Carcharhinid
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Requiem sharks are
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...
s of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Carcharhinidae in the order
Carcharhiniformes Carcharhiniformes , the ground sharks, are the largest order of sharks, with over 270 species. They include a number of common types, such as catsharks, swellsharks, and the sandbar shark. Members of this order are characterized by the presen ...
. They are migratory, live-bearing sharks of warm seas (sometimes of
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estu ...
or fresh water) and include such species as the
tiger shark The tiger shark (''Galeocerdo cuvier'') is a species of requiem shark and the last extant member of the family Galeocerdonidae. It is a large macropredator, capable of attaining a length over . Populations are found in many tropical and tempera ...
,
bull shark The bull shark (''Carcharhinus leucas''), also known as the Zambezi shark (informally zambi) in Africa and Lake Nicaragua shark in Nicaragua, is a species of requiem shark commonly found worldwide in warm, shallow waters along coasts and in ri ...
,
lemon shark The lemon shark (''Negaprion brevirostris'') is a species of shark from the family Carcharhinidae and is classified as a Vulnerable species by the International Union for the Conservation of Nature. Lemon sharks can grow to in length. They are ...
,
spinner shark The spinner shark (''Carcharhinus brevipinna'') is a species of requiem shark, in the family Carcharhinidae, named for the spinning leaps it makes as a part of its feeding strategy. This species occurs in tropical and warm temperate waters worldw ...
,
blacknose shark The blacknose shark (''Carcharhinus acronotus'') is a species of requiem shark, belonging to the family Carcharhinidae, common in the tropical and subtropical waters of the western Atlantic Ocean. This species generally inhabits coastal seagrass ...
, blacktip shark,
grey reef shark The grey reef shark (''Carcharhinus amblyrhynchos'', sometimes misspelled ''amblyrhynchus'' or ''amblyrhinchos'') is a species of requiem shark, in the family Carcharhinidae. One of the most common reef sharks in the Indo-Pacific, it is found as ...
, blacktip reef shark,
silky shark The silky shark (''Carcharhinus falciformis''), also known by numerous names such as blackspot shark, gray whaler shark, olive shark, ridgeback shark, sickle shark, sickle-shaped shark and sickle silk shark, is a species of requiem shark, in the ...
, dusky shark, blue shark,
copper shark The copper shark (''Carcharhinus brachyurus''), bronze whaler, or narrowtooth shark, is a species of requiem shark, family Carcharhinidae, and the only member of its genus found mostly at temperate latitudes. It is distributed in a number of sepa ...
,
oceanic whitetip shark The oceanic whitetip shark (''Carcharhinus longimanus''), also known as shipwreck shark, Brown Milbert's sand bar shark, brown shark, lesser white shark, nigano shark, oceanic white-tipped whaler, and silvertip shark, is a large pelagic requiem ...
, and
whitetip reef shark The whitetip reef shark (''Triaenodon obesus'') is a species of requiem shark, in the family Carcharhinidae, and the only member of its genus. A small shark that does not usually exceed in length, this species is easily recognizable by its slen ...
. Family members have the usual carcharhiniform characteristics. Their eyes are round, and one or two
gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
slits fall over the
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
base. Most species are
viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the ...
, the young being born fully developed. They vary widely in size, from as small as adult length in the Australian sharpnose shark, up to adult length in the
tiger shark The tiger shark (''Galeocerdo cuvier'') is a species of requiem shark and the last extant member of the family Galeocerdonidae. It is a large macropredator, capable of attaining a length over . Populations are found in many tropical and tempera ...
.Compagno, L.J.V
Family Carcharhinidae - Requiem sharks
in Froese, R. and D. Pauly. Editors. 2010
FishBase
World Wide Web electronic publication, version (10/2013).
Scientists assume that size and shape of their pectoral fins have the right dimensions to minimize transport cost. Requiem sharks tend to live in more tropical areas, but tend to migrate. Females release a chemical in the ocean in order to let the males know they are ready to mate. Typical mating time for these sharks are around spring to autumn. Requiem sharks are involved in a large proportion of attacks on humans, among the top five species; however, due to the difficulty in identifying individual species, a degree of inaccuracy exists in attack records.


