Cattle, or cows (female) and bull
s (male), are the most common type of large domesticated ungulate
s. They are a prominent modern member of the subfamily Bovinae
, are the most widespread species
of the genus
'', and are most commonly classified collectively as ''Bos taurus''.
Cattle are commonly raised as livestock
for meat (beef
, see beef cattle
), for milk
(see dairy cattle
), and for hides
, which are used to make leather
. They are used as riding animal
s and draft animal
en or bullocks
, which pull cart
s and other implements). Another product of cattle is their dung
, which can be used to create manure
. In some regions, such as parts of India, cattle have significant religious meaning
. Cattle, mostly small breeds such as the Miniature Zebu
, are also kept as pet
Around 10,500 years ago, taurine cattle
were domesticated from as few as 80 progenitors in central Anatolia
, the Levant
and Western Iran
[ Op. cit. in ]
According to the Food and Agriculture Organization
(FAO), there are approximately 1.5 billion cattle in the world as of 2018.
In 2009, cattle became one of the first livestock animals to have a fully mapped genome
Cattle were originally identified as three separate species: ''Bos taurus'', the European or "taurine" cattle
(including similar types from Africa and Asia); ''Bos indicus'', the Indicine or "zebu"
; and the extinct ''Bos primigenius'', the aurochs
. The aurochs is ancestral to both zebu and taurine cattle. These have been reclassified as one species, ''Bos taurus'', with three subspecies:
Complicating the matter is the ability of cattle to interbreed
with other closely related species. Hybrid individuals and even breeds exist, not only between taurine cattle and zebu (such as the sanga cattle
, ''Bos taurus africanus''), but also between one or both of these and some other members of the genus
s (the dzo
or yattle), banteng
, and gaur
. Hybrids such as the beefalo
breed can even occur between taurine cattle and either species of bison
, leading some authors to consider them part of the genus ''Bos'', as well. The hybrid origin of some types may not be obvious – for example, genetic testing
of the Dwarf Lulu
breed, the only taurine-type cattle in Nepal, found them to be a mix of taurine cattle, zebu, and yak. However, cattle cannot be successfully hybridized with more distantly related bovines such as water buffalo
or African buffalo
The aurochs originally ranged throughout Europe, North Africa, and much of Asia. In historical times, its range became restricted to Europe, and the last known individual died in Mazovia
, Poland, in about 1627. Breeders have attempted to recreate cattle of similar appearance to aurochs by crossing traditional types of domesticated cattle, creating the Heck cattle
''Cattle'' did not originate as the term for bovine animals. It was borrowed from Anglo-Norman
, itself from medieval Latin 'principal sum of money, capital', itself derived in turn from Latin 'head'. ''Cattle'' originally meant movable personal property
, especially livestock of any kind, as opposed to real property
(the land, which also included wild or small free-roaming animals such as chickens—they were sold as part of the land). The word is a variant of ''chattel
'' (a unit of personal property) and closely related to ''capital
'' in the economic sense. The term replaced earlier Old English
'cattle, property', which survives today as ''fee'' (cf. , , ).
The word "cow" came via Anglo-Saxon
(plural ''cȳ''), from Common Indo-European
) = "a bovine animal", compare fa|gâv|script=Latn, sa|go-|script=Latn, cy|buwch. The plural ''cȳ'' became ''ki'' or ''kie'' in Middle English, and an additional plural ending was often added, giving ''kine'', ''kien'', but also ''kies'', ''kuin'' and others. This is the origin of the now archaic English plural, "kine". The Scots language
singular is coo or cou
, and the plural is "kye".
In older English sources such as the King James Version
of the Bible, "cattle" refers to livestock, as opposed to "deer" which refers to wildlife. "Wild cattle" may refer to feral
cattle or to undomesticated species of the genus ''Bos
''. Today, when used without any other qualifier, the modern meaning of "cattle" is usually restricted to domesticated bovines.
In general, the same words are used in different parts of the world, but with minor differences in the definitions. The terminology described here contrasts the differences in definition between the United Kingdom and other British-influenced parts of the world such as Canada, Australia, New Zealand, Ireland and the United States.
* An "intact" (i.e., not castrated
) adult male is called a ''bull''.
* An adult female that has had a calf (or two, depending on regional usage) is a ''cow''.
* A young female before she has had a calf of her own and is under three years of age is called a ''heifer
'' ( ).
[Delbridge, Arthur, The Macquarie Dictionary, 2nd ed., Macquarie Library, North Ryde, 1991]
A young female that has had only one calf is occasionally called a ''first-calf heifer''.
* Young cattle of both sexes are called ''calves
'' until they are weaned
, then ''weaners'' until they are a year old in some areas; in other areas, particularly with male beef cattle, they may be known as ''feeder calves'' or simply feeders. After that, they are referred to as ''yearlings'' or ''stirks'' if between one and two years of age.
* A castrated male is called a ''steer'' in the United States; older steers are often called ''bullocks'' in other parts of the world, but in North America this term refers to a young bull. Piker bullocks are micky bulls (uncastrated young male bulls) that were caught, castrated and then later lost.
In Australia, the term ''Japanese ox'' is used for grain-fed steers in the weight range of 500 to 650 kg that are destined for the Japanese meat trade. In North America, draft cattle under four years old are called working steers. Improper or late castration on a bull results in it becoming a coarse steer known as a ''stag'' in Australia, Canada and New Zealand. In some countries, an incompletely castrated male is known also as a ''rig
* A castrated male (occasionally a female or in some areas a bull) kept for draft or riding purposes is called an ''ox
'' (plural ''oxen''); ''ox'' may also be used to refer to some carcass products from any adult cattle, such as ox-hide, ox-blood, oxtail, or ox-liver.
* A ''springer'' is a cow or heifer close to calving.
* In all cattle species, a female twin of a bull usually becomes an infertile partial intersex
, and is called a ''freemartin
* A wild, young, unmarked bull is known as a ''micky'' in Australia.
[Coupe, Sheena (ed.), Frontier Country, Vol. 1, Weldon Russell Publishing, Willoughby, 1989, ]
* An unbranded
bovine of either sex is called a ''maverick'' in the US and Canada.
* Neat (horned oxen, from which neatsfoot oil
is derived), beef (young ox) and beefing (young animal fit for slaughtering) are obsolete terms, although ''poll'', ''pollard'' and ''polled cattle
'' are still terms in use for naturally hornless animals, or in some areas also for those that have been disbudded
* Cattle raised for human consumption are called ''beef cattle
''. Within the American beef cattle industry, the older term beef (plural beeves) is still used to refer to an animal of either sex. Some Australian, Canadian, New Zealand and British people use the term ''beast''.
