Bakonydraco
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''Bakonydraco'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
period (
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. ...
stage) of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Cr ...
.


Etymology

''Bakonydraco'' was named in 2005 by paleontologists Attila Ősi,
David Weishampel Professor David Bruce Weishampel (born November 16, 1952) is an American palaeontologist in the Center for Functional Anatomy and Evolution at Johns Hopkins University School of Medicine. Weishampel received his Ph.D. in Geology from the Universi ...
, and Jianu Coralia. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
is ''Bakonydraco galaczi''. The genus name refers to the Bakony Mountains and to
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''draco'', "
dragon A dragon is a reptilian legendary creature that appears in the folklore of many cultures worldwide. Beliefs about dragons vary considerably through regions, but dragons in western cultures since the High Middle Ages have often been depicted a ...
". The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''galaczi'' honors Professor András Galácz, who helped the authors in the Iharkút Research Program, where fossils are since 2000 found in
open-pit mining Open-pit mining, also known as open-cast or open-cut mining and in larger contexts mega-mining, is a surface mining technique of extracting rock or minerals from the earth from an open-air pit, sometimes known as a borrow. This form of mini ...
of
bauxite Bauxite is a sedimentary rock with a relatively high aluminium content. It is the world's main source of aluminium and gallium. Bauxite consists mostly of the aluminium minerals gibbsite (Al(OH)3), boehmite (γ-AlO(OH)) and diaspore (α-AlO ...
, among them the remains of pterosaurs, the first ever discovered in Hungary.


Description

''Bakonydraco'' is based on
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
MTM Gyn/3, a nearly complete
mandibula In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
, a fusion of the lower
jaws Jaws or Jaw may refer to: Anatomy * Jaw, an opposable articulated structure at the entrance of the mouth ** Mandible, the lower jaw Arts, entertainment, and media * Jaws (James Bond), a character in ''The Spy Who Loved Me'' and ''Moonraker'' * ...
. Also assigned to it, as
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...
, is MTM Gyn/4, 21: parts from another jaw's
symphysis A symphysis (, pl. symphyses) is a fibrocartilaginous fusion between two bones. It is a type of cartilaginous joint, specifically a secondary cartilaginous joint. # A symphysis is an amphiarthrosis, a slightly movable joint. # A growing togethe ...
(the front parts, having fused into a single blade-like structure, of the two lower jaws); azhdarchid
wing A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is e ...
bones and
neck The neck is the part of the body on many vertebrates that connects the head with the torso. The neck supports the weight of the head and protects the nerves that carry sensory and motor information from the brain down to the rest of the body. In ...
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e from the same area may also belong to it. The lower jaws are toothless and the two halves of the mandibula are frontally fused for about half of its overall length, forming a long, pointed section that is compressed side-to-side and also expanded vertically, giving it a somewhat
spear A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head. The head may be simply the sharpened end of the shaft itself, as is the case with fire hardened spears, or it may be made of a more durable material fastene ...
head- or
arrow An arrow is a fin-stabilized projectile launched by a bow. A typical arrow usually consists of a long, stiff, straight shaft with a weighty (and usually sharp and pointed) arrowhead attached to the front end, multiple fin-like stabilizers ...
head-like shape from the side. This expansion occurs both on the lower edge and on the top surface, where the most extreme point corresponds with a transverse ridge which separates the straight back half of the symphysis from the pointed end in the front. The jaws of MTM Gyn/3 are 29 centimeters (11.4 inches) long, and the
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
of the genus is estimated to be 3.5 to 4 meters (11.5 to 13.1 feet), which is medium-sized for a pterosaur. Because the jaws are relatively taller than other azhdarchids, and reminiscent of '' Tapejara'', it could have been a piscivore (feeding on small
fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
), or a
frugivore A frugivore is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance and ...
.


Classification

Initially, ''Bakonydraco'' was assigned to the family
Azhdarchidae Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early C ...
, however, paleontologists Brian Andres and Timothy Myers in 2013 had proposed that ''Bakonydraco'' actually belonged to the family
Tapejaridae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primi ...
, in a position slightly more basal than both '' Tapejara'' and '' Tupandactylus''. Indeed, the original paper describing this species compared the holotype jaw to '' Tapejara'' and ''
Sinopterus ''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. T ...
'', implicating its affinities to this clade (or at least a large amount of
convergence Convergence may refer to: Arts and media Literature *''Convergence'' (book series), edited by Ruth Nanda Anshen *Convergence (comics), "Convergence" (comics), two separate story lines published by DC Comics: **A four-part crossover storyline that ...
). If ''Bakonydraco'' is a tapejarid, it represents the only
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
record of Tapejaridae known to date (aside from the slightly older ''
Caiuajara dobruskii ''Caiuajara'' is an extinct genus of tapejarid pterosaur from the Late Cretaceous period (Santonian stage) of Brazil. It is known from a single type species, ''Caiuajara dobruskii''. Discovery and naming In 1971, the labourers Alexandre Dobru ...
''). A more recent phylogenetic study reinforces this placement. The cladogram on the left follows the 2014
phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
by Brian Andres and colleagues that also recovered ''Bakonydraco'' within the family Tapejaridae, more specifically within the tribe
Tapejarini Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primit ...
. In 2020, in a phylogenetic analysis conducted by David Martill and colleagues, ''Bakonydraco'' was once again found within the Tapejaridae, this time consisting of two lineages: the
Tapejarinae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primit ...
and the
Sinopterinae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The m ...
, ''Bakonydraco'' was recovered within the subfamily Sinopterinae in the basalmost position, unlike in the analysis by Andres and colleagues. Their cladogram is shown on the right. Topology 1: Andres ''et al.'' (2014). Topology 2: Martill ''et al.'' (2020).


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inval ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...


References

{{Portal bar, Paleontology, Cretaceous, Hungary Late Cretaceous pterosaurs of Europe Tapejaromorphs Taxa named by David B. Weishampel Fossil taxa described in 2005