Etymology

The common name requiem shark may be related to the French word for shark, ''requin'', which is itself of disputed etymology. One derivation of the latter is from Latin '' requiem'' ("rest"), which would thereby create a cyclic etymology (''requiem-requin-requiem''), but other sources derive it from the Old French verb ''reschignier'' ("to grimace while baring teeth"). The scientific name Carcharhinidae was first proposed in 1896 by D.S. Jordan and B.W. Evermann as a subfamily of Galeidae (now replaced by "Carcharhinidae"). The term is derived from Greek ''karcharos'' (sharp or jagged); and ''rhine'' (rasp), both elements describe the jagged, rasp-like skin. Rasp-like skin is typical of
shark skin Shagreen is a type of rawhide (textile) , rawhide consisting of rough untanned skin, historically from a horse's or onager's back, or from shark or sting ray, ray. Etymology The word derives from the French language, French ''chagrin'' and is ...
in general, and is not diagnostic to Carcharhinidae.


Evolutionary history

The oldest member of the family is '' Archaeogaleus lengadocensis'' from the Early Cretaceous (
Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretace ...
) of France. Only a handful of records of the group are known from prior to the beginning of the Cenozoic. Modern carcharinid sharks have extensively diversified in
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
habitats.


Hunting strategies

Requiem sharks are extraordinarily fast and effective hunters. Their elongated, torpedo-shaped bodies make them quick and agile swimmers, so they can easily attack any prey. They have a range of food sources depending on their location and species that includes bony fish, squids, octopuses, lobsters,
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked t ...
s,
marine mammals Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their re ...
,
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
, other
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...
s and
rays Ray may refer to: Fish * Ray (fish), any cartilaginous fish of the superorder Batoidea * Ray (fish fin anatomy), a bony or horny spine on a fin Science and mathematics * Ray (geometry), half of a line proceeding from an initial point * Ray (gra ...
. They are often considered the "garbage cans" of the seas because they will eat almost anything, even non-food items like trash. They are migratory hunters that follow their food source across entire oceans. They tend to be most active at night time, where their impressive eyesight can help them sneak up on unsuspecting prey. Most requiem sharks hunt alone, however some species like the
whitetip reef shark The whitetip reef shark (''Triaenodon obesus'') is a species of requiem shark, in the family Carcharhinidae, and the only member of its genus. A small shark that does not usually exceed in length, this species is easily recognizable by its slen ...
s and
lemon shark The lemon shark (''Negaprion brevirostris'') is a species of shark from the family Carcharhinidae and is classified as a Vulnerable species by the International Union for the Conservation of Nature. Lemon sharks can grow to in length. They are ...
s are cooperative feeders and will hunt in packs through coordinated, timed attacks against their prey.