* Cattle bred specifically for milk production are called ''milking'' or ''dairy cattle
a cow kept to provide milk for one family may be called a ''house cow
'' or ''milker''. A ''fresh cow'' is a dairy term for a cow or first-calf heifer who has recently given birth, or "freshened."
* The adjective applying to cattle in general is usually ''bovine''. The terms ''bull'', ''cow'' and ''calf'' are also used by extension to denote the sex or age of other large animals, including whales, hippopotamus
Singular terminology issue
"Cattle" can only be used in the plural
and not in the singular
: it is a plurale tantum
Thus one may refer to "three cattle" or "some cattle", but not "one cattle". "One head of cattle" is a valid though periphrastic way to refer to one animal of indeterminate or unknown age and sex; otherwise no universally used single-word singular form of ''cattle'' exists in modern English, other than the sex- and age-specific terms such as cow, bull, steer and heifer. Historically, "ox" was not a sex-specific term for adult cattle, but generally this is now used only for working cattle
, especially adult castrated males. The term is also incorporated into the names of other species, such as the musk ox
and "grunting ox" (yak
), and is used in some areas to describe certain cattle products such as ox-hide and oxtail
''Cow'' is in general use as a singular for the collective ''cattle.'' The word ''cow'' is easy to use when a singular is needed and the sex is unknown or irrelevant—when "there is a cow in the road", for example. Further, any herd of fully mature cattle in or near a pasture
is statistically likely to consist mostly of cows, so the term is probably accurate even in the restrictive sense. Other than the few bulls needed for breeding, the vast majority of male cattle are castrated as calves and are used as ox
en or slaughtered for meat before the age of three years. Thus, in a pastured herd, any calves or herd bulls usually are clearly distinguishable from the cows due to distinctively different sizes and clear anatomical differences. Merriam-Webster and Oxford Living Dictionaries recognize the sex-nonspecific use of ''cow'' as an alternate definition, whereas Collins and the OED do not.
ly, more general nonspecific
terms may denote cattle when a singular form is needed. ''Head of cattle'' is usually used only after a numeral. Australian, New Zealand and British farmers use the term ''beast'' or ''cattle beast''. ''Bovine'' is also used in Britain. The term ''critter'' is common in the western United States and Canada, particularly when referring to young cattle. In some areas of the American South (particularly the Appalachian region), where both dairy and beef cattle are present, an individual animal was once called a "beef critter", though that term is becoming archaic
Cattle raised for human consumption are called ''beef cattle
''. Within the beef cattle industry in parts of the United States, the term ''beef'' (plural ''beeves'') is still used in its archaic sense to refer to an animal of either sex. Cows of certain breeds that are kept for the milk they give are called ''dairy cow
s'' or ''milking cows'' (formerly ''milch cows''). Most young male offspring of dairy cows are sold for veal
, and may be referred to as veal calves.
The term ''dogies'' is used to describe orphaned calves in the context of ranch
work in the American West
, as in "Keep them dogies moving". In some places, a cow kept to provide milk for one family is called a "house cow". Other obsolete terms for cattle include "neat" (this use survives in "neatsfoot oil
", extracted from the feet and legs of cattle), and "beefing" (young animal fit for slaughter
term for one of the most common sounds
made by cattle is ''moo'' (also called ''lowing''). There are a number of other sounds made by cattle, including calves ''bawling'', and bulls ''bellowing''. Bawling is most common for cows after weaning of a calf. The bullroarer
makes a sound similar to a bull's territorial call.
Cattle are large quadrupedal ungulate mammal
s with cloven hooves
. Most breeds have horns
, which can be as large as the Texas Longhorn
or small like a scur
. Careful genetic selection has allowed polled
(hornless) cattle to become widespread.
Cattle are ruminant
s, meaning their digestive system
is highly specialized to allow the use of poorly digestible plants as food. Cattle have one stomach
with four compartments, the rumen
, and abomasum
, with the rumen being the largest compartment.
The reticulum, the smallest compartment, is known as the "honeycomb". The omasum's main function is to absorb water and nutrients from the digestible feed. The omasum is known as the "many plies". The abomasum is like the human stomach; this is why it is known as the "true stomach".
Cattle are known for regurgitating
and re-chewing their food, known as cud
chewing, like most ruminants. While the animal is feeding, the food is swallowed without being chewed and goes into the rumen for storage until the animal can find a quiet place to continue the digestion process. The food is regurgitated, a mouthful at a time, back up to the mouth, where the food, now called the cud
, is chewed by the molars, grinding down the coarse vegetation to small particles. The cud is then swallowed again and further digested by specialized microorganism
s in the rumen. These microbes are primarily responsible for decomposing cellulose
and other carbohydrate
s into volatile fatty acids
cattle use as their primary metabolic
fuel. The microbes inside the rumen also synthesize amino acid
s from non-protein nitrogen
ous sources, such as urea
. As these microbes reproduce in the rumen, older generations die and their cells continue on through the digestive tract. These cells are then partially digested in the small intestines, allowing cattle to gain a high-quality protein source. These features allow cattle to thrive on grasses
and other tough vegetation.
Gestation and size
The gestation period
for a cow is about nine months long. A newborn calf's size can vary among breeds, but a typical calf weighs . Adult size and weight vary significantly among breeds and sex. Steers are generally killed before reaching . Breeding stock may be allowed a longer lifespan, occasionally living as long as 25 years. The oldest recorded cow, Big Bertha
, died at the age of 48 in 1993.
On farms it is very common to use artificial insemination
(AI), a medically assisted reproduction technique
consisting of the artificial deposition of semen
in the female's genital tract
It is used in cases where the spermatozoa
can not reach the fallopian tubes
or simply by choice of the owner of the animal. It consists of transferring, to the uterine cavity
, spermatozoa previously collected and processed, with the selection of morphologically more normal and mobile spermatozoa.
A cow's udder
contains two pairs of mammary gland
s, (commonly referred to as ''teats'') creating four "quarters". The front ones are referred to as ''fore quarters'' and the rear ones ''rear quarters''.
Synchronization of cattle ovulation
to benefit dairy farming
may be accomplished via induced ovulation
Bulls become fertile at about seven months of age. Their fertility is closely related to the size of their testicle
s, and one simple test of fertility is to measure the circumference of the scrotum: a young bull is likely to be fertile once this reaches ; that of a fully adult bull may be over .
A bull has a fibro-elastic penis. Given the small amount of erectile tissue, there is little enlargement after erection. The penis is quite rigid when non-erect, and becomes even more rigid during erection. Protrusion is not affected much by erection, but more by relaxation of the retractor penis muscle
and straightening of the sigmoid flexure
The weight of adult cattle varies, depending on the breed. Smaller kinds, such as Dexter and Jersey adults, range between . Large Continental breeds, such as Charolais, Marchigiana, Belgian Blue and Chianina adults range from . British breeds, such as Hereford, Angus, and Shorthorn, mature at between , occasionally higher, particularly with Angus and Hereford. Bulls are larger than cows of the same breed by up to a few hundred kilograms. Chianina bulls can weigh up to ; British bulls, such as Angus and Hereford, can weigh as little as to as much as .
