Chitra indica
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The Indian narrow-headed softshell turtle (''Chitra indica''), also known as the small-headed softshell turtle or the Indo-Gangetic softshell turtle, is an
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
species of
softshell turtle The Trionychidae are a taxonomic family of a number of turtle genera, commonly known as softshell turtles. The family was erected by Leopold Fitzinger in 1826. Softshells include some of the world's largest freshwater turtles, though many can a ...
native to waterways and rivers of the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, In ...
. It is very large (being one of the largest freshwater turtles), feeding on fish, frogs, worms,
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s and
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estim ...
, and even the occasional swimming small
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
or other mammal. ''C. indica'', like other softshell turtles, uses it flexible (and somewhat leathery) shell to dig itself deep into sandy lake and river bottoms; here, it patiently waits (with just its nose, mouth and eyes exposed) for potential prey to swim by. They will also ambush and chase their prey, depending on availability, the time of year, and size of the prey.Das, I.; and Singh, S. (2009).
Chitra indica.
' Conservation Biology of Freshwater Turtles and Tortoises, Chelonian Research Monographs 5
In the past it was included as a subspecies of ''
Chitra chitra The Asian narrow-headed softshell turtle (''Chitra chitra'') is a large species of softshell turtle in the family Trionychidae. The species is endemic to Southeast Asia. Common names ''Chitra chitra'' is also known commonly as Nutaphand's n ...
'', a species restricted to
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
using current taxonomy.


Diagnostic characters

This species is very large with a carapace up to . # Overall color: olive to deep olive-green; # Very complex midline (vertebral) carapacial pattern; # Midline (vertebral) carapacial stripe present; # Complex radiating
costal Costal may refer to: * an adjective related to the rib () in anatomy ** Costal cartilage, a type of cartilage forming bars which serve to prolong the ribs forward ** Costal margin, the medial margin formed by the false ribs ** Costal surface (disa ...
stripes; # Paramedian neck stripes forming a bell-like pattern on
anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Th ...
carapace absent; # Neck stripes do not form a continuous light rim around carapace; # No distinct pair of neck stripes; # Dark speckling on "light" (head and neck) stripes; # Anterior neck "V" divergence point on neck; # 3-4 forelimb lamellae; # No peri-orbital X pattern; # No distinct peri-orbital
ocelli A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l ...
; # No distinct naso-orbital triangular "figure"; # Few, if any, black dots present on chin pattern. Type locality: "India, fl. Ganges, Penang"; restricted by Smith (1931:162) to "
Fatehgarh Fatehgarh is a cantonment town in Farrukhabad district in the state of Uttar Pradesh, India. It is located on the south bank of the Ganges River. It is the administrative headquarters of Farrukhabad District. Fatehgarh derives its name from ...
, Ganges," India = "India: Ganges; Futtaghur" (Gray 1864: 92)


Distribution

This species is found in the
Sutlej The Sutlej or Satluj River () is the longest of the five rivers that flow through the historic crossroads region of Punjab in northern India and Pakistan. The Sutlej River is also known as ''Satadru''. It is the easternmost tributary of the In ...
and
Indus The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmir, ...
river basins of
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
, and
Ganges The Ganges ( ) (in India: Ganga ( ); in Bangladesh: Padma ( )). "The Ganges Basin, known in India as the Ganga and in Bangladesh as the Padma, is an international river to which India, Bangladesh, Nepal and China are the riparian states." is ...
,
Godavari The Godavari ( IAST: ''Godāvarī'' od̪aːʋəɾiː is India's second longest river after the Ganga river and drains into the third largest basin in India, covering about 10% of India's total geographical area. Its source is in Trimbakesh ...
,
Mahanadi The Mahanadi is a major river in East Central India. It drains an area of around and has a total length of . Mahanadi is also known for the Hirakud Dam. The river flows through the states of Chhattisgarh and Odisha and finally merged with Bay ...
and other rivers basins of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
and
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...
. Although widespread, it occurs in low densities, even within protected areas. It is threatened by hunting and habitat loss. It prefers clear, large or medium rivers with sandy bottoms. It spends most of time concealed below the sand, sometimes with only the tip of the nose exposed.


