Castanea mollissima
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''Castanea mollissima'' (), also known as the Chinese chestnut, is a member of the family
Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as ever ...
, and a species of
chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrel ...
native to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...
, and
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic ...
.


Description

It is a
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to 20 m tall with a broad crown. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are alternate, simple, 10–22 cm long and 4.5–8 cm broad, with a toothed margin. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s are produced in
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
s 4–20 cm long, with the female flowers at the base of the catkin and males on the rest. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a densely spiny cupule 4–8 cm diameter, containing two or three glossy brown nuts; these are 2–3 cm diameter on wild trees. The scientific name ''mollissima'' derives from the softly downy shoots and young leaves.


Taxonomy

Synonyms: ''Castanea bungeana'' Blume; ''C. duclouxii'' Dode; ''C. fargesii'' Dode; ''C. formosana'' (Hayata) Hayata; ''C. hupehensis'' Dode; ''C. mollissima'' var. ''pendula'' X. Y. Zhou & Z. D. Zhou; ''C. sativa'' Miller var. ''formosana'' Hayata; ''C. sativa'' var.'' mollissima'' (Blume) Pampanini; ''C. vulgaris'' Lamarck var. ''yunnanensis'' Franchet. In
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making ...
, Chinese chestnut (
Vietnamese language Vietnamese ( vi, tiếng Việt, links=no) is an Austroasiatic language originating from Vietnam where it is the national and official language. Vietnamese is spoken natively by over 70 million people, several times as many as the rest of the ...
: ''hạt dẻ'', Tày language: ''mác lịch'') which are grown in Trùng Khánh district, Cao Bằng province have highest quality with 3.3-5.4%
glucose Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, u ...
, 43.36- 46.47% glucid, 1.16 – 2%
lipid Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids in ...
, 3.12 – 3.62%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
analyzed by Vietnam National Vegetable and Fruit Researching Institution in 1999.


Distribution and habitat

Chinese chestnut has been cultivated in East Asia for millennia and its exact original range cannot be determined. In the provinces of
Anhui Anhui , (; formerly romanized as Anhwei) is a landlocked province of the People's Republic of China, part of the East China region. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze Riv ...
,
Fujian Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its ...
,
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibe ...
,
Guangdong Guangdong (, ), alternatively romanized as Canton or Kwangtung, is a coastal province in South China on the north shore of the South China Sea. The capital of the province is Guangzhou. With a population of 126.01 million (as of 2020 ...
,
Guangxi Guangxi (; ; alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an autonomous region of the People's Republic of China, located in South China and bordering Vietnam ...
,
Guizhou Guizhou (; Postal romanization, formerly Kweichow) is a landlocked Provinces of China, province in the Southwest China, southwest region of the China, People's Republic of China. Its capital and largest city is Guiyang, in the center of the pr ...
,
Hebei Hebei or , (; alternately Hopeh) is a northern province of China. Hebei is China's sixth most populous province, with over 75 million people. Shijiazhuang is the capital city. The province is 96% Han Chinese, 3% Manchu, 0.8% Hui, and ...
,
Henan Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is a ...
,
Hubei Hubei (; ; alternately Hupeh) is a landlocked province of the People's Republic of China, and is part of the Central China region. The name of the province means "north of the lake", referring to its position north of Dongting Lake. The p ...
,
Hunan Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangx ...
,
Jiangsu Jiangsu (; ; pinyin: Jiāngsū, alternatively romanized as Kiangsu or Chiangsu) is an eastern coastal province of the People's Republic of China. It is one of the leading provinces in finance, education, technology, and tourism, with it ...
,
Jiangxi Jiangxi (; ; formerly romanized as Kiangsi or Chianghsi) is a landlocked province in the east of the People's Republic of China. Its major cities include Nanchang and Jiujiang. Spanning from the banks of the Yangtze river in the north int ...
,
Liaoning Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmo ...
, Nei Mongol,
Qinghai Qinghai (; alternately romanized as Tsinghai, Ch'inghai), also known as Kokonor, is a landlocked province in the northwest of the People's Republic of China. It is the fourth largest province of China by area and has the third smallest po ...
,
Shaanxi Shaanxi (alternatively Shensi, see § Name) is a landlocked province of China. Officially part of Northwest China, it borders the province-level divisions of Shanxi (NE, E), Henan (E), Hubei (SE), Chongqing (S), Sichuan (SW), Gansu (W), N ...
,
Shandong Shandong ( , ; ; Chinese postal romanization, alternately romanized as Shantung) is a coastal Provinces of China, province of the China, People's Republic of China and is part of the East China region. Shandong has played a major role in His ...
,
Shanxi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-leve ...
,
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of t ...
, Xizang,
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
, and
Zhejiang Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Ji ...
, and also to
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the no ...
and
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic ...
. It grows close to sea level in the north of its range, and at altitudes of up to 2,800 m in the south of the range. The species prefers full sun and acidic, loamy
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
, and has a medium growth rate.Flora of Taiwan
''Castanea mollissima''
/ref>