The world record for the heaviest bull was , a Chianina
named Donetto, when he was exhibited at the Arezzo
show in 1955.
[Friend, John B., Cattle of the World, Blandford Press, Dorset, 1978]
The heaviest steer was eight-year-old 'Old Ben', a Shorthorn
cross weighing in at in 1910.
[McWhirter, Norris & Ross, ''Guinness Book of Records'', Redwood Press, Trowbridge, 1968]
In the United States, the average weight of beef cattle has steadily increased, especially since the 1970s, requiring the building of new slaughterhouses able to handle larger carcasses. New packing plants in the 1980s stimulated a large increase in cattle weights. Before 1790 beef cattle averaged only net; and thereafter weights climbed steadily.
In laboratory studies, young cattle are able to memorize the locations of several food sources and retain this memory for at least 8 hours, although this declined after 12 hours.
Fifteen-month-old heifers learn more quickly than adult cows which have had either one or two calvings, but their longer-term memory is less stable. Mature cattle perform well in spatial learning tasks and have a good long-term memory in these tests. Cattle tested in a radial arm maze
are able to remember the locations of high-quality food for at least 30 days. Although they initially learn to avoid low-quality food, this memory diminishes over the same duration.
Under less artificial testing conditions, young cattle showed they were able to remember the location of feed for at least 48 days. Cattle can make an association between a visual stimulus and food within 1 day—memory of this association can be retained for 1 year, despite a slight decay.
Calves are capable of discrimination learning and adult cattle compare favourably with small mammals in their learning ability in the Closed-field Test
They are also able to discriminate between familiar individuals, and among humans. Cattle can tell the difference between familiar and unfamiliar animals of the same species (conspecifics). Studies show they behave less aggressively toward familiar individuals when they are forming a new group.
Calves can also discriminate between humans based on previous experience, as shown by approaching those who handled them positively and avoiding those who handled them aversively.
Although cattle can discriminate between humans by their faces alone, they also use other cues such as the color of clothes when these are available.
In audio play-back studies, calves prefer their own mother's vocalizations compared to the vocalizations of an unfamiliar mother.
In laboratory studies using images, cattle can discriminate between images of the heads of cattle and other animal species.
They are also able to distinguish between familiar and unfamiliar conspecifics. Furthermore, they are able to categorize
images as familiar and unfamiliar individuals.
When mixed with other individuals, cloned
calves from the same donor form subgroups, indicating that kin discrimination occurs and may be a basis of grouping behaviour. It has also been shown using images of cattle that both artificially inseminated and cloned calves have similar cognitive capacities of kin and non-kin discrimination.
Cattle can recognize familiar individuals. Visual individual recognition is a more complex mental process than visual discrimination. It requires the recollection of the learned idiosyncratic identity of an individual that has been previously encountered and the formation of a mental representation.
By using 2-dimensional images of the heads of one cow (face, profiles, views), all the tested heifers showed individual recognition of familiar and unfamiliar individuals from their own breed. Furthermore, almost all the heifers recognized unknown individuals from different breeds, although this was achieved with greater difficulty. Individual recognition was most difficult when the visual features of the breed being tested were quite different from the breed in the image, for example, the breed being tested had no spots whereas the image was of a spotted breed.
Cattle use visual/brain lateralisation
in their visual scanning of novel and familiar stimuli. Domestic cattle prefer to view novel stimuli with the left eye, i.e. using the right brain hemisphere (similar to horses, Australian magpies, chicks, toads and fish) but use the right eye, i.e. using the left hemisphere, for viewing familiar stimuli.
Temperament and emotions
In cattle, temperament can affect production traits such as carcass and meat quality or milk yield as well as affecting the animal's overall health and reproduction. Cattle temperament is defined as "the consistent behavioral and physiological difference observed between individuals in response to a stressor or environmental challenge and is used to describe the relatively stable difference in the behavioral predisposition of an animal, which can be related to psychobiological mechanisms".
Generally, cattle temperament is assumed to be multidimensional. Five underlying categories of temperament traits have been proposed:
In a study on Holstein–Friesian heifers learning to press a panel to open a gate for access to a food reward, the researchers also recorded the heart rate and behavior of the heifers when moving along the race towards the food. When the heifers made clear improvements in learning, they had higher heart rates and tended to move more vigorously along the race. The researchers concluded this was an indication that cattle may react emotionally to their own learning improvement.
Negative emotional states are associated with a bias toward negative responses towards ambiguous cues in judgement tasks. After separation from their mothers, Holstein calves showed such a cognitive bias indicative of low mood. A similar study showed that after hot-iron disbudding (dehorning
), calves had a similar negative bias indicating that post-operative pain following this routine procedure results in a negative change in emotional state.
In studies of visual discrimination, the position of the ears has been used as an indicator of emotional state.
When cattle are stressed other cattle can tell by the chemicals released in their urine.
Cattle are very gregarious
and even short-term isolation is considered to cause severe psychological stress
. When Aubrac
and Friesian heifers are isolated, they increase their vocalizations and experience increased heart rate
and plasma cortisol
concentrations. These physiological changes are greater in Aubracs. When visual contact is re-instated, vocalizations rapidly decline, regardless of the familiarity of the returning cattle, however, heart rate decreases are greater if the returning cattle are familiar to the previously-isolated individual. Mirrors have been used to reduce stress in isolated cattle.
Cattle use all of the five widely recognized sensory modalities. These can assist in some complex behavioural patterns, for example, in grazing behaviour. Cattle eat mixed diets, but when given the opportunity, show a partial preference of approximately 70% clover and 30% grass. This preference has a diurnal pattern, with a stronger preference for clover in the morning, and the proportion of grass increasing towards the evening.
Vision is the dominant sense in cattle and they obtain almost 50% of their information visually.
Cattle are a prey animal and to assist predator detection, their eyes are located on the sides of their head rather than the front. This gives them a wide field of view of 330° but limits binocular vision
(and therefore stereopsis
) to 30° to 50° compared to 140° in humans.
This means they have a blind spot directly behind them. Cattle have good visual acuity,
but compared to humans, their visual accommodation is poor.
Cattle have two kinds of color receptors
in the cone cells
of their retina
s. This means that cattle are dichromatic
, as are most other non-primate land mammals. There are two to three rods per cone in the fovea centralis
but five to six near the optic papilla.
Cattle can distinguish long wavelength colors (yellow, orange and red) much better than the shorter wavelengths (blue, grey and green). Calves are able to discriminate between long (red) and short (blue) or medium (green) wavelengths, but have limited ability to discriminate between the short and medium. They also approach handlers more quickly under red light.
