The Chironomoidea are a superfamily within the order Diptera, suborder

Nematocera The Nematocera (the name means "thread-horns") are a suborder of elongated flies with thin, segmented antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies but species from suborder Brachycera (the name means "sh ...

infraorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...

Culicomorpha The Culicomorpha are an infraorder of Nematocera, including mosquitoes, black flies, and several extant and extinct families of insects. They originated 176 million years ago, in the Triassic period. There are phylogenetic patterns that are used ...

. This superfamily contains the

families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideal ...

Chironomidae The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many specie ...

Ceratopogonidae Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic. Ceratopogonidae are holomet ...

, Simuliidae, and

Thaumaleidae Thaumaleidae, the solitary midges or trickle midges, are a group of nematoceran flies related to the Ceratopogonidae, Chironomidae, and the Simuliidae. They are small, stocky, yellow to brown flies (3–4 mm). Very few species are known f ...

Description

Chironomoidea have four life stages: the egg, the worm-like larva, the pupa and the winged adult.

Ecology

Chironomoidea show a range of habitats and diets. Using the family Chironomidae as an example, larvae occur most commonly in aquatic vegetation and benthic debris, but also in sand covered in fine organic material, pools on granite outcrops, wood

snags In forest ecology, a snag refers to a standing, dead or dying tree, often missing a top or most of the smaller branches. In freshwater ecology it refers to trees, branches, and other pieces of naturally occurring wood found sunken in rivers and ...

, muddy lake beds and hygropetric seepages. Pupae may occur near the surface of water, in submerged substrata or amongst benthic debris. Larvae may feed on deposits of organic detritus (gathering collectors), filter

diatoms A diatom (New Latin, Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group com ...

and fine particles of detritus from the water column (filtering collectors), chew or bore into live or dead plant matter (shredders), scrape algae, bacteria and diatoms from surfaces (scrapers) or prey on other invertebrates (predators). Adults generally feed on sugar-rich substances such as nectar. Adult females of the families Ceratopogonidae and Simuliidae also feed on blood and can transmit diseases.

Phylogeny

The phylogeny of the Chironomoidea is disputed. Analyses of the male genital tract,

ribosomal RNA Ribosomal ribonucleic acid (rRNA) is a type of non-coding RNA which is the primary component of ribosomes, essential to all cells. rRNA is a ribozyme which carries out protein synthesis in ribosomes. Ribosomal RNA is transcribed from ribosom ...

and

transcriptome The transcriptome is the set of all RNA transcripts, including coding and non-coding, in an individual or a population of cells. The term can also sometimes be used to refer to all RNAs, or just mRNA, depending on the particular experiment. The t ...

have showed that the superfamily is not monophyletic.

Evolution

A fossil chironomoid larva of the genus ''Anisinodus'' (family unknown) indicates that the superfamily existed during the early

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...

References

{{Taxonbar, from=Q2214088 Diptera superfamilies