before butchering the corpse with their stone tools. While the study does not rule out the hunting of mammoths by early humans, it instead indicates that such an event was probably rare and potentially more dangerous and less rewarding than scavenging. In response, other scientists found no reason to abandon the traditional idea that Clovis points were used to hunt big-game, one suggesting that such spears could have been thrown or thrust at areas of the torso that were not protected by ribs, with the wounds not killing the mammoths instantly, but the hunters could follow their prey until it had bled to death.

Petroglyph A petroglyph is an image created by removing part of a rock surface by incising, picking, carving, or abrading, as a form of rock art. Outside North America, scholars often use terms such as "carving", "engraving", or other descriptions ...

s in the

Colorado Plateau The Colorado Plateau, also known as the Colorado Plateau Province, is a physiographic and desert region of the Intermontane Plateaus, roughly centered on the Four Corners region of the southwestern United States. This province covers an area o ...

have been interpreted as depictions of either Columbian mammoths or mastodons. A bone fragment from

Vero Beach Vero Beach is a city in and the seat of Indian River County, Florida, United States. Vero Beach is the second most populous city in Indian River County. Abundant in beaches and wildlife, Vero Beach is located on Florida's Treasure Coast. It is thi ...

, Florida, estimated to be 13,000-years old and possibly the earliest known example of art in the Americas, is engraved with either a mammoth or a mastodon. While the authenticity of this depiction is based on continuity of mineralisation across the markings, other possible indicators are inconclusive at present. Petroglyphs from the San Juan River in Utah have been suggested to be 11,000–13,000-years old and to include depictions of two Columbian mammoths; the mammoths' domed heads distinguish them from mastodons. They are also shown with two "fingers" on their trunks, a feature known from European depictions of mammoths. The tusks are short, which may indicate they are meant to be females. A carving of a bison (possibly the extinct ''

Bison antiquus ''Bison antiquus'', the antique bison or ancient bison, is an extinct species of bison that lived in Late Pleistocene North America until around 10,000 years ago. It was one of the most common large herbivores on the North American continent dur ...

'') is superimposed on one of the mammoth carvings and may be a later addition. Geological dating of the San Juan River depictions in 2013 have shown them to be less than 4000 years old, after mammoths and mastodons went extinct, and they may instead be an arrangement of unrelated elements. Other possible depictions of Columbian mammoths have been shown to be either misinterpretations or fraudulent. The Columbian mammoth is the

state fossil Most American states have made a state fossil designation, in many cases during the 1980s. It is common to designate one species in which fossilization has occurred, rather than a single specimen, or a category of fossils not limited to a single ...

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

and

South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...

Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the sout ...

's state fossil is "Archie", a Columbian mammoth specimen found in the state in 1922. "Archie" is currently on display at Elephant Hall in

Lincoln, Nebraska Lincoln is the capital city of the U.S. state of Nebraska and the county seat of Lancaster County. The city covers with a population of 292,657 in 2021. It is the second-most populous city in Nebraska and the 73rd-largest in the United ...

, and is the largest mounted mammoth specimen in the United States.

Extinction

Columbian and woolly mammoths both disappeared during the late Pleistocene and early

Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...

, alongside most of the

Pleistocene megafauna Pleistocene megafauna is the set of large animals that lived on Earth during the Pleistocene epoch. Pleistocene megafauna became extinct during the Quaternary extinction event resulting in substantial changes to ecosystems globally. The role of ...

. The most recent Columbian mammoth remains have been dated around 10,900 years ago. This extinction formed part of the

Quaternary extinction event The Quaternary period (from 2.588 ± 0.005 million years ago to the present) has seen the extinctions of numerous predominantly megafaunal species, which have resulted in a collapse in faunal density and diversity and the extinction of key ecolog ...

, which began 40,000 years ago and peaked between 14,000 and 11,500 years ago. Scientists do not know whether these extinctions happened abruptly or were drawn out. During this period, 40 mammal species disappeared from North America, almost all of which weighed over ; the extinction of the mammoths cannot be explained in isolation. Scientists are divided over whether climate change, hunting, or a combination of the two, drove the extinction of the Columbian mammoths. According to the climate-change hypothesis, warmer weather led to the shrinking of suitable habitat for Columbian mammoths, which turned from parkland to forest, grassland, and semidesert, with less diverse vegetation. The "overkill hypothesis" attributes the extinction to hunting by humans, an idea first proposed by geoscientist Paul S. Martin in 1967; more recent research on this subject has varied in conclusions. A 2002 study concluded that the archeological record did not support the "overkill hypothesis", given that only 14 Clovis sites (12 with mammoth remains and two with mastodon remains) out of 76 examined provided strong evidence of hunting. In contrast, a 2007 study found that the Clovis record indicated the highest frequency of prehistoric exploitation of proboscideans for subsistence in the world, and supported the "overkill hypothesis". A 2019 study that used mathematical modelling to simulate correlations between migrations of humans and Columbian mammoths also supported the "overkill hypothesis". Whatever the actual cause of extinction, large mammals are generally more susceptible to hunting pressure than smaller ones due to their smaller population size and low reproduction rates. On the other hand, large mammals are generally less vulnerable to climatic stresses since they have greater fat deposits at their disposal and can migrate long distances to escape food shortages.

References

External links

* *
Mammoth Mystery
–

Nova A nova (plural novae or novas) is a transient astronomical event that causes the sudden appearance of a bright, apparently "new" star (hence the name "nova", which is Latin for "new") that slowly fades over weeks or months. Causes of the dramat ...

documentary about the fighting mammoths of Nebraska
Footage showing removal of a Columbian mammoth skull at the Hot Springs Mammoth Site
{{Taxonbar, from=Q1698713 Mammoths Pleistocene proboscideans Mammoth Columbian Holocene extinctions Mammoth Columbian Fossil taxa described in 1857 Taxa named by Hugh Falconer Pleistocene mammals of North America