Whilst having good color sensitivity, it is not as good as humans or sheep
A common misconception about cattle (particularly bulls) is that they are enraged by the color red (something provocative is often said to be "like a red flag to a bull"). This is a myth. In bullfighting
, it is the movement of the red flag or cape that irritates the bull and incites it to charge.
Cattle have a well-developed sense of taste and can distinguish the four primary tastes (sweet, salty, bitter and sour). They possess around 20,000 taste buds. The strength of taste perception depends on the individual's current food requirements. They avoid bitter-tasting foods (potentially toxic) and have a marked preference for sweet (high calorific value) and salty foods (electrolyte
balance). Their sensitivity to sour-tasting foods helps them to maintain optimal ruminal pH
Plants have low levels of sodium and cattle have developed the capacity of seeking salt by taste and smell. If cattle become depleted of sodium salts, they show increased locomotion directed to searching for these. To assist in their search, the olfactory and gustatory receptors able to detect minute amounts of sodium salts increase their sensitivity as biochemical disruption develops with sodium salt depletion.
Cattle hearing ranges from 23 Hz
to 35 kHz. Their frequency of best sensitivity is 8 kHz and they have a lowest threshold of −21 db
(re 20 μN/m−2
), which means their hearing is more acute than horses (lowest threshold of 7 db).
Sound localization acuity thresholds are an average of 30°. This means that cattle are less able to localise sounds compared to goats (18°), dogs (8°) and humans (0.8°).
Because cattle have a broad foveal fields of view covering almost the entire horizon, they may not need very accurate locus information from their auditory systems to direct their gaze to a sound source.
Vocalizations are an important mode of communication amongst cattle and can provide information on the age, sex, dominance status and reproductive status of the caller. Calves can recognize their mothers using vocalizations; vocal behaviour may play a role by indicating estrus and competitive display by bulls.
Olfaction and gustation
Cattle have a range of odiferous glands over their body including interdigital
glands, indicating that olfaction probably plays a large role in their social life. Both the primary olfactory system using the olfactory bulb
s, and the secondary olfactory system using the vomeronasal organ
This latter olfactory system is used in the flehmen
response. There is evidence that when cattle are stressed, this can be recognised by other cattle and this is communicated by alarm substances in the urine.
The odour of dog faeces induces behavioural changes prior to cattle feeding, whereas the odours of urine from either stressed or non-stressed conspecifics and blood have no effect.
In the laboratory, cattle can be trained to recognise conspecific individuals using olfaction only.
In general, cattle use their sense of smell to "expand" on information detected by other sensory modalities. However, in the case of social and reproductive behaviours, olfaction is a key source of information.
Cattle have tactile sensations detected mainly by mechanoreceptor
s and nociceptor
s in the skin and muzzle. These are used most frequently when cattle explore their environment.
There is conflicting evidence for magnetoreception
in cattle. One study reported that resting and grazing cattle tend to align their body axes in the geomagnetic north–south (N-S) direction. In a follow-up study, cattle exposed to various magnetic fields directly beneath or in the vicinity of power lines trending in various magnetic directions exhibited distinct patterns of alignment. However, in 2011, a group of Czech
researchers reported their failed attempt to replicate the finding using Google Earth
Under natural conditions, calves stay with their mother until weaning at 8 to 11 months. Heifer and bull calves are equally attached to their mothers in the first few months of life.
Cattle are considered to be "hider" type animals, but in the artificial environment of small calving pens, close proximity between cow and calf is maintained by the mother at the first three calvings but this changes to being mediated by the calf after these. Primiparous dams show a higher incidence of abnormal maternal behavior.
Beef-calves reared on the range suckle an average of 5.0 times every 24 hours with an average total time of 46 min spent suckling. There is a diurnal rhythm in suckling activity with peaks between 05:00–07:00, 10:00–13:00 and 17:00–21:00.
Studies on the natural weaning of zebu cattle (''Bos indicus'') have shown that the cow weans her calves over a 2-week period, but after that, she continues to show strong affiliatory behavior with her offspring and preferentially chooses them for grooming and as grazing partners for at least 4–5 years.
Semi-wild Highland cattle
heifers first give birth at 2 or 3 years of age, and the timing of birth is synchronized with increases in natural food quality. Average calving interval is 391 days, and calving mortality within the first year of life is 5%.
Dominance and leadership
One study showed that over a 4-year period, dominance relationships within a herd of semi-wild highland cattle were very firm. There were few overt aggressive conflicts and the majority of disputes were settled by agonistic
(non-aggressive, competitive) behaviors that involved no physical contact between opponents (e.g. threatening and spontaneous withdrawing). Such agonistic behavior reduces the risk of injury. Dominance status depended on age and sex, with older animals generally being dominant to young ones and males dominant to females. Young bulls gained superior dominance status over adult cows when they reached about 2 years of age.
As with many animal dominance hierarchies, dominance-associated aggressiveness does not correlate with rank position, but is closely related to rank distance between individuals.
Dominance is maintained in several ways. Cattle often engage in mock fights where they test each other's strength in a non-aggressive way. Licking is primarily performed by subordinates and received by dominant animals. Mounting is a playful behavior shown by calves of both sexes and by bulls and sometimes by cows in estrus, however, this is not a dominance related behavior as has been found in other species.
The horns of cattle are "honest signals
" used in mate selection. Furthermore, horned cattle attempt to keep greater distances between themselves and have fewer physical interactions than hornless cattle. This leads to more stable social relationships.
In calves, the frequency of agonistic behavior decreases as space allowance increases, but this does not occur for changes in group size. However, in adult cattle, the number of agonistic encounters increases as the group size increases.
When grazing, cattle vary several aspects of their bite, i.e. tongue and jaw movements, depending on characteristics of the plant they are eating. Bite area decreases with the density of the plants but increases with their height. Bite area is determined by the sweep of the tongue; in one study observing steers, bite area reached a maximum of approximately . Bite depth increases with the height of the plants. By adjusting their behavior, cattle obtain heavier bites in swards that are tall and sparse compared with short, dense swards of equal mass/area. Cattle adjust other aspects of their grazing behavior in relation to the available food; foraging velocity decreases and intake rate increases in areas of abundant palatable forage.
Cattle avoid grazing areas contaminated by the faeces of other cattle more strongly than they avoid areas contaminated by sheep, but they do not avoid pasture contaminated by rabbit faeces.
On 24 April 2009, edition of the journal ''Science'', a team of researchers led by the National Institutes of Health and the US Department of Agriculture
reported having mapped the bovine genome
The scientists found cattle have about 22,000 genes, and 80% of their genes are shared with humans, and they share about 1000 genes with dogs and rodents, but are not found in humans. Using this bovine "HapMap", researchers can track the differences between the breeds that affect the quality of meat and milk yields.
