The brown roofed turtle (''Pangshura smithii'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of turtle in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Geoemydidae The Geoemydidae (formerly known as Bataguridae) are one of the largest and most diverse families in the order Testudines (turtles), with about 70 species. The family includes the Eurasian pond and river turtles and Neotropical wood turtles. Mem ...

. The species is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

South Asia South Asia is the southern subregion of Asia, which is defined in both geographical Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth descr ...

. Two subspecies are recognized.

Etymology

The specific name, ''smithii'', is in honor of Scottish

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...

Andrew Smith.

Description

The carapace of ''P. smithii'' is much depressed and feebly keeled. The nuchal shield is small, trapezoidal, and broadest posteriorly. The first vertebral has sinuous lateral borders and is usually a little narrower in front than behind. The second vertebral is shortest, broader than long, and usually with straight or slightly convex posterior border. The third vertebral is considerably longer than broad, subquadrangular, and its posterior border is straight or slightly convex. The fourth vertebral is longest, tapering anteriorly and forming a narrow suture with the third. The fifth vertebral is much broader than the others. The large plastron is feebly angulated laterally. The front lobe is rounded. The hind lobe is angularly notched and as long as or a little shorter than the width of the bridge. The longest median suture is that between the abdominals, which about equals the length of the front lobe. The gulars are usually shorter than the suture between the humerals, their suture with the latter shields forming a right angle. The inguinal is large, and the axillary is smaller. The head is moderate. The snout is short, obtuse, and feebly prominent. The jaws have denticulated edges. The upper jaw is not notched mesially. The alveolar surface of the upper jaw is broad, and the median ridge is nearer the inner than the outer border. There are bony choanae between the orbits. The width of the lower jaw at the symphysis is less than the diameter of the orbit. The fore limbs have large transverse scales. The carapace is pale olive-brown above, and the dorsal keel is usually blackish. The plastral shields and the lower surface of the marginals are dark brown, bordered with yellow. The straight-line carapace length is . Brown roofed turtles exhibit ZZ/ZW sex determination, in contrast to the

temperature-dependent sex determination Temperature-dependent sex determination (TSD) is a type of environmental sex determination in which the temperatures experienced during embryonic/larval development determine the sex of the offspring. It is only observed in reptiles and teleost fish ...

of most turtles.

Subspecies and geographic ranges

Two subspecies are recognized, including the nominotypical subspecies. *''P. s. smithii'' , is found in the Indus,

Ganges The Ganges ( ) (in India: Ganga ( ); in Bangladesh: Padma ( )). "The Ganges Basin, known in India as the Ganga and in Bangladesh as the Padma, is an international river to which India, Bangladesh, Nepal and China are the riparian states." is ...

, and

Brahmaputra The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. I ...

River drainages in

Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

, and

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...

. *''P. s. pallidipes'' , is found in the northern tributaries of the Ganges in India and

Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...

Habitat

The preferred natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s of ''P. smithii'' are freshwater swamps and rivers. These creatures also prefer to bask in the sun, twice a day. They often prefer muddy areas. www.reptile-database.org.

References

External links