Ecology

When cultivated close to other species of chestnut (including Japanese chestnut, ''C. crenata'';
American chestnut The American chestnut (''Castanea dentata'') is a large, fast-growing deciduous tree of the beech family native to eastern North America. As is true of all species in genus Castanea, the American chestnut produces burred fruit with edible nuts. ...
, ''C. dentata''; and sweet chestnut, ''C. sativa''), Chinese chestnut readily cross-pollinates with them to form hybrids.Sisco, P. H., ''et al.'' 2005 An Improved Genetic Map for ''Castanea mollissima''/''Castanea dentata'' and its Relationship to the Genetic Map of ''Castanea sativa''. ''Acta Hort''. 693
Abstract.
/ref> Chinese chestnuts have evolved over a long period of time in coexistence with the bark fungal disease
chestnut blight The pathogenic fungus ''Cryphonectria parasitica'' (formerly ''Endothia parasitica'') is a member of the Ascomycota (sac fungi). This necrotrophic fungus is native to East Asia and South East Asia and was introduced into Europe and North America ...
(''Cryphonectria parasitica'', formerly ''Endothia parasitica''), and have evolved a very successful resistance to the blight, probably more so than any other species of chestnut, so that, although it is not immune, it typically sustains no more than minor damage when infected. It's important to realize, though, that Chinese chestnut trees vary considerably in blight resistance. Some individuals are quite susceptible while others are essentially immune to the disease. Japanese chestnut is also comparatively resistant to blight, with European chestnut somewhat less so. In the 1890s, Chinese and Japanese chestnuts were imported to the United States with the intention of utilizing them as orchard trees due to their small, compact size compared to the towering American chestnut. The results unfortunately were disastrous as the imported Asian species introduced blight to which ''C. dentata'' lacked any resistance. The disease was first noticed on a tree in the Bronx Zoo in 1904 and quickly spread out of control, ravaging American chestnut trees. Within 30 years, there were virtually none left in their native range. An active program has been pursued in North America to cross-breed the Chinese and American chestnuts to try to maximize various desirable traits of the American chestnut, such as larger stature, greater nut sweetness, while also isolating and carrying the blight resistance from the Chinese chestnut.FAO
Chestnut blight
/ref>


Uses

The nuts are edible, and the tree is widely cultivated in eastern
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
; over 300
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s have been selected for nut production, subdivided into five major regional groups: Northern,
Yangtze River The Yangtze or Yangzi ( or ; ) is the longest river in Asia, the third-longest in the world, and the longest in the world to flow entirely within one country. It rises at Jari Hill in the Tanggula Mountains (Tibetan Plateau) and flows ...
Valley,
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of t ...
and
Guizhou Guizhou (; Postal romanization, formerly Kweichow) is a landlocked Provinces of China, province in the Southwest China, southwest region of the China, People's Republic of China. Its capital and largest city is Guiyang, in the center of the pr ...
, Southern, and Southwestern. Besides that, the Dandong chestnut (belonging to the Japanese chestnut – ''Castanea crenata'') is a major cultivar in
Liaoning Liaoning () is a coastal province in Northeast China that is the smallest, southernmost, and most populous province in the region. With its capital at Shenyang, it is located on the northern shore of the Yellow Sea, and is the northernmo ...
Province.Economic forest trees
.
Some cultivars, such as 'Kuling', 'Meiling', and 'Nanking', have large nuts up to 4 cm diameter. The nuts are sweet, and considered by some to have the best taste of any chestnut,Biodiversity of China

though others state they are not as good as the
American chestnut The American chestnut (''Castanea dentata'') is a large, fast-growing deciduous tree of the beech family native to eastern North America. As is true of all species in genus Castanea, the American chestnut produces burred fruit with edible nuts. ...
.''Castanea mollissima: Chinese chestnut''
By Edward F. Gilman and Dennis G. Watson. This is one of a series of documents from the Environmental Horticulture Department, Florida Cooperative Extension Service
Institute of Food and Agricultural Sciences
University of Florida. Original publication date November 1993. Revised December 2006.
The nuts also provide a significant food source for
wildlife Wildlife refers to undomesticated animal species, but has come to include all organisms that grow or live wild in an area without being introduced by humans. Wildlife was also synonymous to game: those birds and mammals that were hunted ...
.


References


Citations


Sources

*


External links

* {{Taxonbar , from = Q699584 mollissima Edible nuts and seeds Crops originating from China Trees of China Trees of Korea Trees of Taiwan Trees of Vietnam