Calophyllum inophyllum
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''Calophyllum inophyllum'' is a large
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
plant, commonly called tamanu, oil-nut, mastwood, beach calophyllum or beautyleaf. It is native to tropical
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
and
Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as ...
. Due to its importance as a source of timber for the traditional
shipbuilding Shipbuilding is the construction of ships and other floating vessels. It normally takes place in a specialized facility known as a shipyard. Shipbuilders, also called shipwrights, follow a specialized occupation that traces its roots to befo ...
of large
outrigger ship Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull. They can range from small dugout canoes to large plank-built vessels. Outrigger ...
s, it has been spread in prehistoric times by the migrations of the
Austronesian peoples The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austro ...
to the islands of
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
and
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, along with other members of the genus ''
Calophyllum ''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands. History Members of the genus ''Calophyll ...
''. It has since been naturalized in regions in the East African coast. It is also a source of the culturally important tamanu oil.


Names

''Calophyllum inophyllum'' is also known as Alexandrian laurel balltree, beach touriga, Borneo-mahogany, Indian doomba oiltree, Indian-laurel, laurelwood, red poon, satin touriga, and tacamahac-tree. In Island Southeast Asia and
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
, they are also commonly known as bintangur, bitaog, tamanu, or kamani. In India it is known as Polanga (ପୋଲାଙ୍ଗ) in Odia, Punnag (पुन्नाग) in
Marathi Marathi may refer to: *Marathi people, an Indo-Aryan ethnolinguistic group of Maharashtra, India *Marathi language, the Indo-Aryan language spoken by the Marathi people *Palaiosouda, also known as Marathi, a small island in Greece See also * * ...
and Punnai in
Tamil Tamil may refer to: * Tamils, an ethnic group native to India and some other parts of Asia **Sri Lankan Tamils, Tamil people native to Sri Lanka also called ilankai tamils **Tamil Malaysians, Tamil people native to Malaysia * Tamil language, nativ ...
languages.


Description


Tree and leaves

''Calophyllum inophyllum'' is a low-branching and slow-growing tree, it spreads with a broad and irregular crown. It usually reaches in height.


Flowers

Flowering can occur perennially, but usually two distinct flowering periods are observed twice a year, in late spring from April to June and in late autumn from October to December. The flower is wide and occurs in
racemose A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
or paniculate
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
s consisting of four to 15 flowers. It has a sweet fragrance which attracts many insects to pollinate it.


Fruit

The fruit (the ballnut) is a round, green drupe measuring in diameter. When ripe, the fruit is wrinkled and its color varies from yellow to brownish-red.


Distribution and habitat

''Calophyllum inophyllum'' is native to Africa in: Comoros;
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi ...
;
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
;
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label= Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It ...
;
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
;
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...
;
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
(including
Pemba Island Pemba Island ( ar, الجزيرة الخضراء ''al-Jazīra al-khadrā'', literally "The Green Island"; sw, Pemba kisiwa) is a Tanzanian island forming part of the Zanzibar Archipelago, lying within the Swahili Coast in the Indian Ocean. Geo ...
of the
Zanzibar Archipelago The Zanzibar Archipelago ( ar, أرخبيل زنجبار, sw, Funguvisiwa la Zanzibar) consists of several islands lying off the coast of East Africa south of the Somali sea. The archipelago is also known as the Spice Islands. There are four ...
); south, southeast and east Asia in:
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
;
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
; China (on
Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
); southern India; Andaman and Nicobar Islands
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
; Japan (
Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yona ...
);
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
;
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
;
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
; the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
; Sri Lanka;
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
;
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
;
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
; the northwestern, southwestern and south central Pacific Region in: the
Cook Islands ) , image_map = Cook Islands on the globe (small islands magnified) (Polynesia centered).svg , capital = Avarua , coordinates = , largest_city = Avarua , official_languages = , lan ...
; Fiji; French Polynesia (
Marquesas The Marquesas Islands (; french: Îles Marquises or ' or '; Marquesan: ' (North Marquesan) and ' (South Marquesan), both meaning "the land of men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in t ...
and
Society Islands The Society Islands (french: Îles de la Société, officially ''Archipel de la Société;'' ty, Tōtaiete mā) are an archipelago located in the South Pacific Ocean. Politically, they are part of French Polynesia, an overseas country of the ...
);
Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...
; the
Marshall Islands The Marshall Islands ( mh, Ṃajeḷ), officially the Republic of the Marshall Islands ( mh, Aolepān Aorōkin Ṃajeḷ),'' () is an independent island country and microstate near the Equator in the Pacific Ocean, slightly west of the Intern ...
;
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
; the
Northern Mariana Islands The Northern Mariana Islands, officially the Commonwealth of the Northern Mariana Islands (CNMI; ch, Sankattan Siha Na Islas Mariånas; cal, Commonwealth Téél Falúw kka Efáng llól Marianas), is an unincorporated territory and commonw ...
;
Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country and microstate in the western Pacific. The nation has approximately 340 islands and connects the western chain of the ...
; and
Samoa Samoa, officially the Independent State of Samoa; sm, Sāmoa, and until 1997 known as Western Samoa, is a Polynesian island country consisting of two main islands ( Savai'i and Upolu); two smaller, inhabited islands ( Manono and Apolima); ...
; and in Australia in:
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an Australian territory in the central and central northern regions of Australia. The Northern Territory shares its borders with Western Aust ...
and
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
. Now, it is widely cultivated in all
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
regions of the world. Because of its decorative leaves, fragrant
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s, and spreading crown, it is best known as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
. This tree often grows in
coast The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in ...
al regions, as well as nearby
lowland Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland. Definitions Upland and lowland are portions of p ...
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s. However, it has also been cultivated successfully in inland areas at moderate altitudes. It tolerates varied kinds of
soil Soil, also commonly referred to as earth or dirt Dirt is an unclean matter, especially when in contact with a person's clothes, skin, or possessions. In such cases, they are said to become dirty. Common types of dirt include: * Debri ...
, coastal
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class o ...
,
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay par ...
, or even degraded soil.