Behavioral traits of cattle can be as heritable
as some production traits, and often, the two can be related. The heritability of fear varies markedly in cattle from low (0.1) to high (0.53); such high variation is also found in pigs and sheep, probably due to differences in the methods used.
The heritability of temperament (response to isolation during handling) has been calculated as 0.36 and 0.46 for habituation
to handling. Rangeland assessments show that the heritability of aggressiveness in cattle is around 0.36.
Quantitative trait loci
(QTLs) have been found for a range of production and behavioral characteristics for both dairy and beef cattle.
Domestication and husbandry
Cattle occupy a unique role in human history
, having been domesticated since at least the early neolithic
Archaeozoological and genetic data indicate that cattle were first domesticated from wild aurochs
(''Bos primigenius'') approximately 10,500 years ago. There were two major areas of domestication: one in the Near East
(specifically central Anatolia
, the Levant
and Western Iran
), giving rise to the taurine line, and a second in the area that is now Pakistan, resulting in the indicine line.
Modern mitochondrial DNA variation indicates the taurine line may have arisen from as few as 80 aurochs
tamed in the upper reaches of Mesopotamia
near the villages of Çayönü Tepesi
in what is now southeastern Turkey and Dja'de el-Mughara
in what is now northern Iraq
Although European cattle are largely descended from the taurine lineage, gene flow from African cattle (partially of indicine origin) contributed substantial genomic components to both southern European cattle breeds and their New World descendants.
A study on 134 breeds showed that modern taurine cattle originated from Africa, Asia, North and South America, Australia, and Europe.
Some researchers have suggested that African taurine cattle are derived from a third independent domestication from North African aurochsen.
Usage as money
As early as 9000 BC both grain and cattle were used as money or as ''barter'' (the ''first grain remains
'' found, considered to be evidence of pre-agricultural practice date to 17,000 BC).
Some evidence also exists to suggest that other animals, such as camels
, may have been used as currency in some parts of the world. One of the advantages of using cattle as currency is that it allows the seller to set a fixed price. It even created the standard pricing. For example, two chickens were traded for one cow as cows were deemed to be more valuable than chickens.
Cattle are often raised by allowing herds to graze on the grasses
of large tracts of rangeland
. Raising cattle in this manner allows the use of land that might be unsuitable for growing crops. The most common interactions with cattle involve daily feeding
, cleaning and milking
. Many routine husbandry practices involve ear tag
, loading, medical operations
, vaccinations and hoof
care, as well as training for agricultural shows and preparations. Also, some cultural differences
occur in working with cattle; the cattle husbandry of Fulani
men rests on behavioural techniques
, whereas in Europe, cattle are controlled primarily by physical means, such as fence
s. Breeders use cattle husbandry to reduce ''M. bovis'' infection
susceptibility by selective breeding
and maintaining herd health to avoid concurrent disease.
Cattle are farmed for beef, veal, dairy, and leather. They are less commonly used for conservation grazing
, or simply to maintain grassland for wildlife, such as in Epping Forest
, England. They are often used in some of the most wild places for livestock. Depending on the breed, cattle can survive on hill grazing, heaths, marshes, moors and semidesert. Modern cattle are more commercial than older breeds and, having become more specialized, are less versatile. For this reason, many smaller farmers still favor old breeds, such as the Jersey
In Portugal, Spain, southern France
and some Latin American countries, bulls are used in the activity of bullfighting
'' in India is a bull taming sport radically different from European bullfighting, humans are unarmed and bulls are not killed. In many other countries bullfighting is illegal. Other activities such as bull riding
are seen as part of a rodeo
, especially in North America. Bull-leaping
, a central ritual in Bronze Age Minoan
culture (see Sacred Bull
), still exists in southwestern France. In modern times, cattle are also entered into agricultural competitions
. These competitions can involve live cattle or cattle carcases in hoof and hook events.
In terms of food intake by humans, consumption of cattle is less efficient than of grain or vegetables with regard to land use
, and hence cattle grazing consumes more area than such other agricultural production when raised on grains.
[Edward O. Wilson, ''The Future of Life'', 2003, Vintage Books, 256 pages ]
Nonetheless, cattle and other forms of domesticated animals can sometimes help to use plant resources in areas not easily amenable to other forms of agriculture. Bulls are sometimes used as guard animals
The average sleep time of a domestic cow is about 4 hours a day. Cattle do have a stay apparatus
, but do not sleep standing up; they lie down to sleep deeply.
In spite of the urban legend
, cows cannot be tipped over by people pushing on them.
The meat of adult cattle is known as beef
, and that of calves
. Other animal parts are also used as food products, including blood, liver
. Cattle also produce milk
, and dairy cattle
are specifically bred to produce the large quantities of milk processed and sold for human consumption. Cattle today are the basis of a multibillion-dollar industry worldwide. The international trade
in beef for 2000 was over $30 billion and represented only 23% of world beef production. Approximately 300 million cattle, including dairy cattle, are slaughtered each year for food. The production of milk, which is also made into cheese, butter, yogurt
, and other dairy products, is comparable in economic size to beef production, and provides an important part of the food supply for many of the world's people. Cattle hides, used for leather to make shoes, couch
es and clothing, are another widespread product. Cattle remain broadly used as draft animals in many developing countries
, such as India. Cattle are also used in some sporting games, including rodeo
Cattle meat production
Source: Helgi Library,
[| Cattle Meat Production | 12 February 2014]
World Bank, FAOSTAT
About half the world's meat comes from cattle.
Certain breeds of cattle, such as the Holstein-Friesian
, are used to produce milk
which can be processed into dairy products such as milk, cheese
. Dairy cattle are usually kept on specialized dairy farms designed for milk production. Most cows are milked twice per day, with milk processed at a dairy, which may be onsite at the farm or the milk may be shipped to a dairy plant for eventual sale of a dairy product.
Lactation is induced in heifers and spayed cows by a combination of physical and psychological stimulation, by drugs, or by a combination of those methods. For mother cows to continue producing milk, they give birth to one calf per year. If the calf is male, it generally is slaughtered at a young age to produce veal
. They will continue to produce milk until three weeks before birth.
Over the last fifty years, dairy farming has become more intensive to increase the yield of milk produced by each cow. The Holstein-Friesian
is the breed of dairy cow most common in the UK, Europe and the United States. It has been bred selectively to produce the highest yields of milk of any cow. Around 22 litres per day is average in the UK.
