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A bone bed is any
geological Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Eart ...
stratum In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ei ...
or deposit that contains
bone A bone is a Stiffness, rigid Organ (biology), organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red blood cell, red and white blood cells, store minerals, provid ...
s of whatever kind. Inevitably, such deposits are
sedimentary Sedimentary rocks are types of rock (geology), rock that are formed by the accumulation or deposition of mineral or organic matter, organic particles at Earth#Surface, Earth's surface, followed by cementation (geology), cementation. Sedimentati ...
in nature. Not a formal term, it tends to be used more to describe especially dense collections such as
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...
. It is also applied to
breccia Breccia () is a rock composed of large angular broken fragments of minerals or rocks cemented together by a fine-grained matrix. The word has its origins in the Italian language, in which it means "rubble". A breccia may have a variety of di ...
ted and stalagmitic deposits on the floor of caves, which frequently contain osseous remains. In a more restricted sense, the term is used to describe certain thin layers of bony fragments, which occur in well-defined geological strata. One of the best-known of these is the
Ludlow Bone Bed Ludlow () is a market town in Shropshire, England. The town is significant in the history of the Welsh Marches and in relation to Wales. It is located south of Shrewsbury and north of Hereford, on the A49 road (Great Britain), A49 road which ...
, which is found at the base of the Downton Sandstone in the Upper Ludlow series. At
Ludlow Ludlow () is a market town in Shropshire, England. The town is significant in the history of the Welsh Marches and in relation to Wales. It is located south of Shrewsbury and north of Hereford, on the A49 road which bypasses the town. The t ...
(
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
) itself, two such beds are actually known, separated by about . of strata. Although quite thin, the Ludlow Bone Bed can be followed from that town into
Gloucestershire Gloucestershire ( abbreviated Glos) is a county in South West England. The county comprises part of the Cotswold Hills, part of the flat fertile valley of the River Severn and the entire Forest of Dean. The county town is the city of Gl ...
, for a distance of . It is almost completely made up of fragments of spines,
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
and
scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number w ...
of
ganoid fish In most biological nomenclature, a scale ( grc, λεπίς, lepís; la, squāma) is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran (butterfly and moth) species, scales are plates on the surface o ...
. Another well-known bed, formerly known as the
Bristol Bristol () is a city, ceremonial county and unitary authority in England. Situated on the River Avon, it is bordered by the ceremonial counties of Gloucestershire to the north and Somerset to the south. Bristol is the most populous city in ...
or Lias Bone Bed, exists in the form of several thin layers of
mica Micas ( ) are a group of silicate minerals whose outstanding physical characteristic is that individual mica crystals can easily be split into extremely thin elastic plates. This characteristic is described as perfect basal cleavage. Mica is ...
ceous
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
, with the remains of fish and
saurian Sauria is the clade containing the most recent common ancestor of archosaurs (such as crocodilians, dinosaurs, etc.) and lepidosaurs ( lizards and kin), and all its descendants. Since most molecular phylogenies recover turtles as more closel ...
s, which occur in the
Rhaetic Black Paper Shale Rhaetic or Raetic (), also known as Rhaetian, was a language spoken in the ancient region of Rhaetia in the eastern Alps in pre-Roman and Roman times. It is documented by around 280 texts dated from the 5th up until the 1st century BC, which were ...
s that lie above the
Keuper The Keuper is a lithostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Keuper consists of dolomite, shales or claystones and evaporites that were deposited during the Middle and Late ...
marl Marl is an earthy material rich in carbonate minerals, clays, and silt. When hardened into rock, this becomes marlstone. It is formed in marine or freshwater environments, often through the activities of algae. Marl makes up the lower part o ...
s, in the south-west of England. A similar bone bed has been traced on the same geological horizon in Brunswick,
Hanover Hanover (; german: Hannover ; nds, Hannober) is the capital and largest city of the German state of Lower Saxony. Its 535,932 (2021) inhabitants make it the 13th-largest city in Germany as well as the fourth-largest city in Northern Germany ...
(
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
), in
Franconia Franconia (german: Franken, ; Franconian dialect: ''Franggn'' ; bar, Frankn) is a region of Germany, characterised by its culture and Franconian dialect (German: ''Fränkisch''). The three administrative regions of Lower, Middle and Upper Fr ...
and in
Tübingen Tübingen (, , Swabian: ''Dibenga'') is a traditional university city in central Baden-Württemberg, Germany. It is situated south of the state capital, Stuttgart, and developed on both sides of the Neckar and Ammer rivers. about one in thr ...
(Germany). A bone bed has also been observed at the base of the
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
series, in certain parts of the south-west of England. Bone beds are also recorded in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
,
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
,
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Terrestrial bonebed examples are: the Triassic ''
Metoposaurus ''Metoposaurus'' meaning "front lizard" is an extinct genus of stereospondyl temnospondyl amphibian, known from the Late Triassic of Germany, Italy, Poland, and Portugal. https://www.app.pan.pl/archive/published/app47/app47-535.pdf This mostly a ...
'' bone bed from
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
, the ''
Mapusaurus ''Mapusaurus'' () was a giant carcharodontosaurid carnosaurian dinosaur from the early Late Cretaceous (early Turonian stage), approximately 93.9 to 89.6 million years ago, of what is now Argentina. Discovery ''Mapusaurus'' was excavated be ...
'' bone bed at Cañadón del Gato, in
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, the ''
Allosaurus ''Allosaurus'' () is a genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch (Kimmeridgian to late Tithonian). The name "''Allosaurus''" means "different lizard" alluding to ...