Eating habits

Buried in sand, the Indian narrow-headed softshell turtle waits for its prey to come into its reach. When this happens, the turtle's head extends out of its shell at a high velocity in order to grab and then eat the prey. In a 2009 video from
Animal Planet Animal Planet (stylized in all lowercase since 2018) is an American multinational pay television channel owned by the Warner Bros. Discovery Networks unit of Warner Bros. Discovery. First established on June 1, 1996, the network is primarily ...
's show ''
River Monsters ''River Monsters'' is a British and United States, American wildlife Documentary film, documentary Television show, television programme produced for Animal Planet by Icon Films of Bristol, United Kingdom. It is hosted by extreme Angling, ang ...
'', this turtle is seen quickly extending its head and long retractable neck out of its shell.


Conservation

The Nepali government has established a turtle breeding center in
Chitwan National Park , iucn_category = II , location = Central Terai of Nepal , established = 1973 , nearest_city = Bharatpur , map = Nepal Bagmati Province#Nepal#India#South Asia , relief = 1 , label = Chitwan National Park , label_position = top , coordina ...
and granted permission to a
non-governmental organization A non-governmental organization (NGO) or non-governmental organisation (see spelling differences) is an organization that generally is formed independent from government. They are typically nonprofit entities, and many of them are active in ...
to rescue and conserve the turtles in eastern Nepal. Despite this, and the turtles' status as an endangered species, it is not on the Nepali government's list of highly protected wildlife. Conservationists have claimed the Nepali government's actions do not go far enough in protecting the turtles from natural and anthropogenic threats, such as increased flooding due to the worsening of
monsoons A monsoon () is traditionally a seasonal reversing wind accompanied by corresponding changes in precipitation but is now used to describe seasonal changes in atmospheric circulation and precipitation associated with annual latitudinal oscill ...
caused by climate change, damming of rivers, and gravel mining. In 2022, after two decades of breeding attempts, 41 newborn turtles hatched at San Diego Zoo.


Threats

The turtles are often caught by humans, both as
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
and for consumption, as turtle meat and eggs are considered a delicacy.


References


Bibliography

* Database entry includes a brief justification of why this species is endangered and the criteria used * Boulenger, G.A. 1889 Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). British Museum, London, 311 pp. * Engstrom, Tag N., H. Bradley Shaffer, and William P. McCord. 2002 Phylogenetic Diversity of Endangered and Critically Endangered Southeast Asian Softshell Turtles (TrionychidaChitra). Biological Conservation 104 (2):173-179 * Gray, J. E. 1831 A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed ol. 9 Whittaker, Treacher and Co., London: 481 + 110 pp. * Gray, J. E. 1831 Synopsis Reptilium or short descriptions of the species of reptiles. Part I: Cataphracta, tortoises, crocodiles, and enaliosaurians. Treuttel, Wurz & Co., London, 85 pp. * Gray, J.E. 1844 Catalogue of Tortoises, Crocodilians, and Amphisbaenians in the Collection of the British Museum. British Museum (Natural History), London. viii + 80 p. * Gray, J. E. 186
Revision of the species of Trionychidae found in Asia and Africa, with descriptions of some new species
Proc. Zool. Soc. London 1864: 76-98 * Webb, R.G. 1980 Gray, Hardwicke, Buchanan-Hamilton, and drawings of Indian softshell turtles (Family Trionychidae). Amphibia-Reptilia 1: 61–74.


External links

* {{Taxonbar, from=Q2078104 Chitra (genus) Reptiles of Pakistan Reptiles of India Reptiles described in 1831 Taxa named by John Edward Gray