Most cattle are not kept solely for hides, which are usually a by-product of beef production. Hides are most commonly used for leather, which can be made into a variety of product, including shoes. In 2012 India was the world's largest producer of cattle hides.
cattle are defined as being 'cattle that are not domesticated or cultivated'. Populations of feral cattle are known to come from and exist in: Australia
, United States of America
and many islands, including New Guinea
, Juan Fernández Islands
), Tristan da Cunha
and Île Amsterdam
two islands of Kuchinoshima
and Kazura Island next to Naru Island
. Chillingham cattle
is sometimes regarded as a feral breed. Aleutian wild cattle
s can be found on Aleutian Islands
. The "Kinmen cattle" which is dominantly found on Kinmen Island
is mostly domesticated while smaller portion of the population is believed to live in the wild due to accidental releases.
Other notable examples include cattle in the vicinity of Hong Kong (in the Shing Mun Country Park
, among Sai Kung District
and Lantau Island
and on Grass Island
), and semi-feral animals in Yangmingshan
in cattle include methanogen
s that produce methane
as a byproduct of enteric fermentation
, which cattle belch out. The same volume of atmospheric methane
has a higher global warming potential
than atmospheric carbon dioxide
[ See Table 8.7.]
Methane belching from cattle can be reduced with genetic selection, immunization, rumen defaunation, diet modification, decreased antibiotic use, and grazing management, among others.
A report from the Food and Agriculture Organization
(FAO) states that the livestock sector is "responsible for 18% of greenhouse gas emissions".
[Steinfeld, H. et al. 2006,]
Livestock's Long Shadow: Environmental Issues and Options
Livestock, Environment and Development, FAO.
estimates that cattle and other livestock emit about 80 to 93 Megatonnes of methane per year, accounting for an estimated 37% of anthropogenic methane emissions,
and additional methane is produced by anaerobic fermentation of manure in manure lagoon
s and other manure storage structures. The net change in atmospheric methane content was recently about 1 Megatonne per year, and in some recent years there has been no increase in atmospheric methane content. While cattle fed forage actually produce more methane than grain-fed cattle, the increase may be offset by the increased carbon recapture of pastures, which recapture three times the CO2
of cropland used for grain.
One of the cited changes suggested to reduce greenhouse gas emissions is intensification of the livestock industry, since intensification leads to less land for a given level of production. This assertion is supported by studies of the US beef production system, suggesting practices prevailing in 2007 involved 8.6% less fossil fuel use, 16.3% less greenhouse gas emissions, 12.1% less water use, and 33.0% less land use, per unit mass of beef produced, than those used in 1977. The analysis took into account not only practices in feedlot
s, but also feed production (with less feed needed in more intensive production systems), forage-based cow-calf operations and back-grounding before cattle enter a feedlot (with more beef produced per head of cattle from those sources, in more intensive systems), and beef from animals derived from the dairy industry.
The number of American cattle kept in confined feedlot conditions fluctuates. From 1 January 2002 through 1 January 2012, there was no significant overall upward or downward trend in the number of US cattle on feed for slaughter, which averaged about 14.046 million head over that period.
[USDA. 2011. Agricultural Statistics 2011. US Government Printing Office, Washington. 509 pp. Table 7.6.] [USDA. 2012. Cattle. ]
Previously, the number had increased; it was 12.453 million in 1985. Cattle on feed (for slaughter) numbered about 14.121 million on 1 January 2012, i.e. about 15.5% of the estimated inventory of 90.8 million US cattle (including calves) on that date. Of the 14.121 million, US cattle on feed (for slaughter) in operations with 1000 head or more were estimated to number 11.9 million.
Cattle feedlots in this size category correspond to the regulatory definition of "large" concentrated animal feeding operations
(CAFOs) for cattle other than mature dairy cows or veal calves.
[US Code of Federal Regulations 40 CFR 122.23]
Significant numbers of dairy, as well as beef cattle, are confined in CAFOs, defined as "new and existing operations which stable or confine and feed or maintain for a total of 45 days or more in any 12-month period more than the number of animals specified" where "
ops, vegetation, forage growth, or post-harvest residues are not sustained in the normal growing season over any portion of the lot or facility." They may be designated as small, medium and large. Such designation of cattle CAFOs is according to cattle type (mature dairy cows, veal calves or other) and cattle numbers, but medium CAFOs are so designated only if they meet certain discharge criteria, and small CAFOs are designated only on a case-by-case basis.
A CAFO that discharges pollutants is required to obtain a permit, which requires a plan to manage nutrient runoff, manure, chemicals, contaminants, and other wastewater pursuant to the US Clean Water Act
. The regulations involving CAFO permitting have been extensively litigated.
Commonly, CAFO wastewater and manure nutrients are applied to land at agronomic rates for use by forages or crops, and it is often assumed that various constituents of wastewater and manure, e.g. organic contaminants and pathogens, will be retained, inactivated or degraded on the land with application at such rates; however, additional evidence is needed to test reliability of such assumptions
. Concerns raised by opponents of CAFOs have included risks of contaminated water due to feedlot
soil erosion, human and animal exposure to toxic chemicals, development of antibiotic resistant bacteria
and an increase in ''E. coli
'' contamination. While research suggests some of these impacts can be mitigated by developing wastewater treatment systems
and planting cover crops in larger setback zones, the Union of Concerned Scientists
released a report in 2008 concluding that CAFOs are generally unsustainable and externalize costs
An estimated 935,000 cattle operations were operating in the US in 2010.
[USDA. 2011. Agricultural Statistics 2011. US Government Printing Office, Washington. 509 pp. Table 7.1.]
In 2001, the US Environmental Protection Agency (EPA) tallied 5,990 cattle CAFOs then regulated, consisting of beef (2,200), dairy (3,150), heifer (620) and veal operations (20). Since that time, the EPA has established CAFOs as an enforcement priority. EPA enforcement highlights for fiscal year 2010 indicated enforcement actions against 12 cattle CAFOs for violations that included failures to obtain a permit, failures to meet the terms of a permit, and discharges of contaminated water.
Another concern is manure
, which if not well-managed, can lead to adverse environmental consequences. However, manure also is a valuable source of nutrients and organic matter when used as a fertilizer. Manure was used as a fertilizer on about of US cropland in 2006, with manure from cattle accounting for nearly 70% of manure applications to soybeans and about 80% or more of manure applications to corn, wheat, barley, oats and sorghum. Substitution of manure for synthetic fertilizers in crop production can be environmentally significant, as between 43 and 88 megajoule
s of fossil fuel energy would be used per kg of nitrogen in manufacture of synthetic nitrogenous fertilizers.
Grazing by cattle at low intensities can create a favourable environment for native herbs and forb
s by mimicking the native grazers who they displaced; in many world regions, though, cattle are reducing biodiversity
due to overgrazing
. A survey of refuge managers on 123 National Wildlife Refuges in the US tallied 86 species of wildlife considered positively affected and 82 considered negatively affected by refuge cattle grazing or haying. Proper management of pastures, notably managed intensive rotational grazing
and grazing at low intensities can lead to less use of fossil fuel energy, increased recapture of carbon dioxide, fewer ammonia emissions into the atmosphere, reduced soil erosion, better air quality, and less water pollution.