Classification

The 60 species of requiem shark are grouped into 12 genera: * Genus '' Galeocerdo'' J. P. Müller &
Henle Henle can refer to: * Friedrich Gustav Jakob Henle, a German physician, pathologist and anatomist (1809–1885) ** Loop of Henle in the kidney, named after Henle *Fritz Henle, a photographer, known as "Mr. Rollei" for his use of the 2.25" square for ...
, 1837
** '' Galeocerdo cuvier'' Péron & Lesueur, 1822 (tiger shark) * Genus ''
Scoliodon ''Scoliodon'' is a genus of requiem sharks in the family Carcharhinidae. It was formerly thought to include only a single Indo-Pacific species, the spadenose shark (''S. laticaudus''), but recent taxonomic research has found an additional spe ...
'' J. P. Müller & Henle, 1838 ** ''
Scoliodon laticaudus The spadenose shark (''Scoliodon laticaudus'') is a species of requiem shark in the family Carcharhinidae. It is common in the tropical Indian and western Pacific Oceans, where it forms large schools in shallow water. A small shark reaching a len ...
'' J. P. Müller & Henle, 1838 (spadenose shark) ** '' Scoliodon macrorhynchos'' Bleeker, 1852 (Pacific spadenose shark) * Genus '' Carcharhinus'' Blainville, 1816 **'' Carcharhinus acronotus'' Poey, 1860 (blacknose shark) ** '' Carcharhinus albimarginatus'' Rüppell, 1837 (silvertip shark) ** '' Carcharhinus altimus'' S. Springer, 1950 (bignose shark) ** '' Carcharhinus amblyrhynchoides'' Whitley, 1934 (graceful shark) ** ''
Carcharhinus amblyrhynchos The grey reef shark (''Carcharhinus amblyrhynchos'', sometimes misspelled ''amblyrhynchus'' or ''amblyrhinchos'') is a species of requiem shark, in the family Carcharhinidae. One of the most common reef sharks in the Indo-Pacific, it is found as ...
'' Bleeker, 1856 (grey reef shark) ** '' Carcharhinus amboinensis'' J. P. Müller &
Henle Henle can refer to: * Friedrich Gustav Jakob Henle, a German physician, pathologist and anatomist (1809–1885) ** Loop of Henle in the kidney, named after Henle *Fritz Henle, a photographer, known as "Mr. Rollei" for his use of the 2.25" square for ...
, 1839
(pigeye shark) ** ''
Carcharhinus borneensis The Borneo shark (''Carcharhinus borneensis'') is a species of requiem shark, and part of the family Carcharhinidae. Extremely rare, it is known only from inshore waters around Mukah in northwestern Borneo, though it may once have been more wide ...
'' Bleeker, 1858 (Borneo shark) ** '' Carcharhinus brachyurus'' Günther, 1870 (copper shark) ** '' Carcharhinus brevipinna'' J. P. Müller & Henle, 1839 (spinner shark) ** '' Carcharhinus cautus'' Whitley, 1945 (nervous shark) ** '' Carcharhinus cerdale'' C. H. Gilbert, 1898 (Pacific smalltail shark) ** '' Carcharhinus coatesi'' Whitley, 1939 (Coates's shark) ** '' Carcharhinus dussumieri'' J. P. Müller & Henle, 1839 (whitecheek shark) ** ''
Carcharhinus falciformis The silky shark (''Carcharhinus falciformis''), also known by numerous names such as blackspot shark, gray whaler shark, olive shark, ridgeback shark, sickle shark, sickle-shaped shark and sickle silk shark, is a species of requiem shark, in the f ...
'' J. P. Müller & Henle, 1839 (silky shark) ** '' Carcharhinus fitzroyensis'' Whitley, 1943 (creek whaler) ** '' Carcharhinus galapagensis''
Snodgrass The family name Snodgrass is said to originate from lands in the parish of Irvine, Ayrshire, Scotland, known as Snodgrasse, or Snodgers, at a bend in the River Garnock at 55°38' north, 4°42' west, which were rented out in plots. Both forms are ...
& Heller, 1905
(Galapagos shark) ** '' Carcharhinus hemiodon'' J. P. Müller & Henle, 1839 (Pondicherry shark) ** '' Carcharhinus humani'' W. T. White & Weigmann, 2014 (Human's whaler shark) ** '' Carcharhinus isodon'' J. P. Müller & Henle, 1839 (finetooth shark) ** '' Carcharhinus leiodon''
Garrick Garrick may refer to: * Garrick (name), for the name's origin and people with either the surname or given name, the most famous being: ** David Garrick (1717–1779), English actor * Garrick Club, a London gentlemen's club named in honour of David ...
, 1985
(smoothtooth blacktip shark) ** '' Carcharhinus leucas'' J. P. Müller &
Henle Henle can refer to: * Friedrich Gustav Jakob Henle, a German physician, pathologist and anatomist (1809–1885) ** Loop of Henle in the kidney, named after Henle *Fritz Henle, a photographer, known as "Mr. Rollei" for his use of the 2.25" square for ...
, 1839
(bull shark) ** '' Carcharhinus limbatus'' J. P. Müller & Henle, 1839 (blacktip shark) ** '' Carcharhinus longimanus'' Poey, 1861 (oceanic whitetip shark) ** '' Carcharhinus macloti'' J. P. Müller & Henle, 1839 (hardnose shark) ** '' Carcharhinus melanopterus'' Quoy &
Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon and naturalist. Biography Gaimard was born at Saint-Zacharie on January 31, 1793. He studied medicine at the naval medical school in Toulon, subsequent ...
, 1824
