Celebes warty pig
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The Celebes warty pig (''Sus celebensis''), also called Sulawesi warty pig or Sulawesi pig, is a species in the pig genus ('' Sus'') that lives on Sulawesi in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. It survives in most habitats and can live in altitudes of up to . It has been
domesticated Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
and introduced to a number of other islands in Indonesia.


Description

The Celebes warty pig is a medium-sized pig, and quite variable in size and appearance. Although a number of subspecies have been recognised, it is now regarded as a
monotypic taxon In biology, a monotypic taxon is a taxonomic group ( taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
. It is the only pig species that has been domesticated apart from the
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
; being semi-domesticated may have had an influence on the variability of its appearance. This pig has a head-and-body length of between and a long tail, with males generally being larger than females. The back is rounded and the legs short. The colour is greyish-black, sometimes tinged with red or yellow on the flanks. There are three pairs of facial warts and a fringe of pale bristles on the snout and more bristles on the cheeks. The crown and nape are topped by a short crest of dark bristles, while a dark mid-dorsal stripe extends from the crest towards the tufted tail. Young pigs have longitudinal stripes, but these fade as the piglets grow.


Distribution and habitat

The Celebes warty pig occurs in Sulawesi, being plentiful in central, eastern and south-eastern parts of the island but uncommon in the northeastern and southern parts. It also occurs naturally on the nearby smaller islands of
Buton Buton (also Butung, Boeton or Button) is an island in Indonesia located off the southeast peninsula of Sulawesi. It covers roughly 4,727 square kilometers in area, or about the size of Madura; it is the 129th largest island in the world and ...
, Muna, Kabeana,
Peleng Peleng is an island off the east coast of Sulawesi, Indonesia and is the largest island of the Banggai Islands (''Kepulauan Banggai''). It is surrounded by the Banda Sea and Molucca Sea and has an area of 2,406 km². Some of the smaller i ...
,
Lembeh Lembeh is an island off the north east coast of Sulawesi near the city of Bitung. Lembeh Island is 25 km long and 2 km wide. Lembeh Island is separated from the mainland of Sulawesi by a narrow stretch of water known as the Lembeh Str ...
and the
Togian Islands The Togian (or Togean) Islands are an archipelago of 56 islands and many offshore islets, situated in the Gulf of Tomini, off the coast of Central Sulawesi, in Indonesia. The largest islands are Batudaka, Togean, Talatako and Una-Una. Ther ...
. Besides this, it has been domesticated and introduced to various other islands, has hybridised with '' Sus scrofa'', and has become
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
in some places, giving rise to a number of different pig populations. It inhabits various habitat types including rainforest, swamp, cultivated land and grassland, at altitudes up to .


Ecology

This pig tends to move around in small groups of up to nine individuals, led by a dominant male and including several females and their offspring. Foraging takes place mainly in the early morning and the evening; the diet consists mainly of roots, shoots, leaves and fallen fruit, but also includes carrion,
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s and small
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () (chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with c ...
s. Breeding takes place throughout the year. Gestation periods are probably between four and five months, and litter sizes can be up to eight, but in one study, averaged about two. The most important natural
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
is the
reticulated python The reticulated python (''Malayopython reticulatus'') is a python species native to South and Southeast Asia. It is the world's longest snake, and is among the three heaviest. It is listed as least concern on the IUCN Red List because of its w ...
.


Conservation status

The chief threats faced by this pig are an expanding human population, deforestation, with conversion of the land to agricultural use, and hunting for human consumption. Even in areas such as national parks, where the species is protected, hunting still occurs and the meat is traded in local markets. This over-hunting and loss of habitat has led the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
to assess the pig's conservation status as being "
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
".


Cave art

A cave painting of a Celebes warty pig in the Leang Tedongnge cave on Sulawesi has been dated to an age of at least 45,500 years ago, making it the oldest known cave painting. A layer of calcite over the painting was dated using
uranium Uranium is a chemical element with the symbol U and atomic number 92. It is a silvery-grey metal in the actinide series of the periodic table. A uranium atom has 92 protons and 92 electrons, of which 6 are valence electrons. Uranium is weak ...
-series isotope dating to give a minimum age for the painting. The image was made using
red ochre Ochre ( ; , ), or ocher in American English, is a natural clay earth pigment, a mixture of ferric oxide and varying amounts of clay and sand. It ranges in colour from yellow to deep orange or brown. It is also the name of the colours produced ...
, and is accompanied by images of human hands made by blowing ochre on to the cave wall around the hands. The painting depicts a male pig, including the male's pair of facial warts. It was first seen by Westerners in 2017.


Geology

At one point in time there may have been a portion of the land which shifted upward resulted in increased diversification. The researchers found out that the Sulawesi who roam the island today were preceded by other ancestors who once expanded from the innermost area of the island to the outermost part of it.


References

Frantz, L., Rudzinski, A., Nugraha, A., Evin, A., Burton, J., Hulme-Beaman, A., Linderholm, A., Barnett, R., Vega, R., Irving-Pease, E. K., Haile, J., Allen, R., Leus, K., Shephard, J., Hillyer, M., Gillemot, S., van den Hurk, J., Ogle, S., Atofanei, C., Thomas, M. G., … Larson, G. (2018). Synchronous diversification of Sulawesi's iconic artiodactyls driven by recent geological events. Proceedings. Biological sciences, 285(1876), 20172566. https://doi.org/10.1098/rspb.2017.2566


External links

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The Ultimate Ungulate's entry on Sus celebensis
{{Taxonbar, from=Q535406 Sus (genus) Mammals of Sulawesi Endemic fauna of Indonesia Mammals described in 1843 Taxa named by Salomon Müller Taxa named by Hermann Schlegel