The veterinary discipline dealing with cattle and cattle diseases (bovine veterinary) is called buiatrics. Veterinarians and professionals working on cattle health issues are pooled in the World Association for Buiatrics, founded in 1960. National associations and affiliates also exist.
Cattle diseases were in the center of attention in the 1980s and 1990s when the Bovine spongiform encephalopathy
(BSE), also known as mad cow disease, was of concern. Cattle might catch and develop various other diseases, like blackleg
, foot rot
In most states, as cattle health is not only a veterinarian issue, but also a public health issue, public health
and food safety
standards and farming regulations directly affect the daily work of farmers who keep cattle. However, said rules change frequently and are often debated. For instance, in the U.K., it was proposed in 2011 that milk from tuberculosis
-infected cattle should be allowed to enter the food chain. Internal food safety regulations might affect a country's trade policy as well. For example, the United States has just reviewed its beef import rules according to the "mad cow standards"; while Mexico forbids the entry of cattle who are older than 30 months.
is commonly used in India for internal medical purposes. It is distilled and then consumed by patients seeking treatment for a wide variety of illnesses. At present, no conclusive medical evidence shows this has any effect. However, an Indian medicine containing cow urine has already obtained U.S. patents.
is caused by the bacteria from the genus Treponema. It differs from foot rot and can appear under unsanitary conditions such as poor hygiene or inadequate hoof trimming, among other causes. It primarily affects dairy cattle and has been known to lower the quantity of milk produced, however the milk quality remains unaffected. Cattle are also susceptible to ringworm
caused by the fungus, ''Trichophyton verrucosum
'', a contagious skin disease which may be transferred to humans exposed to infected cows.
Effect of high stocking density
Stocking density refers to the number of animals within a specified area. When stocking density reaches high levels, the behavioural needs of the animals may not be met. This can negatively influence health, welfare and production performance.
The effect of overstocking in cows can have a negative effect on milk production
rates which are two very important traits for dairy farmers. Overcrowding of cows in barns has been found to reduced feeding, resting and rumination.
Although they consume the same amount of dry matter within the span of a day, they consume the food at a much more rapid rate, and this behaviour in cows can lead to further complications. The feeding behaviour of cows during their post-milking period is very important as it has been proven that the longer animals can eat after milking, the longer they will be standing up and therefore causing less contamination to the teat ends.
This is necessary to reduce the risk of mastitis as infection has been shown to increase the chances of embryonic loss
. Sufficient rest is important for dairy cows because it is during this period that their resting blood flow increases up to 50%, this is directly proportionate to milk production.
Each additional hour of rest can be seen to translate to 2 to 3.5 more pounds of milk per cow daily. Stocking densities of anything over 120% have been shown to decrease the amount of time cows spend lying down.
is an important stress hormone; its plasma concentrations increase greatly when subjected to high levels of stress.
[Sjaasted O.V., Howe K., Sand O., (2010) Physiology of Domestic Animals. 3rd edition. Sunderland: Sinaver Association. Inc]
Increased concentration levels of cortisol have been associated with significant increases in gonadotrophin levels and lowered progestin levels. Reduction of stress is important in the reproductive state of cows as an increase in gonadotrophin
and lowered progesterone
levels may impinge on the ovulatory
process and to reduce the chances of successful implantation. A high cortisol level will also stimulate the degradation of fats and proteins which may make it difficult for the animal to sustain its pregnancy if implanted successfully.
Animal welfare concerns
activists have criticized the treatment of cattle, claiming that common practices in cattle husbandry, slaughter
and entertainment unnecessarily cause fear, stress, and pain. They advocate for abstaining from the consumption of cattle-related animal products and cattle-based entertainment.
The following husbandry practices have been criticized by animal welfare and animal rights groups: branding
, ear tag
ging, nose ringing
, tail docking
, the use of veal crates
and cattle prod
s. There are concerns that the stress and negative health impacts induced by high stocking density such as in concentrated animal feeding operations
, auctions, and during transport may be detrimental to their welfare,
and has also been criticized.
The treatment of dairy cows
faces additional criticism. To produce milk from dairy cattle
, most calves are separated from their mothers soon after birth and fed milk replacement in order to retain the cows' milk for human consumption.
Animal welfare advocates are critical of this practice, stating that this breaks the natural bond between the mother and her calf.
The welfare of veal calves
is also a concern.
In order to continue lactation
, dairy cows are bred every year, usually through artificial insemination
Because of this, some individuals have posited that dairy production is based on the sexual exploitation of cows. Although the natural life expectancy of cattle could be as much as twenty years,
after about five years, a cow's milk production has dropped; at which point most dairy cows are sent to slaughter.
While leather is often a by-product of slaughter, in some countries, such as India and Bangladesh, cows are raised primarily for their leather. These leather industries often make their cows walk long distances across borders to be killed in neighboring provinces and countries where cattle slaughter is legal. Some cows die along the long journey, and sometimes exhausted animals are abused to keep them moving. These practices have faced backlash from various animal rights groups.
Animal treatment in rodeo
is targeted most often at bull riding
but also calf roping
and steer roping
, with the opposition saying that rodeos are unnecessary and cause stress, injury, and death to the animals. In Spain, the Running of the bulls faces opposition
due to the stress and injuries incurred by the bulls during the event. Bullfighting is opposed
as a blood sport
in which bulls are forced to suffer severe stress and death.
Oxen (singular ox) are cattle trained as draft animal
s. Often they are adult, castrated
males of larger breeds, although females and bulls are also used in some areas. Usually, an ox
is over four years old due to the need for training and to allow it to grow to full size. Oxen are used for plowing
, transport, hauling cargo, grain-grinding by trampling or by powering machines, irrigation
by powering pumps, and wagon
drawing. Oxen were commonly used to skid logs in forests, and sometimes still are, in low-impact, select-cut logging. Oxen are most often used in teams of two, paired, for light work such as cart
ing, with additional pairs added when more power is required, sometimes up to a total of 20 or more.
Oxen can be trained to respond to a teamster
's signals. These signals are given by verbal commands or by noise (whip cracks). Verbal commands vary according to dialect and local tradition. Oxen can pull harder and longer than horses. Though not as fast as horses, they are less prone to injury because they are more sure-footed.
Many oxen are used worldwide, especially in developing countries
. About 11.3 million draft oxen are used in sub-Saharan Africa. In India, the number of draft cattle in 1998 was estimated at 65.7 million head. About half the world's crop production is thought to depend on land preparation (such as plowing) made possible by animal traction.
