Bos mutus
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The wild yak (''Bos mutus'') is a large, wild
bovine Bovines (subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship betwe ...
native to the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
. It is the ancestor of the
domestic yak The domestic yak (''Bos grunniens''), also known as the Tartary ox, grunting ox or hairy cattle, is a species of long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Kachin Sta ...
(''Bos grunniens'').


Taxonomy

The ancestor of the wild and domestic yak is thought to have diverged from ''
Bos primigenius The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocene; ...
'' at a point between one and five million years ago. The wild yak is now normally treated as a separate species from the domestic yak (''Bos grunniens'').


Description

The wild yak is among the largest extant bovid species. Adults stand about tall at the shoulder, and weigh . The head and body length is , not counting the tail of .Han Jianlin, M. Melletti, J. Burton, 2014, Wild yak (Bos mutus Przewalski, 1883), Ecology, Evolution and Behavior of Wild Cattle: Implications for Conservation, Chapter 1, p.203,
Cambridge University Press Cambridge University Press is the university press of the University of Cambridge. Granted letters patent by King Henry VIII in 1534, it is the oldest university press in the world. It is also the King's Printer. Cambridge University Pre ...
The females are about one-third the weight and are about 30% smaller in their linear dimensions when compared to bull wild yaks. Domesticated yaks are somewhat smaller. They are heavily built animals with a bulky frame, sturdy legs, and rounded cloven hooves. To protect against the cold, the
udder An udder is an organ formed of two or four mammary glands on the females of dairy animals and ruminants such as cattle, goats, and sheep. An udder is equivalent to the breast in primates and elephantine pachyderms. The udder is a single mass han ...
in females and the
scrotum The scrotum or scrotal sac is an anatomical male reproductive structure located at the base of the penis that consists of a suspended dual-chambered sac of skin and smooth muscle. It is present in most terrestrial male mammals. The scrotum co ...
in males are small, and covered in a layer of hair. Females have four
teat A teat is the projection from the mammary glands of mammals from which milk flows or is ejected for the purpose of feeding young. In many mammals the teat projects from the udder. The number of teats varies by mammalian species and often corr ...
s. Both sexes have long shaggy hair, with a dense woolly undercoat over the chest, flanks, and thighs for insulation against the cold. In males especially, this undercoat may form a long "skirt" that can reach the ground. The tail is long and horse-like, rather than tufted like the tails of cattle or bison. The coat is typically black or dark brown, covering most of the body, with a grey muzzle (although some wild golden-brown individuals have been reported). Wild yaks with gold coloured hair are known as the wild golden yak (). They are considered an endangered subspecies in China, with an estimated population of 170 left in the wild. Two morphological types have been identified, so-called ''Qilian'' and ''Kunlun''.


Distribution and habitat

Wild yaks once ranged up to southern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
to the east of Lake Baikal, but became extinct in Russia around the 17th century. Today, wild yaks are found primarily in northern
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa, Taman ...
and western
Qinghai Qinghai (; alternately romanized as Tsinghai, Ch'inghai), also known as Kokonor, is a landlocked province in the northwest of the People's Republic of China. It is the fourth largest province of China by area and has the third smallest po ...
, with some populations extending into the southernmost parts of
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
, and into
Ladakh Ladakh () is a region administered by India as a union territory which constitutes a part of the larger Kashmir region and has been the subject of dispute between India, Pakistan, and China since 1947. (subscription required) Quote: "Jammu ...
in India. Small, isolated populations of wild yak are also found farther afield, primarily in western Tibet and eastern Qinghai. In historic times, wild yaks were also found in
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainou ...
, but they are now considered extinct there. The primary habitat of wild yaks consists of treeless uplands between , dominated by mountains and
plateau In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ...
s. They are most commonly found in alpine tundra with a relatively thick carpet of grasses and sedges rather than the more barren steppe country. The wild yak was thought to be regionally extinct in
Nepal Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...
in the 1970s, but was rediscovered in
Humla Humla District ( ne, हुम्ला जिल्ला), a part of Karnali Province, is one of the seventy-seven districts of Nepal. The district, with Simikot as its district headquarters, covers an area of and has population of 50,858 a ...
in 2014. This discovery later made the species to be painted on Nepal's currency.


