''Canarium ovatum'', the pili (

Central Bikol Central Bikol commonly called Bikol Naga, also known simply as Bikol, is an Austronesian language spoken by the Bicolanos, primarily in the Bicol Region of southern Luzon, Philippines. It is spoken in the northern and western part of Camarines ...

and Filipino: ''pili'', ;), is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...

of tropical tree belonging to the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

Canarium ''Canarium'' is a genus of about 100 species of tropical and subtropical trees, in the family Burseraceae. They grow naturally across tropical Africa, south and southeast Asia, Indochina, Malesia, Australia and western Pacific Islands; includi ...

''. It is one of approximately 600 species in the family

Burseraceae The Burseraceae are a moderate-sized family of 17-19 genera and about 540 species of flowering plants. The actual numbers differ according to the time period in which a given source is written describing this family. The Burseraceae are also k ...

. ''C. ovatum'' are native to the Philippines. They are commercially cultivated in the Philippines for their edible nuts and is believed to be indigenous to that country. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...

and

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

are often vulgarized with the umbrella term of " Java almond" which mixes multiple species of the same genus, ''

''.

Description

The ''C. ovatum'' tree is a symmetrically-shaped evergreen, averaging tall with resinous wood and resistance to strong winds. It is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...

, with flowers borne on

cymose An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

inflorescence at the leaf

axil A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...

s of young shoots. As in

papaya The papaya (, ), papaw, () or pawpaw () is the plant species ''Carica papaya'', one of the 21 accepted species in the genus '' Carica'' of the family Caricaceae. It was first domesticated in Mesoamerica, within modern-day southern Mexico and ...

and

rambutan Rambutan (; taxonomic name: ''Nephelium lappaceum'') is a medium-sized tropical tree in the family Sapindaceae. The name also refers to the edible fruit produced by this tree. The rambutan is native to Southeast Asia. It is closely related to s ...

, functional

hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have ...

s exist in ''C. ovatum''.

Pollination Pollination is the transfer of pollen from an Stamen, anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by Anemophily, wind. Pollinating agents can ...

is by insects. Flowering is frequent and

s ripen through a prolonged period of time. The ovary contains three

locule A locule (plural locules) or loculus (plural loculi) (meaning "little place" in Latin) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usu ...

s, each with two ovules; most of the time only one ovule develops. The fruit of ''C. ovatum'' is a

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part ( exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or ''pyrena'') of hardened endocarp with a seed (''kernel ...

, long, in diameter, and weighs . The skin (

exocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...

) is smooth, thin, shiny, and turns purplish black when the fruit ripens; the pulp (

mesocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Agg ...

) is fibrous, fleshy, and greenish yellow in color, and the hard shell (

endocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...

) within protects a normally

dicotyledonous The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...

embryo. The basal end of the shell (endocarp) is pointed and the

apical Apical means "pertaining to an apex". It may refer to: *Apical ancestor, refers to the last common ancestor of an entire group, such as a species (biology) or a clan (anthropology) *Apical (anatomy), an anatomical term of location for features loc ...

end is more or less blunt; between the seed and the hard shell (endocarp) is a thin, brownish, fibrous seed coat developed from the inner layer of the endocarp. This thin coat usually adheres tightly to the shell and/or the seed. Much of the kernel weight is made up of the

cotyledon A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...

s, which are about 4.1–16.6% of the whole fruit; it is composed of approximately 8%

carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...

, 11.5–13.9%

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...

, and 70% fat. Kernels from some trees may be bitter, fibrous or have a turpentine odor.

Distribution and habitat

''C. ovatums distribution range is the Philippines. ''C. ovatum'' is a tropical tree preferring deep, fertile, well drained

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...

, warm temperatures, and well distributed

rainfall Rain is water droplets that have condensed from atmospheric water vapor and then fall under gravity. Rain is a major component of the water cycle and is responsible for depositing most of the fresh water on the Earth. It provides water f ...

. It cannot tolerate the slightest frost or low temperature. Refrigeration of seeds at resulted in loss of viability after 5 days. Seed germination is highly recalcitrant, reduced from 98 to 19% after 12 weeks of storage at room temperature; seeds stored for more than 137 days did not germinate. Asexual propagations using

marcotting Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...

, budding, and grafting were too inconsistent to be used in commercial production. Young shoots of ''C. ovatum'' were believed to have functional internal

phloem Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is c ...

s, which rendered

bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...

ringing ineffective as a way of building up carbohydrate levels in the wood. Success in marcottage may be cultivar-dependent. The production standard for a mature ''C. ovatum'' tree is between of in-shell nut, with the harvest season from May to October and peaking between June and August. There are high variations in kernel qualities and production between seedling trees. Most ''C. ovatum'' kernels tend to stick to the shell when fresh, but come off easily after being dried to 3–5% moisture ( for 27 to 28 h). Shelled nuts, with a moisture content of 2.5–4.6%, can be stored in the shade for one year without deterioration of quality.

Cultivation and uses

In the Philippines, production centers are located in the

Bicol region Bicol, known formally as the Bicol Region or colloquially as Bicolandia ( bcl, Rehiyon kan Bikol; Rinconada Bikol: ''Rehiyon ka Bikol''; Waray Sorsogon, Masbateño: ''Rehiyon san Bikol''; tl, Rehiyon ng Bikol), is an administrative region of ...

, provinces of

Sorsogon Sorsogon, officially the Province of Sorsogon ( Bikol: ''Probinsya kan Sorsogon''; Waray: ''Probinsya han Sorsogon''; tl, Lalawigan ng Sorsogon), is a province in the Philippines located in the Bicol Region. It is the southernmost province in ...

Albay Albay,