Religion, traditions and folklore
The cow is mentioned often in the Quran. The second and longest surah of the Quran is named Al-Baqara
("The Cow"). Out of the 286 verses of the surah, seven mention cows (Al Baqarah 67–73). The name of the surah derives from this passage in which Moses
orders his people to sacrifice a cow in order to resurrect a man murdered by an unknown person.
Cattle are venerated within the Hinduism|Hindu
religion of India. In the Vedic period
they were a symbol of plenty
and were frequently slaughtered. In later times they gradually acquired their present status. According to the ''Mahabharata
'', they are to be treated with the same respect 'as one's mother'. In the middle of the first millennium, the consumption of beef began to be disfavoured by lawgivers.
Although there has never been any cow-goddesses or temples dedicated to them,
cows appear in numerous stories from the Veda
s and Purana
s. The deity Krishna
was brought up in a family of cowherders, and given the name Govinda
(protector of the cows). Also, Shiva
is traditionally said to ride on the back of a bull named Nandi
Milk and milk products were used in Vedic rituals.
In the postvedic period products of the cow—milk, curd, ghee, but also cow dung and urine (gomutra
), or the combination of these five (panchagavya
)—began to assume an increasingly important role in ritual purification
Veneration of the cow has become a symbol of the identity of Hindus as a community,
especially since the end of the 19th century. Slaughter of cows (including oxen, bulls and calves) is forbidden by law in several states of the Indian Union. McDonald's
outlets in India do not serve any beef burgers. In Maharaja Ranjit Singh
's empire of the early 19th century, the killing of a cow was punishable by death.
thumb|upright=0.9| An idealized depiction of girl cow herders in 19th-century Norway by Knud Bergslien
* The Evangelist St. Luke
is depicted as an ox in Christian art
* In Judaism
, as described in , the ashes of a sacrificed unblemished red heifer
that has never been yoked can be used for ritual purification
of people who came into contact with a corpse.
* The ox is one of the 12-year cycle of animals which appear in the Chinese zodiac
related to the Chinese calendar
. See: Ox (Zodiac)
* The constellation Taurus
represents a bull.
* An apocryphal story has it that a cow started the Great Chicago Fire
by kicking over a kerosene
lamp. Michael Ahern, the reporter who created the cow story, admitted in 1893 that he had fabricated it for more colorful copy.
* On 18 February 1930, Elm Farm Ollie
became the first cow to fly in an airplane and also the first cow to be milked in an airplane.
* The first known law requiring branding in North America was enacted on 5 February 1644, by Connecticut. It said that all cattle and pigs had to have a registered brand or earmark by 1 May 1644.
* The is a traditional toy from the Aizu
region of Japan that is thought to ward off illness.
* The case of ''Sherwood v. Walker
''—involving a supposedly barren heifer that was actually pregnant—first enunciated the concept of mutual mistake
as a means of destroying the meeting of the minds
in contract law.
* The Fulani
of West Africa are the world's largest nomadic cattle-herders.
* The Maasai tribe
of East Africa traditionally believe their god Engai
entitled them to divine rights to the ownership of all cattle on earth.
[Patrick Mendis 2007. Glocalization: The Human Side of Globalization.. p160]
Cattle are typically represented in heraldry
by the bull.
File:Coat of arms of the Azores.svg|Arms of the Azores
File:Mecklenburg Arms.svg|Arms of Mecklenburg region, Germany
File:Turin coat of arms.svg|Arms of Turin, Italy
File:Coat of arms of Kaunas.svg|Arms of Kaunas, Lithuania
File:POL Bielsk Podlaski COA.svg|Arms of Bielsk Podlaski, Poland
File:POL COA Ciołek.svg|Arms of Ciołek, Poland
File:POL Turek COA PioM.svg|Arms of Turek, Poland
For 2013, the FAO estimated global cattle numbers at 1.47 billion.
[FAOSTAT. gricultural statistics databaseFood and Agriculture Organization of the United Nations, Rome. ]
Regionally, the FAO estimate for 2013 includes: Asia 497 million; South America 350 million; Africa 307 million; Europe 122 million; North America 102 million; Central America 47 million; Oceania 40 million; and Caribbean 9 million.
Didactic model of Bovine-FMVZ USP-23.jpeg|Didactic model of Bovine
Modelo didático bovino (fundo branco).jpg|Bovine anatomical model
Didactic model of a bovine muscular system-FMVZ USP-24.jpeg|Didactic model of a bovine muscular system
* 1966 anti-cow slaughter agitation
* :Category:Individual cattle
* British Cattle Health Initiative
* Bulls and Cows
* Cattle age determination
* Intensive animal farming
* List of cattle breeds
* List of domesticated animals
* Bhattacharya, S. 2003Cattle ownership makes it a man's world
''Newscientist.com''. Retrieved 26 December 2006.
* Cattle Today (CT). 2006. WebsiteBreeds of cattle
''Cattle Today''. Retrieved 26 December 2006
* Clay, J. 2004. ''World Agriculture and the Environment: A Commodity-by-Commodity Guide to Impacts and Practices''. Washington, DC: Island Press. .
* Clutton-Brock, J. 1999. ''A Natural History of Domesticated Mammals''. Cambridge: Cambridge University Press. .
* – A visual textbook containing History/Origin, Phenotype & Statistics of 45 breeds.
* Huffman, B. 2006''The ultimate ungulate page''
''UltimateUngulate.com''. Retrieved 26 December 2006.
* Invasive Species Specialist Group (ISSG). 2005''Bos taurus''
''Global Invasive Species Database''.
* Johns, Catherine. 2011 ''Cattle: History, Myth, Art''. London: The British Museum Press. 978-0-7141-5084-0
* Nowak, R.M. and Paradiso, J.L. 1983. ''Walker's Mammals of the World''. Baltimore, MD: The Johns Hopkins University Press.
* Oklahoma State University (OSU). 2006''Breeds of Cattle''
Retrieved 5 January 2007.
* Public Broadcasting Service (PBS). 2004
''PBS Nature''. Retrieved 5 January 2007.
* Rath, S. 1998. ''The Complete Cow''. Stillwater, MN: Voyageur Press. .
* Raudiansky, S. 1992. ''The Covenant of the Wild''. New York: William Morrow and Company, Inc. .
* Spectrum Commodities (SC). 2006
''Spectrumcommodities.com''. Retrieved 5 January 2007.
* Voelker, W. 1986. ''The Natural History of Living Mammals''. Medford, NJ: Plexus Publishing, Inc. .
* Yogananda, P. 1946. ''The Autobiography of a Yogi''. Los Angeles: Self Realization Fellowship. .
Category:Mammals described in 1758
Category:Articles containing video clips
Category:Taxa named by Carl Linnaeus