Carpinus betulus
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''Carpinus betulus'', the European or common hornbeam, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of tree in the birch
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Betulaceae,
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to Western Asia and central, eastern, and southern Europe, including southern England. It requires a warm climate for good growth, and occurs only at elevations up to . It grows in mixed stands with oak, and in some areas
beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engl ...
, and is also a common tree in scree forests. Hornbeam was also known as 'Yoke Elm'.


Description

It is a deciduous small to medium-size
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
reaching heights of , rarely , and often has a fluted and crooked
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
. The bark is smooth and greenish-grey, even in old trees. The buds, unlike those of the beech, are long at the most, and pressed close to the twig. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are alternate, long, with prominent veins giving a distinctive corrugated texture, and a serrated margin. It is
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is ...
, and the wind-pollinated male and female
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in '' Salix''). They contain many, usually unisexual flowers, arrang ...
s appear in early summer after the leaves. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a small long nut, partially surrounded by a three-pointed leafy
involucre In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or o ...
long; it matures in autumn. The wood is heavy and hard, and is used for tools and building constructions. It also burns hot and slowly, making it very suitable for firewood. This was the reason for lopping and hence indirectly the saving of
Epping Forest Epping Forest is a area of ancient woodland, and other established habitats, which straddles the border between Greater London and Essex. The main body of the forest stretches from Epping in the north, to Chingford on the edge of the London ...
, where the hornbeam was a favoured pollarding tree. Hornbeam was frequently coppiced and
pollarded Pollarding is a pruning system involving the removal of the upper branches of a tree, which promotes the growth of a dense head of foliage and branches. In ancient Rome, Propertius mentioned pollarding during the 1st century BCE. The practice oc ...
in the past in England. It is still infrequently managed using these traditional methods, but mainly for non-commercial conservation purposes. As a woodland tree traditionally managed in this way, it is particularly frequent in the
ancient woodland In the United Kingdom, an ancient woodland is a woodland that has existed continuously since 1600 or before in England, Wales and Northern Ireland (or 1750 in Scotland). Planting of woodland was uncommon before those dates, so a wood present in 16 ...
s of south
Essex Essex () is a Ceremonial counties of England, county in the East of England. One of the home counties, it borders Suffolk and Cambridgeshire to the north, the North Sea to the east, Hertfordshire to the west, Kent across the estuary of the Riv ...
,
Hertfordshire Hertfordshire ( or ; often abbreviated Herts) is one of the home counties in southern England. It borders Bedfordshire and Cambridgeshire to the north, Essex to the east, Greater London to the south, and Buckinghamshire to the west. For gov ...
and north
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it faces ...
where it typically occupies more than half of most ancient woods and
wood pasture Silvopasture (''silva'' is forest in Latin) is the practice of integrating trees, forage, and the grazing of domesticated animals in a mutually beneficial way. It utilizes the principles of managed grazing, and it is one of several distinct forms ...
s. Because it stands up well to cutting back and has dense foliage, it has been much used in
landscape gardening Landscape architecture is the design of outdoor areas, landmarks, and structures to achieve environmental, social-behavioural, or aesthetic outcomes. It involves the systematic design and general engineering of various structures for constructio ...
, mainly as tall hedges and for
topiary Topiary is the horticultural practice of training perennial plants by clipping the foliage and twigs of trees, shrubs and subshrubs to develop and maintain clearly defined shapes, whether geometric or fanciful. The term also refers to plants w ...
. It was the classic tree used in
French formal garden The French formal garden, also called the (), is a style of garden based on symmetry and the principle of imposing order on nature. Its epitome is generally considered to be the Gardens of Versailles designed during the 17th century by the ...
s for hedges in
bosquet In the French formal garden, a ''bosquet'' (French, from Italian ''bosco'', "grove, wood") is a formal plantation of trees in a wide variety of forms, some open at the bottom and others not. At a minimum a bosquet can be five trees of identical s ...
s, as in the
Gardens of Versailles The Gardens of Versailles (french: Jardins du château de Versailles ) occupy part of what was once the ''Domaine royal de Versailles'', the royal demesne of the château of Versailles. Situated to the west of the palace, the gardens cover so ...
, and in their English equivalent, the
garden wilderness In the Western history of gardening, from the 16th to early 19th centuries, a wilderness was a highly artificial and formalized type of woodland, forming a section of a large garden. Though examples varied greatly, a typical English style was a ...
. The leaves provide food for some animals, including
Lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described speci ...
such as the
case-bearer __NOTOC__ The Coleophoridae are a family of small moths, belonging to the huge superfamily Gelechioidea. Collectively known as case-bearers, casebearing moths or case moths, this family is represented on all continents, but the majority are foun ...
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
''
Coleophora anatipennella ''Coleophora anatipennella'' is a moth of the case-bearer family (Coleophoridae). Taxonomy It was described by Jacob Hübner in 1796. It is the type species of its genus (''Coleophora'') and, via that, of its family. It is not completely und ...
''. There are a number of notable forests where ''C. betulus'' is a dominant tree species, among which are: * Epping Forest, Essex/London, UK * Halltorp Nature Reserve,
Öland Öland (, ; ; sometimes written ''Øland'' in other Scandinavian languages, and often ''Oland'' internationally; la, Oelandia) is the second-largest Swedish island and the smallest of the traditional provinces of Sweden. Öland has an area ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...


