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The bean weevils or seed beetles are a subfamily (Bruchinae) of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, now placed in the family Chrysomelidae, though they have historically been treated as a separate family. They are
granivore Seed predation, often referred to as granivory, is a type of plant-animal interaction in which granivores (seed predators) feed on the seeds of plants as a main or exclusive food source,Hulme, P.E. and Benkman, C.W. (2002) "Granivory", pp. 13 ...
s, and typically infest various kinds of
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s or beans, living most of their lives inside a single seed. The subfamily includes about 1,650 species and are found worldwide. Bean weevils are generally compact and oval in shape, with small heads somewhat bent under. Sizes range from 1 to 22 mm for some tropical species. Colors are usually black or brown, often with mottled patterns. Although their mandibles may be elongated, they do not have the long snouts characteristic of true
weevil Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...
s. Adults deposit eggs on seeds, then the larvae chew their way into the seed. When ready to
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in thei ...
te, the larvae typically cut an exit hole, then return to their feeding chamber. Adult weevils have a habit of feigning death and dropping from a plant when disturbed. Host plants tend to be legumes, but species will also be found in
Convolvulaceae Convolvulaceae (), commonly called the bindweeds or morning glories, is a family of about 60 genera and more than 1,650 species. These species are primarily herbaceous vines, but also include trees, shrubs and herbs. The tubers of several spe ...
, Arecaceae, and
Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and durian. There are also some genera containing familiar ...
, and several species are considered pests. One characteristic of the beetles which can be seen in the photo is that the
elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...
are short, not quite reaching the tip of the abdomen. Several species are native to Great Britain, but there are also records of several
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
from stored products in warehouses and dwellings, although these species cannot proliferate outside of heated buildings in that climate.


