biological taxonomy In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...

, circumscription is the content of a

taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...

, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or

Botanical Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...

Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in

molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

. New evidence may suggest that a traditional circumscription should be revised, particularly if the old circumscription is shown to be

paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ...

(a group containing some but not all of the descendants of the common ancestor). For example, the family Pongidae contained orangutans ('' Pongo''), chimpanzees ('' Pan'') and gorillas (''

Gorilla Gorillas are herbivorous, predominantly ground-dwelling great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four ...

''), but not humans (''

Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely rela ...

''), which are placed in

Hominidae The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the ...

. Once molecular phylogenetic data showed that

chimpanzee The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative t ...

s were more closely related to

human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, cultu ...

s than to

gorilla Gorillas are herbivorous, predominantly ground-dwelling great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four ...

s or

orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genu ...

s, it became clear that Pongidae is a paraphyletic group, and the circumscription of Hominidae was changed to include all four extant genera of great apes. Sometimes, systematists propose circumscriptions that do not rectify problems with paraphyly. For example, the broadly circumscribed moth family Pyralidae (or superfamily Pyraloidea) can be split into two families, Pyralidae and Crambidae, which are reciprocally monophyletic sister taxa. An example of a botanical group with unstable circumscription is Anacardiaceae, a

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...

s. Some experts favor a circumscription in which this family includes the

Blepharocaryaceae Anacardioideae is a subfamily of plants in the family Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of t ...

Julianaceae Anacardioideae is a subfamily of plants in the family Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of t ...

, and

Podoaceae Anacardioideae is a subfamily of plants in the family Anacardiaceae The Anacardiaceae, commonly known as the cashew family or sumac family, are a family of flowering plants, including about 83 genera with about 860 known species. Members of t ...

, which are sometimes considered to be separate families.Stevens, P. F. (2001 onwards)
Angiosperm Phylogeny Website
Version 9, June 2008 nd more or less continuously updated since

References

{{reflist Logic Taxonomy (biology)

See also

References