Ecology

In England, trees appear to prefer soils with a pH from 3.6 to 4.6 but tolerate up to 7.6. They are found on soils with moderate clay content and avoid soils with particularly high or low clay content. ''Carpinus betulus'' likes full sun or partial shade, moderate
soil fertility Soil fertility refers to the ability of soil to sustain agricultural plant growth, i.e. to provide plant habitat and result in sustained and consistent yields of high quality.
and moisture. It has a shallow, wide-spreading root system and is marked by the production of stump sprouts when cut back. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s often do not germinate till the spring of the second year after sowing. The hornbeam is a prolific seeder and is marked by vigorous natural regeneration.


Fossil record

Three
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
fruits of ''Carpinus betulus'' have been extracted from
borehole A borehole is a narrow shaft bored in the ground, either vertically or horizontally. A borehole may be constructed for many different purposes, including the extraction of water ( drilled water well and tube well), other liquids (such as petrol ...
samples of the
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million ...
fresh water deposits in Nowy Sacz Basin,
West Carpathians The Western Carpathians are a mountain range and geomorphological province that forms the western part of the Carpathian Mountains. The mountain belt stretches from the Low Beskids range of the Eastern Carpathians along the border of Poland wit ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
.


Cultivation & uses

''Carpinus betulus'' is widely cultivated as an
ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
, for planting in gardens and parks throughout north west Europe. There are several
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s, notably: * ''C. betulus'' 'Fastigiata' or 'Pyramidalis', a very fastigiate tree when young, which has become a popular urban
street tree Urban forestry is the care and management of single trees and tree populations in urban settings for the purpose of improving the urban environment. Urban forestry involves both planning and management, including the programming of care and ...
in the United Kingdom and other countries. * ''C. betulus'' 'Frans Fontaine', a similar fastigiate tree to 'Fastigiata' Both the species and the cultivar ''C. betulus'' 'Fastigiata' have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


Notable examples

*''The Last Tree'' - The only tree that survived the devastation of the
Battle of Delville Wood The Battle of Delville Wood was a series of engagements in the 1916 Battle of the Somme in the First World War, between the armies of the German Empire and the British Empire. Delville Wood , was a thick tangle of trees, chiefly beech and ...
in 1916. It is preserved as part of the Delville Wood South African National Memorial near
Longueval Longueval () is a commune in the Somme department in Hauts-de-France in northern France. Geography Longueval is located northwest of Amiens on the D919 road, at the junction with the D8. Longueval is found in the north-east of the départemen ...
. *''Lincoln's Hornbeam'' - Planted by
President Lincoln Abraham Lincoln ( ; February 12, 1809 – April 15, 1865) was an American lawyer, politician, and statesman who served as the 16th president of the United States from 1861 until his assassination in 1865. Lincoln led the nation thro ...
at the
United States Botanic Garden The United States Botanic Garden (USBG) is a botanical garden on the grounds of the United States Capitol in Washington, D.C., near Garfield Circle. The Botanic Garden is supervised by the Congress through the Architect of the Capitol, who ...
in the Capitol grounds, Washington D.C.


Gallery

Image:CarpinusBetulusBark.jpg, Bark of a mature tree File:Herchies JPG09.jpg, Tree in winter File:Carpinus betulus trunk.jpg, Veteran tree File:La Reid AR1aJPG.jpg, Hornbeam-covered walk () File:Carpinus betulus MHNT.BOT.2004.0.494.jpg, Fruit and seeds File:Carpinus betulus bud.jpg, Leaf bud File:Carpinus betulus 'Fastigiata' Dublin P1310089.jpg, Carpinus betulus 'Fastigiata' in
Dublin Dublin (; , or ) is the capital and largest city of Ireland. On a bay at the mouth of the River Liffey, it is in the province of Leinster, bordered on the south by the Dublin Mountains, a part of the Wicklow Mountains range. At the 2016 ...
, Ireland where it is a common street tree File:Carpinus betulus 'Fastigiata' Eindhoven 21 MG 3470.jpg, Mature Carpinus betulus 'Fastigiata' in
Eindhoven Eindhoven () is a city and municipality in the Netherlands, located in the southern province of North Brabant of which it is its largest. With a population of 238,326 on 1 January 2022,Poundbury Poundbury is an experimental planned community or urban development, urban extension on the western outskirts of Dorchester, Dorset, Dorchester in the Counties of England, county of Dorset, England. The development is led by the Duchy of Cornwal ...
, Dorset, UK


References


External links


Den virtuella floran: ''Carpinus betulus'' distribution''Carpinus betulus''
- information, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote '' in situ'' an ...
(EUFORGEN) {{Authority control betulus Flora of Europe Flora of Western Asia Flora of England Flora of Italy Trees of Asia Trees of Europe Plants described in 1753 Taxa named by Carl Linnaeus Garden plants of Asia Garden plants of Europe Ornamental trees