''-dominated
Cleveland-Lloyd Dinosaur Quarry Jurassic National Monument, at the site of the Cleveland-Lloyd Dinosaur Quarry, well known for containing the densest concentration of Jurassic dinosaur fossils ever found, is a paleontological site located near Cleveland, Utah, in the San Rafa ...
of
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
, the
Dinosaur National Monument Dinosaur National Monument is an American national monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green and Yampa rivers. Although most of the monument area is in ...
on the boundary of Utah and
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
, an ''
Albertosaurus ''Albertosaurus'' (; meaning "Alberta lizard") is a genus of tyrannosaurid theropod dinosaurs that lived in western North America during the Late Cretaceous Period, about 71 million years ago. The type species, ''A. sarcophagus'', was app ...
'' bonebed from
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
, a ''
Daspletosaurus ''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species ...
'' bone bed from
Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
, the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
John Day Fossil Beds of
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
, a ''
Triceratops ''Triceratops'' ( ; ) is a genus of herbivore, herbivorous Chasmosaurinae, chasmosaurine Ceratopsidae, ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous Period (geology), period, about 68 m ...
'' bonebed from Montana, a ''
Centrosaurus ''Centrosaurus'' ( ; ) is a genus of herbivorous ceratopsian dinosaur from the Late Cretaceous of Canada. Their remains have been found in the Dinosaur Park Formation, dating from 76.5 to 75.5 million years ago. Discovery and naming The firs ...
'' bonebed in Alberta, a ''
Styracosaurus ''Styracosaurus'' ( ; meaning "spiked lizard" from the Ancient Greek / "spike at the butt-end of a spear-shaft" and / "lizard") is a genus of herbivorous ceratopsian dinosaur from the Cretaceous Period (Campanian stage), about 75.5 to 74.5&nbs ...
'' bone bed in Alberta, an ''
Edmontosaurus annectens ''Edmontosaurus annectens'' (meaning "connected lizard from Edmonton") is a species of flat-headed and duck-billed (hadrosaurid) dinosaur from the very end of the Cretaceous Period (geology), Period, in what is now North America. Remains of ''E. ...
'' bone bed in
Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
, an ''
Edmontosaurus regalis ''Edmontosaurus regalis'' is a species of comb-crested hadrosaurid (duck-billed) dinosaur. Fossils of ''E. regalis'' have been found in rocks of western North America that date from the late Campanian stage of the Cretaceous Period 73 million yea ...
'' bone bed in Alberta, a ''
Gryposaurus ''Gryposaurus'' (meaning "hooked-nosed (Ancient Greek, Greek ''grypos'') lizard"; sometimes incorrectly translated as "griffin (Latin ''gryphus'') lizard") was a genus of hadrosaur, duckbilled dinosaur that lived about 80 to 75 million years ...
'' bone bed in the
Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...
, a ''
Pachyrhinosaurus ''Pachyrhinosaurus'' (meaning in Greek "thick-nosed lizard", from ' (), thick; ' (), nose; and (), lizard) is an extinct genus of centrosaurine ceratopsid dinosaur from the Late Cretaceous period of North America. The first examples were discove ...
'' bone bed in the Wapiti Formation, and the
Nemegt Basin The Nemegt Basin is a geographical area in the northwestern Gobi Desert, in Ömnögovi Province, southern Mongolia. It is known locally as the "Valley of the Dragons", since it is a source of many fossil finds, including dinosaurs, dinosaur eggs, ...
in the
Gobi Desert The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...
region of Mongolia, specifically the ''
Saurolophus ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 million ...
'' bone bed known as the Dragon's Tomb.
Bentiaba The Bentiaba (or ''Rio de São Nicolau'' or Saint Nicolas River) is a river in southern Angola. Its mouth is at the Atlantic Ocean near the commune of Bentiaba in Namibe Province. The riverbank has produced a number of Cretaceous fossils includin ...
,
Angola , national_anthem = " Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
, is an example of a marine bonebed with numerous mosasaurs and plesiosaurs. Another example of a marine bonebed is the Sharktooth Hill Bonebed located in the
Temblor Formation The Temblor Formation is a geologic formation in California. It preserves fossils dating back from the Late Oligocene to the Middle Miocene of the Neogene period. It is notable for the famous Sharktooth Hill deposit (otherwise known as Ernst Q ...
in California. Fossil bonebeds don't always consist of one single species, but rather many species of organisms. There are several of the bonebeds known throughout North America. Two of the best examples include the Mixson's Bone Bed of Florida, whose geological settings preserved the remains of ''Ambelodon'', ''
Aepycamelus ''Aepycamelus'' is an extinct genus of camelids that lived during the Miocene 20.6–4.9 million years ago, existing for about . Its name is derived from the Homeric Greek , "high and steep" and κάμηλος – "camel"; thus, "high camel"; ''a ...
'', and ''
Cormohipparion ''Cormohipparion'' is an extinct genus of horse belonging to the tribe Hipparionini that lived in North America during the late Miocene to Pliocene ( Hemphillian to Blancan in the NALMA classification). This ancient species of horse grew up to l ...
'', and the Agate Fossil Beds in
Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the southwe ...
which has the fossils of abundant creatures such as ''
Menoceras ''Menoceras'' ("Crescent Horns") is a genus of extinct, small rhinoceros endemic to most of southern North America and ranged as far south as Panama during the early Miocene epoch. It lived from around 30.7—19.7 Ma, existing for approximately ...
'', ''
Stenomylus ''Stenomylus'' is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek (, "narrow") and (, "molar"). ''Stenomylus'' was extremely diminutive compared to ...
'', and ''
Daphoenodon ''Daphoenodon'' is an extinct genus of terrestrial carnivore, which lived in the early Miocene and belonged to the family Amphicyonidae ("bear dogs") of the suborder Caniformia Caniformia is a suborder within the order Carnivora consistin ...
''.


References

{{Reflist Fossils