Uses


Shipbuilding

Mastwood is notable for its ability to grow to massive sizes in sandy or rocky beaches of island and coastal habitats, as well as its habit of sending out arching large trunks over the water where its seeds are dispersed via the currents. Due to these characters, mastwood are of particular importance to traditional
shipbuilding Shipbuilding is the construction of ships and other floating vessels. It normally takes place in a specialized facility known as a shipyard. Shipbuilders, also called shipwrights, follow a specialized occupation that traces its roots to befo ...
of the larger Austronesian outrigger ships and were carried with them as they migrated to Oceania and Madagascar. Other species of the genus ''
Calophyllum ''Calophyllum'' is a genus of tropical flowering plants in the family Calophyllaceae. They are mainly distributed in Asia, with some species in Africa, the Americas, Australasia, and the Pacific Islands. History Members of the genus ''Calophyll ...
'' were also used similarly, like '' Calophyllum soulattri'', '' Calophyllum peekelii'', and '' Calophyllum goniocarpum''. They were comparable in importance to how
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...
s were in European shipbuilding and timber industries. Various parts of the mastwood were integral to the manufacture of
outrigger canoe Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull. They can range from small dugout canoes to large plank-built vessels. Outrigger ...
s among various
Austronesian peoples The Austronesian peoples, sometimes referred to as Austronesian-speaking peoples, are a large group of peoples in Taiwan, Maritime Southeast Asia, Micronesia, coastal New Guinea, Island Melanesia, Polynesia, and Madagascar that speak Austro ...
. The large curving limbs were commonly carved into the
dugout canoe A dugout canoe or simply dugout is a boat made from a hollowed tree. Other names for this type of boat are logboat and monoxylon. ''Monoxylon'' (''μονόξυλον'') (pl: ''monoxyla'') is Greek – ''mono-'' (single) + '' ξύλον xylon'' ( ...
that formed the
keel The keel is the bottom-most longitudinal structural element on a vessel. On some sailboats, it may have a hydrodynamic and counterbalancing purpose, as well. As the laying down of the keel is the initial step in the construction of a ship, in Br ...
of the Austronesian outriggers ships. The
strake On a vessel's hull, a strake is a longitudinal course of planking or plating which runs from the boat's stempost (at the bows) to the sternpost or transom (at the rear). The garboard strakes are the two immediately adjacent to the keel on ea ...
s, which are attached to the keel by the uniquely Austronesian technique of "sewing" them with a combination of dowels and lashed lugs instead of nails, can also be made from mastwood, but it is more commonly made from other "softer" timber species like ''
Artocarpus ''Artocarpus'' is a genus of approximately 60 trees and shrubs of Southeast Asian and Pacific origin, belonging to the mulberry family, Moraceae. Most species of ''Artocarpus'' are restricted to Southeast Asia; a few cultivated species are more w ...
''. Other pieces became masts,
outrigger An outrigger is a projecting structure on a boat, with specific meaning depending on types of vessel. Outriggers may also refer to legs on a wheeled vehicle that are folded out when it needs stabilization, for example on a crane that lifts ...
floats, and outrigger spars. Smaller curving limbs can also be carved into the ribs of the boat. In many parts of Polynesia, mastwood groves planted in ''
marae A ' (in New Zealand Māori, Cook Islands Māori, Tahitian), ' (in Tongan), ' (in Marquesan) or ' (in Samoan) is a communal or sacred place that serves religious and social purposes in Polynesian societies. In all these languages, the term a ...
'' were considered sacred and abodes of spirits. Mastwood were also carved into religious objects like ''
tiki In Māori mythology, Tiki is the first man created by either Tūmatauenga or Tāne. He found the first woman, Marikoriko, in a pond; she seduced him and he became the father of Hine-kau-ataata. By extension, a tiki is a large or small wooden, ...
''. They are also commonly mentioned in the chants and folklore of Polynesia. In Australia, the 1889 book ''The Useful Native Plants of Australia'' records "During a debate on the Pearl Fisheries Bill in the Queensland Assembly, a clause was specially inserted to protect trees of this species at
Thursday Island Thursday Island, colloquially known as TI, or in the Kawrareg dialect, Waiben or Waibene, is an island of the Torres Strait Islands, an archipelago of at least 274 small islands in the Torres Strait. TI is located approximately north of Cape ...
. A fine of £10 is inflicted on any person who cuts down or injures this or a cocoa-nut tree, or any other tree bearing edible fruit. This clause is, of course, in the interest of the aboriginals.".