(blacktip reef shark) ** '' Carcharhinus obscurus'' Lesueur, 1818 (dusky shark) ** '' Carcharhinus perezi'' Poey, 1876 (Caribbean reef shark) ** '' Carcharhinus plumbeus'' Nardo, 1827 (sandbar shark) ** '' Carcharhinus porosus''
Ranzani Camillo Ranzani (22 June 1775 – 23 April 1841, Bologna ) was an Italian Priesthood (Catholic Church), priest and a naturalist. He was director of the Museum of Natural History of Bologna from 1803 to 1841 (now the Museum of Comparative Anatomy, ...
, 1839
(smalltail shark) ** '' Carcharhinus sealei'' Pietschmann, 1913 (blackspot shark) ** '' Carcharhinus signatus'' Poey, 1868 (night shark) ** '' Carcharhinus sorrah'' J. P. Müller & Henle, 1839 (spot-tail shark) ** '' Carcharhinus tilstoni'' Whitley, 1950 (Australian blacktip shark) ** †'' Carcharhinus tingae'' ** '' Carcharhinus tjutjot'' Bleeker, 1852 (Indonesian whaler shark) ** '' Carcharhinus obsolerus''
White White is the lightest color and is achromatic (having no hue). It is the color of objects such as snow, chalk, and milk, and is the opposite of black. White objects fully reflect and scatter all the visible wavelengths of light. White o ...
, Kyne, and
Harris Harris may refer to: Places Canada * Harris, Ontario * Northland Pyrite Mine (also known as Harris Mine) * Harris, Saskatchewan * Rural Municipality of Harris No. 316, Saskatchewan Scotland * Harris, Outer Hebrides (sometimes called the Isle of ...
, 2019
(lost shark) * Genus '' Glyphis'' Agassiz, 1843 ** '' Glyphis gangeticus'' (J. P. Müller & Henle, 1839) (Ganges shark) ** '' Glyphis garricki'' Compagno, W. T. White & Last, 2008 (northern river shark) ** ''
Glyphis glyphis The speartooth shark (''Glyphis glyphis'') is a rare species of river shark, belonging to the family Carcharhinidae. It inhabits coastal marine waters and tidal reaches of large tropical rivers in northern Australia and New Guinea. Despite bei ...
'' (J. P. Müller & Henle, 1839) (speartooth shark) ** '' Glyphis'' sp. not yet described (Mukah river shark) * Genus '' Lamiopsis''
Gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
, 1862
** '' Lamiopsis temminckii'' (J. P. Müller & Henle, 1839) (broadfin shark) ** '' Lamiopsis tephrodes'' (Fowler, 1905) (Borneo broadfin shark) * Genus '' Nasolamia'' Compagno &
Garrick Garrick may refer to: * Garrick (name), for the name's origin and people with either the surname or given name, the most famous being: ** David Garrick (1717–1779), English actor * Garrick Club, a London gentlemen's club named in honour of David ...
, 1983
** '' Nasolamia velox'' (Gilbert, 1898) (whitenose shark) * Genus '' Negaprion'' Whitley, 1940 ** '' Negaprion acutidens'' (Rüppell, 1837) (sicklefin lemon shark) ** '' Negaprion brevirostris'' (Poey, 1868) (lemon shark) ** †'' Negaprion eurybathrodon'' (Blake, 1862) * Genus '' Prionace''
Cantor A cantor or chanter is a person who leads people in singing or sometimes in prayer. In formal Jewish worship, a cantor is a person who sings solo verses or passages to which the choir or congregation responds. In Judaism, a cantor sings and lead ...
, 1849
** '' Prionace glauca'' (
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758)
(blue shark) * Genus '' Rhizoprionodon'' Whitley, 1929 ** '' Rhizoprionodon acutus'' (Rüppell, 1837) (milk shark) ** '' Rhizoprionodon lalandii'' (J. P. Müller & Henle, 1839) (Brazilian sharpnose shark) ** '' Rhizoprionodon longurio'' ( D. S. Jordan & Gilbert, 1882) (Pacific sharpnose shark) ** '' Rhizoprionodon oligolinx'' V. G. Springer, 1964 (grey sharpnose shark) ** '' Rhizoprionodon porosus'' (Poey, 1861) (Caribbean sharpnose shark) ** '' Rhizoprionodon taylori'' ( Ogilby, 1915) (Australian sharpnose shark) ** '' Rhizoprionodon terraenovae'' ( J. Richardson, 1836) (Atlantic sharpnose shark) * Genus '' Loxodon'' J. P. Müller & Henle, 1838 ** '' Loxodon macrorhinus'' (J. P. Müller & Henle, 1839) (sliteye shark) * Genus ''
Isogomphodon The daggernose shark (''Isogomphodon oxyrhynchus'') is a little-known species of requiem shark, in the family Carcharhinidae, and the only extant member of its genus. It inhabits shallow tropical waters off northeastern South America, from Trin ...
'' Gill, 1862 ** '' Isogomphodon oxyrhynchus'' (J. P. Müller & Henle, 1839) (daggernose shark) * Genus '' Triaenodon'' J. P. Müller & Henle, 1837 ** '' Triaenodon obesus'' (Rüppell, 1837) (whitetip reef shark) † = extinct


See also

*
Shark meat Shark meat is a seafood consisting of the flesh of sharks. Several sharks are fished for human consumption, such as porbeagles, shortfin mako shark, requiem shark, and thresher shark, among others. Shark meat is popular in Asia, where it is ...


References


External links


Requiem Shark Photo Gallery
(
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to 88% of the ar ...
)
Elasmo-research


{{Taxonbar, from=Q48191 Taxa named by David Starr Jordan Taxa named by Barton Warren Evermann Extant Valanginian first appearances