Behaviour and ecology

The diet of wild yaks consists largely of grasses and sedges, such as ''
Carex ''Carex'' is a vast genus of more than 2,000 species of grass-like plants in the family Cyperaceae, commonly known as sedges (or seg, in older books). Other members of the family Cyperaceae are also called sedges, however those of genus ''Carex'' ...
'', ''
Stipa ''Stipa'' is a genus of around 300 large perennial hermaphroditic grasses collectively known as feather grass, needle grass, and spear grass. They are placed in the subfamily Pooideae and the tribe Stipeae, which also contains many species for ...
'', and ''
Kobresia ''Kobresia'' is a genus of plants in the sedge family. They are sometimes called bog sedges. These perennial sedges are quite similar to ''Carex'' species in appearance. The genus is widespread across much of Europe, Asia and North America, with ...
''. They also eat a smaller amount of herbs,
winterfat ''Krascheninnikovia'' is a genus of flowering plants in the subfamily Chenopodioideae of the family Amaranthaceae known as winterfat, so-called because it is a nutritious livestock forage. They are known from Eurasia and western North America. Th ...
shrubs, and mosses, and have even been reported to eat lichen. Historically, the main natural predator of the wild yak has been the
Himalayan wolf The Himalayan wolf (''Canis lupus chanco'') is a canine of debated taxonomy. It is distinguished by its genetic markers, with mitochondrial DNA indicating that it is genetically basal to the Holarctic gray wolf, genetically the same wolf as t ...
, but
Himalayan black bear The Himalayan black bear (''Ursus thibetanus laniger'') is a subspecies of the Asian black bear found in the Himalayas of India, Bhutan, Nepal, China, and Pakistan. Description It is distinguished from '' U. t. thibetanus'' by its longer, thicke ...
s, Himalayan brown bears and
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a felid in the genus '' Panthera'' native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because the global population is es ...
s have also been reported as predators in some areas, likely of young or infirm wild yaks.
Thubten Jigme Norbu Thubten Jigme Norbu () (August 16, 1922 – September 5, 2008), recognised as the Taktser Rinpoche, was a Tibetan lama, writer, civil rights activist and professor of Tibetan studies and was the eldest brother of the 14th Dalai Lama, Tenzin G ...
, the elder brother of the
14th Dalai Lama The 14th Dalai Lama (spiritual name Jetsun Jamphel Ngawang Lobsang Yeshe Tenzin Gyatso, known as Tenzin Gyatso (Tibetan: བསྟན་འཛིན་རྒྱ་མཚོ་, Wylie: ''bsTan-'dzin rgya-mtsho''); né Lhamo Thondup), known as ...
, reported on his journey from
Kumbum A Kumbum ( "one hundred thousand holy images") is a multi-storied aggregate of Buddhist chapels in Tibetan Buddhism. The most famous Kumbum forms part of Palcho Monastery. The first Kumbum was founded in the fire sheep year 1427 by a Gyantse p ...
in
Amdo Amdo ( am˥˥.to˥˥ ) is one of the three traditional Tibetan regions, the others being U-Tsang in the west and Kham in the east. Ngari (including former Guge kingdom) in the north-west was incorporated into Ü-Tsang. Amdo is also the ...
to
Lhasa Lhasa (; Lhasa dialect: ; bo, text=ལྷ་ས, translation=Place of Gods) is the urban center of the prefecture-level Lhasa City and the administrative capital of Tibet Autonomous Region in Southwest China. The inner urban area of Lhas ...
in 1950: Wild yaks are herd animals. Herds can contain several hundred individuals, although many are much smaller. Herds consist primarily of females and their young, with a smaller number of adult males. On average female yaks graze 100m higher than males. Females with young tend to choose grazing ground on high, steep slopes. The remaining males are either solitary, or found in much smaller groups, averaging around six individuals. Groups move into lower altitude ranges during the winter. Although wild yaks can become aggressive when defending young, or during the rut, they generally avoid humans, and may flee for great distances if approached.


Reproduction

Wild yaks mate in summer and give birth to a single calf the following spring.Wiener, G.; Jianlin, H.; Ruijun, L. (2003)
"4 The Yak in Relation to Its Environment"
''The Yak, Second Edition''. Bangkok: Regional Office for Asia and the Pacific Food and Agriculture Organization of the United Nations, . Accessed 8 August 2008.
Females typically only give birth every other year.


Conservation

The wild yak is currently listed as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. It was previously classified as
Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and in ...
, but was downlisted in 1996 based on the estimated rate of population decline and current population sizes. The latest assessment in 2008 suggested a total population of no more than 10,000 mature individuals. The wild yak is experiencing threats applied by several sources. Poaching, including commercial poaching, has remained the most serious threat; males are particularly impacted because of their more solitary habits. Disturbance by and interbreeding with livestock herds is also common. This may include the transmission of cattle-borne diseases, although no direct evidence of this has yet been found. Conflicts with herders themselves, as in preventive and retaliatory killings for abduction of domestic yaks by wild herds, also occur but appear to be relatively rare. Recent protection from poaching particularly appears to have stabilized or even increased population sizes in several areas, leading to the IUCN downlisting in 2008. In both China and India, the species is officially protected; in China it is present in a number of large nature reserves.


Impact on humans

The wild yak is a reservoir for
zoonotic diseases A zoonosis (; plural zoonoses) or zoonotic disease is an infectious disease of humans caused by a pathogen (an infectious agent, such as a bacterium, virus, parasite or prion) that has jumped from a non-human (usually a vertebrate) to a human ...
of both bacterial and viral origins. Such bacterial diseases include anthrax,
botulism Botulism is a rare and potentially fatal illness caused by a toxin produced by the bacterium ''Clostridium botulinum''. The disease begins with weakness, blurred vision, feeling tired, and trouble speaking. This may then be followed by weakne ...
,
tetanus Tetanus, also known as lockjaw, is a bacterial infection caused by ''Clostridium tetani'', and is characterized by muscle spasms. In the most common type, the spasms begin in the jaw and then progress to the rest of the body. Each spasm usually ...
, and
tuberculosis Tuberculosis (TB) is an infectious disease usually caused by '' Mycobacterium tuberculosis'' (MTB) bacteria. Tuberculosis generally affects the lungs, but it can also affect other parts of the body. Most infections show no symptoms, i ...
.


References


External links

*ARKive
images and movies of the wild yak ''(Bos grunniens)''
{{Taxonbar, from=Q26547 Mammals described in 1883 Mammals of Nepal Mammals of India Mammals of China Bovines Extant Zanclean first appearances Yaks Taxa named by Nikolay Przhevalsky