Other uses

Aside from shipbuilding, tamanu oil extracted from the fruit kernels were important in Polynesian culture. The oils, as well as poultices made from leaves and flowers, are also commonly used for traditional medicine. The leaves contain compounds that are poisonous to fish and can be used as
fish poison Fish toxins or fish stupefying plants have historically been used by many hunter gatherer cultures to stun fish, so they become easy to collect by hand. Some of these toxins paralyse fish, which can then be easily collected. The process of documen ...
. The sap of the tree is poisonous and is used to make poison arrows in Samoa. The mature fruit is poisonous enough to use as rat bait. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s yield a thick, dark green tamanu oil for medicinal use or hair grease. The nuts are dried before cracking, after which the oil-laden kernel is removed and further dried. The first neoflavone isolated from natural sources (1951) was calophyllolide from ''C. inophyllum'' seeds. The Mavilan, a Tulu-speaking tribe in north
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South ...
in India, use the bark to make a powder that they mix with water and apply it to plants affected by a type of plant disease caused by water that they call ''neeru vembu''.Suresh, K. P. (2010). Indigenous Agricultural Practices among Mavilan Tribe in North Kerala.
/ref> The
fatty acid methyl ester Fatty acid methyl esters (FAME) are a type of fatty acid ester that are derived by transesterification of fats with methanol. The molecules in biodiesel are primarily FAME, usually obtained from vegetable oils by transesterification. They are used ...
s derived from ''C. inophyllum'' seed oil meets the major
biodiesel Biodiesel is a form of diesel fuel derived from plants or animals and consisting of long-chain fatty acid esters. It is typically made by chemically reacting lipids such as animal fat ( tallow), soybean oil, or some other vegetable oil ...
requirements in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
( ASTM D 6751), and
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been de ...
( EN 14214). The average oil yield is 11.7 kg-oil/tree or 4680 kg-oil/hectare. In the northwest coastal areas of Luzon island in Philippines, the oil was used for night lamps. This widespread use started to decline when kerosene became available, and later on electricity. It was also used as fuel to generate electricity to provide power for radios during World War II.


See also

*
Domesticated plants and animals of Austronesia One of the major human migration events was the maritime settlement of the islands of the Indo-Pacific by the Austronesian peoples, believed to have started from at least 5,500 to 4,000 BP (3500 to 2000 BCE). These migrations were accompanied ...


References


External links

* {{Taxonbar, from=Q311471 inophyllum Flora of tropical Asia Flora of the Pacific Trees of Australia Trees of China Flora of the Western Indian Ocean Trees of Japan Flora of Kenya Flora of Mozambique Flora of the Northern Territory Flora of Queensland Flora of the Ryukyu Islands Trees of Taiwan Flora of Tanzania Flora of the Zanzibar Archipelago Malpighiales of Australia Plants described in 1753 Taxa named by Carl Linnaeus Least concern biota of Asia Least concern flora of Oceania Least concern flora of Australia Least concern biota of Queensland Ornamental trees