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{{refimprove, date=July 2020 A bolete is a type of
mushroom A mushroom or toadstool is the fleshy, spore-bearing fruiting body of a fungus, typically produced above ground, on soil, or on its food source. ''Toadstool'' generally denotes one poisonous to humans. The standard for the name "mushroom" is ...
, or
fungal A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...
fruiting body The sporocarp (also known as fruiting body, fruit body or fruitbody) of fungi is a multicellular structure on which spore-producing structures, such as basidia or asci, are borne. The fruitbody is part of the sexual phase of a fungal life cy ...
. It can be identified thanks to a unique mushroom
cap A cap is a flat headgear, usually with a visor. Caps have crowns that fit very close to the head. They made their first appearance as early as 3200 BC. Caps typically have a visor, or no brim at all. They are popular in casual and informal se ...
. The cap is clearly different from the stem. On the underside of the cap there is usually a spongy surface with pores, instead of the
gills A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
typical of mushrooms. However, there are some boletes that are gilled, such as species of ''
Chroogomphus ''Chroogomphus'' is a genus of mushrooms commonly known as pine-spikes or spike-caps based on their shape and because they are often found growing in association with pine trees. The genus is distributed throughout the Northern Hemisphere includ ...
'', '' Gomphidius'', ''
Paxillus ''Paxillus'' is a genus of mushrooms of which most are known to be poisonous or inedible. Species include '' Paxillus involutus'' and '' Paxillus vernalis''. Two former species—''Tapinella panuoides'' and ''Tapinella atrotomentosa''—have now ...
'', ''
Phylloporus ''Phylloporus'' is a genus of fungi in the family Boletaceae (suborder Boletineae). The genus has a cosmopolitan distribution, and contains about 50 species, mostly in tropical The tropics are the regions of Earth surrounding the Equator. T ...
'' and ''
Hygrophoropsis aurantiaca ''Hygrophoropsis aurantiaca'', commonly known as the false chanterelle, is a species of fungus in the family Hygrophoropsidaceae. It is found across several continents, growing in woodland and heathland, and sometimes on woodchips used in gar ...
''. "Bolete" is the English common name for fungus species whose mushroom caps have this appearance. The boletes are classified in the order
Boletales The Boletales are an order of Agaricomycetes containing over 1300 species with a diverse array of fruiting body types. The boletes are the best known members of this group, and until recently, the Boletales were thought to only contain boletes. ...
. Not all members of the order Boletales are boletes. The micromorphology and
molecular phylogeny Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
of the order Boletales have established that it also contains many gilled,
puffball Puffballs are a type of fungus featuring a ball-shaped fruit body that bursts on impact, releasing a cloud of dust-like spores when mature. Puffballs belong to the division Basidiomycota and encompass several genera, including '' Calvatia'', '' ...
, and other fruit body shapes. A similar pore surface is found in
polypore Polypores are a group of fungi that form large fruiting bodies with pores or tubes on the underside (see Delimitation for exceptions). They are a morphological group of basidiomycetes-like gilled mushrooms and hydnoid fungi, and not all polyp ...
s, but these species generally have a different physical structure from boletes, and have different microscopic characteristics than boletes. Many polypores have much firmer, often woody, flesh. Boletes are susceptible to infection by the fungus ''Hypomyces chrysospermus'', also known as the ''
bolete eater ''Hypomyces chrysospermus'', the bolete eater, is a parasitic ascomycete fungus that grows on bolete mushrooms, turning the afflicted host a whitish, golden yellow, or tan color. It is found in Eurasia and North America, as well as southwest We ...
''.


External links


"Evolution & Morphology in the Homobasidiomycetes" by Gary Lincoff & Michael Wood, ''MykoWeb.com''
Mushroom types Fungal morphology and anatomy
Boletes {{refimprove, date=July 2020 A bolete is a type of mushroom, or fungal fruiting body. It can be identified thanks to a unique mushroom cap. The cap is clearly different from the stem. On the underside of the cap there is usually a spongy surfa ...
Basidiomycota cs:Hřib