Genera

This list of genera uses the new classification, treating the bean weevils as a subfamily with six extant tribes, eight or nine subtribes, and one extinct tribe. The former names and ranks for the tribes and subtribes under the old classification, treating the been weevils as a family, are given in parentheses (except for Myanmaropini, which was established only for the new classification). * Tribe Amblycerini Bridwell, 1932 (= Subfamily Amblycerinae) ** Subtribe Amblycerina Bridwell, 1932 (= Tribe Amblycerini) *** '' Amblycerus'' Thunberg, 1815 ** Subtribe Spermophagina Borowiec, 1987 (= Tribe Spermophagini) *** '' Spermophagus'' Schoenherr, 1833 *** '' Zabrotes'' Horn, 1885 * Tribe Bruchini Latreille, 1802 (= Subfamily Bruchinae) ** Subtribe Acanthoscelidina Bridwell, 1946 (= Tribe Acanthoscelidini) *** '' Abutiloneus'' Bridwell, 1946 *** '' Acanthoscelides'' Schilsky, 1905 *** '' Algarobius'' Bridwell, 1946 *** '' Althaeus'' Bridwell, 1946 *** '' Bonaerius'' Bridwell, 1952 *** '' Caryedes'' Hummel, 1827 *** '' Cosmobruchus'' Bridwell, 1931 *** '' Ctenocolum'' Kingsolver & Whitehead, 1974 *** '' Dahlibruchus'' Bridwell, 1931 *** '' Gibbobruchus'' Pic, 1913 *** '' Lithraeus'' Bridwell, 1952 *** '' Margaritabruchus'' Romero & Johnson, 2001 *** '' Meibomeus'' Bridwell, 1946 *** '' Megasennius'' Whitehead & Kingsolver, 1975 *** '' Merobruchus'' Bridwell, 1946 *** '' Mimosestes'' Bridwell, 1946 *** '' Neltumius'' Bridwell, 1946 *** '' Neobruchidius'' Johnson & Romero, 2006 *** '' Palpibruchus'' Borowiec, 1987 *** '' Pectinibruchus'' Kingsolver, 1967 *** '' Penthobruchus'' Kingsolver, 1973 *** '' Pseudopachymerina'' Zacher, 1952 *** '' Pygiopachymerus'' Pic, 1911 *** '' Rhipibruchus'' Bridwell, 1932 *** '' Scutobruchus'' Kingsolver, 1968 *** '' Sennius'' Bridwell, 1946 *** '' Spatulobruchus'' Borowiec, 1987 *** '' Stator'' Bridwell, 1946 *** '' Stylantheus'' Bridwell, 1946 ** Subtribe Bruchidiina Bridwell, 1946 (= Tribe Bruchidiini) *** '' Acanthobruchidius'' Borowiec, 1980 *** '' Borowiecus'' Anton, 1994 *** '' Bruchidius'' Schilsky, 1905 *** '' Callosobruchus'' Pic, 1902 *** '' Conicobruchus'' Decelle, 1951 *** '' Decellebruchus'' Borowiec, 1987 *** '' Horridobruchus'' Borowiec, 1984 *** '' Kingsolverius'' Borowiec, 1987 *** '' Megabruchidius'' Borowiec, 1984 *** '' Palaeoacanthoscelides'' Borowiec, 1985 *** '' Parasulcobruchus'' Anton, 1999 *** '' Pygobruchidius'' Pic, 1951 *** '' Salviabruchus'' Decelle, 1982 *** '' Specularius'' Bridwell, 1938 *** '' Sulcobruchus'' Chûjô, 1937 *** '' Tuberculobruchus'' Decelle, 1951 ** Subtribe Bruchina Latreille, 1802 (= Tribe Bruchini) *** '' Bruchus'' Linnaeus, 1767 ** Subtribe Megacerina Bridwell, 1946 (= Tribe Megacerini) *** '' Megacerus'' Fåhraeus, 1839 * Tribe Eubaptini Bridwell, 1932 (= Subfamily Eubaptinae) ** '' Eubaptus'' Lacordaire, 1945 * Tribe Kytorhinini Bridwell, 1832 (= Subfamily Kytorhininae) ** '' Kytorhinus'' Fischer von Waldheim, 1809 * Tribe Pachymerini Bridwell, 1929 (= Subfamily Pachymerinae) ** Subtribe Caryedontina Bridwell, 1929 (= Tribe Caryedontini) *** '' Aforedon'' Decelle, 1965 *** '' Caryedon'' Schoenherr, 1823 *** '' Caryotrypes'' Decelle, 1968 *** '' Exoctenophorus'' Decelle, 1968 *** '' Mimocaryedon'' Decelle, 1968 ** Subtribe Caryopemina Bridwell, 1929 (= Tribe Caryomepini) *** '' Caryopemon'' Jekel, 1855 *** '' Diedobruchus'' Pic, 1913 *** '' Protocaryopemon'' Borowiec, 1987 ** Subtribe Pachymerina Bridwell, 1929 (= Tribe Pachymerini) *** '' Butiobruchus'' Prevett, 1966 *** '' Caryobruchus'' Bridwell, 1929 *** '' Caryoborus'' Schoenherr, 1833 *** '' Pachymerus'' Thunberg, 1805 ***†'' Mesopachymerus'' Poinar, 2005
Canadian amber Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...
,
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
* Tribe
Rhaebini ''Rhaebus'' (from the Greek: ' curved) is a genus of metallic bean weevil, bean weevils in the subfamily Bean weevil, Bruchinae, and the only member of the tribe Rhaebini.Vencl, Fredric & Leschen, Richard. (2014). Vencl FV & Leschen RAB (2014) Cha ...
Chapuis, 1874 (= Subfamily Rhaebinae) ** '' Rhaebus'' Fischer von Waldheim, 1824 * Tribe † Myanmaropini Legalov ''et al.'', 2020 **†'' Myanmarops'' Legalov ''et al.'', 2020
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, Cenomanian


See also

*''
Caryobruchus gleditsiae ''Caryobruchus gleditsiae'' is a species of beetle in the family Chrysomelidae (formerly Bruchidae). It lives in North America, North and Central America and develops inside the seeds of Arecaceae, palm trees. Adults grow to a maximum length of , ...
''


Notes


References

* John M. Kingsolver, "Bruchidae", in
Ross H. Arnett, Jr. Ross Harold Arnett Jr. (April 13, 1919 – July 16, 1999) was an American entomologist noted for his studies of beetles, and as founder of the ''Coleopterist's Bulletin''. Born in Medina, New York, he was a star student at Cornell University, wher ...
and Michael C. Thomas, '' American Beetles'' (CRC Press, 2002), vol. 2 (note that this reference itself indicates the group is a subfamily, and the change was made only after the chapter was written)


External links


BRUCHBASE
{{Taxonbar, from1=Q217617, from2=Q13167641 Storage pests